Checklist |
Corresponding author: Lars Söderström ( lars.soderstrom@ntnu.no ) Academic editor: Lyubomir Penev
© 2016 Lars Söderström, Anders Hagborg, Matt von Konrat, Sharon Bartholomew-Began, David Bell, Laura Briscoe, Elizabeth Brown, D. Christine Cargill, Denise Pinheiro da Costa, Barbara J. Crandall-Stotler, Endymion Cooper, Gregorio Dauphin, John Engel, Kathrin Feldberg, David Glenny, S. Robbert Gradstein, Xiaolan He, Jochen Heinrichs, Joern Hentschel, Anna Luiza Ilkiu-Borges, Tomoyuki Katagiri, Nadezhda A. Konstantinova, Juan Larraín, David Long, Martin Nebel, Tamás Pócs, Felisa Puche, Elena Reiner-Drehwald, Matt Renner, Andrea Sass-Gyarmati, Alfons Schäfer-Verwimp, José Segarra-Moragues, Raymond E. Stotler, Phiangphak Sukkharak, Barbara Thiers, Jaime Uribe, Jiří Váňa, Juan Villarreal, Martin Wigginton, Li Zhang, Rui-Liang Zhu.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Söderström L, Hagborg A, von Konrat M, Bartholomew-Began S, Bell D, Briscoe L, Brown E, Cargill DC, Costa DP, Crandall-Stotler BJ, Cooper ED, Dauphin G, Engel JJ, Feldberg K, Glenny D, Gradstein SR, He X, Heinrichs J, Hentschel J, Ilkiu-Borges AL, Katagiri T, Konstantinova NA, Larraín J, Long DG, Nebel M, Pócs T, Felisa Puche F, Reiner-Drehwald E, Renner MAM, Sass-Gyarmati A, Schäfer-Verwimp A, Moragues JGS, Stotler RE, Sukkharak P, Thiers BM, Uribe J, Váňa J, Villarreal JC, Wigginton M, Zhang L, Zhu R-L (2016) World checklist of hornworts and liverworts. PhytoKeys 59: 1-821. https://doi.org/10.3897/phytokeys.59.6261
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A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online flora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 7486 species in 398 genera representing 92 families from the two phyla. The checklist has far reaching implications and applications, including providing a valuable tool for taxonomists and systematists, analyzing phytogeographic and diversity patterns, aiding in the assessment of floristic and taxonomic knowledge, and identifying geographical gaps in our understanding of the global liverwort and hornwort flora. The checklist is derived from a working data set centralizing nomenclature, taxonomy and geography on a global scale. Prior to this effort a lack of centralization has been a major impediment for the study and analysis of species richness, conservation and systematic research at both regional and global scales. The success of this checklist, initiated in 2008, has been underpinned by its community approach involving taxonomic specialists working towards a consensus on taxonomy, nomenclature and distribution.
Marchantiophyta , Anthocerophyta , nomenclature, taxonomy
This checklist is dedicated to Elizabeth Brown and Ray Stotler, who sadly left us during our work with it.
“……Charles Darwin wrote of his desire to provide financial support ‘for the formation of a perfect M.S. catalogue of all known plants’ (Darwin 1881, letter 13570). It is a personal embarrassment to me, and should be chastening to us all, that more than 120 years later we still have not delivered on that commitment.” (
The natural world is changing fast (
Version 1.0 of The Plant List (http://www.theplantlist.org/) was released in December 2010 aimed to be comprehensive for species of vascular plants (flowering plants, conifers, ferns and their allies) and of bryophytes (mosses, liverworts and hornworts), as a response to the 2010 Target 1 of the GSPC and to a clear global need for such data. The Plant List is a broad collaboration, coordinated by the Royal Botanic Gardens, Kew, and the Missouri Botanical Garden (MO), involving diverse partnerships.
Earlier,
Liverworts and hornworts (Figure
Habit images of selected lineages of Anthocerotophyta (hornworts) and Marchantiophyta (liverworts). APhaeomegaceros squamuliger, photo by J. Hollinger BHaplomitrium mnioides, photo by L. Zhang CConocephalum japonicum, photo by D. G. Long DColura vitiensis, photo by T. Pócs EPleurozia gigantea, photo by L. Söderström FRiccardia spegazziniana, photo by Juan Larraín.
Liverworts, hornworts and mosses offer microhabitats that are critical to the survival of a tremendous diversity of organisms such as single-celled eukaryotes, protozoa, and numerous groups of invertebrates (
There are estimated to exist only 215 hornwort species. By comparison estimates of liverwort species richness have varied considerably, by as much as 50%, with estimates ranging from 4,500 to 9,000 (e.g.
Below we provide context to the worldwide checklist, including a brief historical account and current informational resources, structure and layout, systematic and classification concepts, and methodology, including a brief overview of the underlying data set and nomenclatural elements.
Figure
Data availability and information needs associated with liverworts and hornworts have been reviewed extensively by
A major nomenclatural work is the Index Hepaticarum (http://www.ville-ge.ch/musinfo/bd/cjb/hepatic/index.php), which includes all effectively published liverwort epithets spanning 12 volumes with the closing date of 1973. The indices were prepared as a purely nomenclatural resource and did not claim to express any particular taxonomic concept. Recently,
The application of molecular phylogenetics continues to generate new insights into the evolutionary history of liverworts and hornworts. In recent years, inferences made from these phylogenies have especially revolutionized liverwort classification (
The systematic treatment as outlined here also includes the addition of subfamily and infrageneric classification; ranks not presented by
The checklist is presented in a taxonomic framework as outlined above with four main sections. The first is arranged systematically with taxa ordered alphabetically within the nearest higher rank. A brief discussion for selected families is provided in the main body of the text under the respective families. This does not appear uniformly for all families, but typically for larger more complicated families. Rather, a brief description is provided mainly where concepts depart from
*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (
There are comprehensive footnotes throughout the systematic section of the checklist. These are applied to selected names representing the rank of genus and below only. Footnotes without any literature citation represent observations from the authors of the respective family. Footnotes are wide ranging in their content. Generally, footnotes can be categorized into the following: i) general statements relating to taxonomic opinion, ii) comments on the type specimen, iii) comments about possible conspecificity, including citation of conflicting opinions, and iv) comment about species complexes.
This section is followed by a list of taxa in genera that we do not recognize. Those names represent taxa that have not been studied recently and have not been recognized in any recent treatment. However, many of those names are old and may gain priority over some recognized taxa once their identity is determined.
The next section is an alphabetical list of taxa of the ranks of species and below. The alphabetical list includes the confidence level, the taxon name and authority as well as a reference to the page of the corresponding entry in the systematic section. Thus, the alphabetical list provides a rapid gateway requiring no prior knowledge of higher taxonomy, but also serves as an index to the corresponding name in the systematic section that includes more detailed information.
The last section is a reference list where full bibliographic citations are given to all references for taxa included in the checklist. Titles and references have all been checked and verified, except in a few cases where we could not get hold of the publications.
Significantly, each accepted taxon is qualified using a three level ranking system that summarizes our knowledge about a taxon. The coding convention we are adopting largely follows that described by
The coding convention:
* Serious doubts. There are doubts about the value of the taxon. It can be that there are conflicting views without any substantial evidence in any direction, conflicting views with substantiating evidence supporting one or both positions, or evidence pointing towards synonymization but it is premature to do it. We have adhered to the principle that it is better to keep a taxon with one star (and preferably a note) than to synonymize it too quickly. Example: Bazzania asymmetrica is conspecific with Bazzania macgregorii in
** Knowledge problem. The taxon is not well known by the person evaluating it. It may be a newly described species or a species originally not well described and not restudied recently. Example: Jungermannia erectii Ajit P.Singh et V.Nath was recently described and has not been independently studied by someone with a global overview. It is therefore difficult to evaluate.
*** Accepted. A good taxon as currently understood based on personal experience or on taxonomic revisions that have been convincingly performed. Nomenclature and/or taxonomic position may, however, be questioned. Elements may be excluded from the taxon, but the taxon with the current type will still be accepted. Example: H. Bischler-Causse revised the genus Marchantia worldwide in a series of publications. Although she had a broad species concept, she also recognized infraspecific taxa. The taxa that she recognized without doubt should be accepted unless new evidence against it exists. Adopting a narrower species concept so that many of her subspecies are elevated to species does not change her view of what a good taxon is.
The foundation of the checklist is in the underlying data set from which it is derived. It was briefly described by
There has been an intense systematic effort focusing on data quality. In all but a few cases nomenclatural data has been verified against original publications. Söderström and Hagborg have checked and confirmed almost every original publication (three publications have not been available) for correct author, title and journal/book citation, date of publication as far as possible, page number for the protologue, and if the name is validly and legitimately published according to the International Code of Nomenclature for algae, fungi and plants (Melbourne Code;
The notes, published in the journal Phytotaxa, have included 38 authors from 13 countries. These are the results of several years of revisions of taxonomic groups by taxonomic experts. Following these revisions and working with participants identifying nomenclatural and taxonomic problems that would impact the worldwide checklist, Söderström, Hagborg and von Konrat led editorial efforts in compiling manuscripts with baseline information and nomenclatural or taxonomic issues that required resolving. The work was then coordinated with the taxonomic experts who drove and led the process and often identified further issues. The series of notes on taxonomy and nomenclature of liverworts and hornworts provided updated nomenclature, corrected invalid and illegitimate names and described new taxa based on studies (mainly molecular) that did not draw nomenclatural conclusions. Those notes are all open access and any changes were effective immediately as the Melbourne Code of Botanical Nomenclature allows publishing on internet from 1 Jan. 2012 (
This checklist builds on all published taxonomic and nomenclatural papers, that have come to our attention until June 30, 2015.
We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 8,078 taxa (species and below) in 7,486 species representing 398 genera, 92 families, 20 orders and 7 classes from the two phyla. The list includes 3,533 species with three stars, 2,988 species with two stars and 915 species with one star. The checklist also has extra utility in that it contains 3,106 references in the bibliography that serve as a powerful bibliographic resource for liverwort and hornwort systematic and taxonomic research.
The marked-up publication form of the current checklist by PhytoKeys provides a virtual instrument with a linked environment both internally (e.g. within an article) and externally (GBIF, IPNI, Tropicos, Wikispecies, etc.) that will undoubtedly help accelerate taxonomic research. The published checklist was the first phase in providing a worldwide list of accepted names. The next phase is to establish a generally recognized online repository to augment the huge underlying informational auxiliary data of over 25,000 publications, almost 39,000 published names, and the over 700,000 geographical observations. Several features of Web-based technology make it an essential tool, including 1) a vehicle to facilitate data access, 2) to unify the vastly scattered data on distributional information and nomenclature, 3) offering dynamic rather than static data in a searchable forum. The broader accessibility to the wealth of auxiliary data will help augment monographic and revisional work for many taxonomic groups, aid in identifying the need for increased floristic and survey work in many regions throughout the world, and have broad implications and applications beyond taxonomic research such as conservation science. The current Early Land Plants Today model has strong participation from taxonomic experts and an online resource will provide an opportunity to expand stakeholders to include ecologists, conservationists, scientists from other disciplines and general interest groups. However, such an effort can only be successful if it comes with sustained funding and infrastructure rather than depending on an ad hoc commitment by a few individuals, however dedicated.
The project will help augment monographic and revisional work for many taxonomic groups and aid in identifying the need for increased floristic and survey work in many regions throughout the world. Although there are many challenges ahead to obtain high quality data, quantifying global liverwort diversity is a tractable, multi-faceted and scientifically important goal, and everyone stands to gain by fostering this endeavour. The success of the project will lie on strong collaboration between institutions and the bryological community in general.
by J.C. Villarreal and D.C. Cargill
Notes on nomenclature and taxonomy can also be found in
*** Anthoceros L., Sp. Pl. 1: 1139, 1753 (
*** Anthoceros adscendens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 24, 1832 (
*** Anthoceros agrestis Paton, J. Bryol. 10 (3): 257, 1979 (
*** Anthoceros alpinus Steph., Sp. Hepat. (Stephani) 6: 425, 1923 (
* Anthoceros angustifolius Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 582, 1846 (
*** Anthoceros angustus Steph., Sp. Hepat. (Stephani) 5: 1001, 1916 (
*** Anthoceros bharadwajii Udar et A.K.Asthana, Proc. Indian Natl. Sci. Acad., B 51 (4): 484, 1985 (
** Anthoceros buettneri Steph., Sp. Hepat. (Stephani) 5: 997, 1916 (
*** Anthoceros capricornii Cargill et G.A.M.Scott, J. Hattori Bot. Lab. 82: 55, 1997 (
*** Anthoceros caucasicus Steph., Izv. Kavkazsk. Muz. 8: 87, 1914 (
** Anthoceros cavernosus Steph., Sp. Hepat. (Stephani) 5: 998, 1916 (
* Anthoceros chambensis Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 281, 1917 (
** Anthoceros chungii Khanna, J. Indian Bot. Soc. 17 (5/6): 316, 1938 (
** Anthoceros crispatus Griff., Not. pl. asiat. 2: 349, 1849 (
** Anthoceros dimorphus Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926 (
*** Anthoceros erectus Kashyap, New Phytol. 14 (1): 9, 1915 (
** Anthoceros expansus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 92, 2015 (
* Anthoceros ferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 5: 1007, 1916 (
*** Anthoceros fragilis Steph., Sp. Hepat. (Stephani) 5: 1006, 1916 (
*** Anthoceros fusiformis Austin, Bull. Torrey Bot. Club 6 (4): 28, 1875 [1876] (
*** Anthoceros fusiformis var. taiwanensis J.Haseg., Acta Phytotax. Geobot. 44 (2): 100, 1993 (
** Anthoceros gasongorii Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (
** Anthoceros granulatus Gottsche, Mexik. Leverm.: 275, 1863 (
** Anthoceros harrisanus (Steph.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 31, 1962 (
* Anthoceros helmsii Steph., Hedwigia 32 (3): 142, 1893 (
** Anthoceros jamesonii Taylor ex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 171, 1855 (
* Anthoceros javanicoides H.A.Mill., Phytologia 47 (4): 319, 1981 (
** Anthoceros jungermannioides Schwein., Spec. Fl. Amer. Crypt.: 25, 1821 (
* Anthoceros kajumas (K.I.Goebel) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
* Anthoceros koshii Khanna, J. Indian Bot. Soc. 15 (4): 237, 1936 (
*** Anthoceros lamellatus Steph., Sp. Hepat. (Stephani) 5: 1000, 1916 (
*** Anthoceros laminifer Steph., J. Linn. Soc., Bot. 29 (201): 266, 1892 (
*** Anthoceros macounii M.Howe, Bull. Torrey Bot. Club 25 (1): 19, 1898 (
*** Anthoceros macrosporus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (
** Anthoceros maritimus Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (
* Anthoceros megasporus Meijer, J. Hattori Bot. Lab. 18: 6, 1957 (
** Anthoceros muscoides Colenso, Trans. & Proc. New Zealand Inst. 16: 361, 1884 (
*** Anthoceros myriandroecius Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 134, 1911 (
** Anthoceros natalensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (
*** Anthoceros neesii Prosk., Leberm. Eur. 2 (9): 1312, 1958 (
** Anthoceros niger Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (
** Anthoceros orizabensis (Steph.) Hässel, Candollea 45 (1): 211, 1990 (
*** Anthoceros pandei Udar et A.K.Asthana, J. Indian Bot. Soc. 64: 305, 1985 (
*** Anthoceros patagonicus Hässel, Candollea 45 (1): 207, 1990 (
*** Anthoceros patagonicus subsp. gremmenii J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 32, 2013 (
** Anthoceros peruvianus Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (
** Anthoceros pinnatus Steph., Bol. Soc. Brot. 4: 154 [182], 1885 [1886] (
*** Anthoceros punctatus L., Sp. Pl. 1: 1139, 1753 (
* Anthoceros pusillus Colenso, Trans. & Proc. New Zealand Inst. 18: 255, 1886 (
*** Anthoceros rosulans J.Haseg., J. Hattori Bot. Lab. 60: 379, 1986 (
*** Anthoceros sambesianus Steph., Sp. Hepat. (Stephani) 5: 996, 1916 (
*** Anthoceros scariosus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 230, 1869 (
** Anthoceros schroederi Steph., 52 (5): 306, 1912 (
** Anthoceros serratus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911 (
** Anthoceros simulans M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 204, 1934 (
** Anthoceros spongiosus Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (
*** Anthoceros subtilis Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (
*** Anthoceros telaganus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (
*** Anthoceros tristanianus J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 33, 2013 (
*** Anthoceros tuberculatus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832 (
** Anthoceros venosus Lindenb. et Gottsche, Syn. Hepat. 4: 584, 1846 (
Excluded from the genus
* Anthoceros aethyopicus Gola, Ann. Bot. (Rome) 13 (1): 73, 1914 (
* Anthoceros brunnthaleri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (
* Anthoceros floribundus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (
* Anthoceros mildbraedii Steph., Sp. Hepat. (Stephani) 6: 428, 1923 (
* Anthoceros parvifrons Steph., 52 (5): 307, 1912 (
* Anthoceros pseudocostus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (
* Anthoceros rossoi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (
*** Folioceros D.C.Bharadwaj, Geophytology 1 (1): 9, 1971 (
*** Folioceros amboinensis (Schiffn.) Piippo, Acta Bot. Fenn. 148: 36, 1993 (
** Folioceros apiahynus (Steph.) Hässel, Candollea 45 (1): 215, 1990 (
** Folioceros argillaceus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (
*** Folioceros assamicus D.C.Bharadwaj, Geophytology 1 (1): 10, 1971 (
** Folioceros dilatatus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (
*** Folioceros dixitianus (Mahab.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros fuciformis (Mont.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros glandulosus (Lehm. et Lindenb.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros incurvus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros indicus D.C.Bharadwaj, Geophytology 8 (1): 114, 1978 (
*** Folioceros kashyapii S.C.Srivast. et A.K.Asthana, Bryologist 92 (2): 219, 1989 (
*** Folioceros mangaloreus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros paliformis D.K.Singh, Bull. Bot. Surv. India 29: 176, 1987 [1989] (
*** Folioceros physocladus D.C.Bharadwaj, Geophytology 8 (1): 115, 1978 (
*** Folioceros pinnilobus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (
*** Folioceros satpurensis D.C.Bharadwaj et K.P.Srivast., Geophytology 8 (1): 112, 1978 (
*** Folioceros udarii A.K.Asthana et S.C.Srivast., Cryptog. Bryol. Lichénol. 7 (2): 151, 1986 (
*** Folioceros verruculosus (J.Haseg.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 383, 2011 (
by J.C. Villarreal and D.C. Cargill
*** Dendroceros Nees, Syn. Hepat. 4: 579, 1846 (
** subg. Apoceros R.M.Schust., Phytologia 63 (3): 200, 1987 (
*** Dendroceros cavernosus J.Haseg., J. Hattori Bot. Lab. 47: 306, 1980 (
*** Dendroceros difficilis Steph., Sp. Hepat. (Stephani) 5: 1009, 1917 (
*** Dendroceros muelleri Steph., Hedwigia 28 (2): 133, 1889 (
*** Dendroceros ogeramnangus Piippo, Acta Bot. Fenn. 148: 40, 1993 (
*** Dendroceros pedunculatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (
** Dendroceros subdifficilis S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 119, 1951 (
** subg. Dendroceros
*** Dendroceros acutilobus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (
*** Dendroceros borbonicus Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 31, 1893 [1894] (
*** Dendroceros crispus (Sw.) Nees, Syn. Hepat. 4: 581, 1846 (
*** Dendroceros foliicola J.Haseg., J. Hattori Bot. Lab. 47: 296, 1980 (
*** Dendroceros japonicus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 15, 1909 (
*** Dendroceros javanicus (Nees) Nees, Syn. Hepat. 4: 582, 1846 (
*** Dendroceros subplanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 20, 1909 (
*** Dendroceros tubercularis S.Hatt., Bot. Mag. (Tokyo) 58 (685): 6, 1944 (
*** Dendroceros validus Steph., Sp. Hepat. (Stephani) 5: 1016, 1917 (
** Dendroceros vesconianus Gottsche, J. Bot. (Morot) 12: 150, 1898 (
** Dendroceros wattsianus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (
Incertae sedis
* Dendroceros adglutinatus (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 580, 1846 (
*** Dendroceros africanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (
** Dendroceros allionii Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (
** Dendroceros australis Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (
** Dendroceros breutelii Nees, Syn. Hepat. 4: 581, 1846 (
** Dendroceros breutelii var. surinamensis Lindenb. et Gottsche, Linnaea 24 (6): 639, 1851 [1852] (
*** Dendroceros cichoraceus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 16, 1858 (
* Dendroceros crassicostatus Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (
** Dendroceros crassinervis (Nees) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 18, 1858 (
*** Dendroceros crispatus (Hook.) Nees, Syn. Hepat. 4: 579, 1846 (
** Dendroceros crispatus var. simplicior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 574, 1885 (
** Dendroceros cucullatus Steph., Sp. Hepat. (Stephani) 6: 429, 1923 (
* Dendroceros exalatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (
* Dendroceros gracilis Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (
*** Dendroceros granulatus Mitt., Fl. vit.: 419, 1871 [1873] (
** Dendroceros herasii M.Infante, J. Bryol. 32 (4): 285, 2010 (
** Dendroceros humboldtensis Hürl., Bauhinia 1 (3): 254, 1960 (
** Dendroceros paivae C.A.Garcia, Sérgio et J.C.Villarreal, Cryptog. Bryol. 33 (1): 5, 2012 (
* Dendroceros rarus Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (
* Dendroceros reticulus Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 37, 1950 [1951] (
** Dendroceros rigidus Steph., Sp. Hepat. (Stephani) 5: 1017, 1917 (
*** Dendroceros seramensis J.Haseg., Acta Phytotax. Geobot. 37 (1/3): 10, 1986 (
** Dendroceros subtropicus C.J.Wild, Trans. Nat. Hist. Soc. Queensland 1: 49, 1893 (
** Dendroceros tahitensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 138, 1873 (
*** Megaceros Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (
** Megaceros aneuriformis Steph., Sp. Hepat. (Stephani) 5: 949, 1916 (
*** Megaceros austronesophilus Cargill et Seppelt, Austral. Syst. Bot. 26 (5): 372, 2013 (
** Megaceros ciliatus K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 74, 1928 (
*** Megaceros denticulatus (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 956, 1916 (
*** Megaceros flagellaris (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 951, 1916 (
*** Megaceros gracilis (Reichardt) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (
** Megaceros leptohymenius (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (
*** Megaceros pellucidus (Colenso) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 115, 1972 (
** Megaceros tjibodensis Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (
Excluded from the genus
* Megaceros flavens (Spruce) Campb., Ann. Bot. (Oxford) 21 (4): 483, 1907 (
* Megaceros jamesonii (Taylor) Steph., Biblioth. Bot. 87 (2): 268, 1916 (
*** Nothoceros (R.M.Schust.) J.Haseg., J. Hattori Bot. Lab. 76: 32, 1994 (
*** Nothoceros aenigmaticus J.C.Villarreal et K.D.McFarland, Bryologist 113 (1): 109, 2010 (
*** Nothoceros canaliculatus (Pagán) J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (
*** Nothoceros endiviifolius (Mont.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (
** Nothoceros fuegiensis (Steph.) J.C.Villarreal, Bryologist 113 (1): 109, 2010 (
*** Nothoceros giganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (
*** Nothoceros minarum (Nees) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (
** Nothoceros renzagliensis J.C.Villarreal, L.V.Campos et Uribe, Syst. Bot. 37 (1): 32, 2012 (
*** Nothoceros schizophyllus (Steph.) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (
*** Nothoceros superbus J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 280, 2007 (
*** Nothoceros vincentianus (Lehm. et Lindenb.) J.C.Villarreal, Bryologist 113 (1): 111, 2010 (
** Phaeomegaceros R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros coriaceus (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros fimbriatus (Gottsche) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros foveatus (J.Haseg.) J.C.Villarreal, Nova Hedwigia 91 (3/4): 352, 2010 (
*** Phaeomegaceros hirticalyx (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros plicatus (Mitt.) J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 85, 2013 (
*** Phaeomegaceros skottsbergii (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (
*** Phaeomegaceros squamuliger (Spruce) J.C.Villarreal, Nova Hedwigia 91 (3/4): 351, 2010 (
** Phaeomegaceros squamuliger subsp. hasselii J.C.Villarreal, Cargill et Goffinet, Nova Hedwigia 91 (3/4): 352, 2010 (
by J.C. Villarreal and D.C. Cargill
*** Phymatoceros Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 113, 2005 (
*** Phymatoceros bulbiculosus (Brot.) Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 114, 2005 (
*** Phymatoceros phymatodes (M.Howe) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 240, 2007 (
by J.C. Villarreal and D.C. Cargill
*** Notothylas Sull. ex A.Gray, Amer. J. Sci. Arts (ser. 2) 1 (1): 74, 1846 (
** subg. Notothylas
*** Notothylas anaporata Udar et D.K.Singh, Rev. Bryol. Lichénol. 45 (2): 202, 1979 (
*** Notothylas breutelii (Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 21, 1858 (
*** Notothylas depressispora J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 26, 1979 (
*** Notothylas dissecta Steph., Sp. Hepat. (Stephani) 5: 1020, 1917 (
*** Notothylas himalayensis Udar et D.K.Singh, J. Bryol. 11 (3): 451, 1981 (
*** Notothylas indica Kashyap, Proc. Lahore Philos. Soc. 4: 54, 1925 (
*** Notothylas javanica (Sande Lac.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 20, 1858 (
*** Notothylas orbicularis (Schwein.) Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 75, 1846 (
*** Notothylas pandei Udar et V.Chandra, Geophytology 7 (2): 142, 1977 (
** subg. Notothyloides A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 106, 1991 (
*** Notothylas khasiana Udar et D.K.Singh, J. Indian Bot. Soc. 60: 112, 1981 (
*** Notothylas pfleidereri Udar et D.K.Singh, Lindbergia 5 (1): 28, 1979 (
Incertae sedis
*** Notothylas decurva (Mitt.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (
*** Notothylas flabellata Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (
*** Notothylas galapagensis M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 203, 1934 (
** Notothylas irregularis Chantanaorr., Acta Bot. Hung. 56 (3/4): 270, 2014 (
*** Notothylas kashyapii D.K.Singh, Indian J. Forest. 23 (4): 386, 2000 (
*** Notothylas nepalensis D.K.Singh, J. Bombay Nat. Hist. Soc. 84 (3): 650, 1987 [1988] (
*** Notothylas temperata J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 20, 1979 (
*** Notothylas udarii D.K.Singh et Semwal, Phytotaxonomy 1: 35, 2001 (
* Notothylas verdoornii Khanna, Rev. Bryol. Lichénol. 6: 118, 1933 (
*** Notothylas vitalii Udar et D.K.Singh, Misc. Bryol. Lichenol. 8 (9): 173, 1980 (
** Notothylas yunnanensis T.Peng et R.L.Zhu, Phytotaxa 156 (3): 157, 2014 (
* Mesoceros Piippo, Acta Bot. Fenn. 148: 30, 1993 (
* Mesoceros mesophoros Piippo, Acta Bot. Fenn. 148: 30, 1993 (
* Mesoceros porcatus Piippo, Haussknechtia, Beih. 9: 279, 1999 (
** Paraphymatoceros Hässel, Phytologia 88 (2): 208, 2006 (
*** Paraphymatoceros diadematus Hässel, Phytologia 88 (2): 209, 2006 (
*** Paraphymatoceros hallii (Austin) Hässel, Phytologia 88 (2): 209, 2006 (
*** Paraphymatoceros pearsonii (M.Howe) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (
*** Paraphymatoceros proskaueri (Stotler, Crand.-Stotl. et W.T.Doyle) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (
*** Phaeoceros Prosk., Bull. Torrey Bot. Club 78 (4): 346, 1951 (
** Phaeoceros austroandinus Hässel, Candollea 44 (2): 721, 1989 (
** Phaeoceros bolusii (Sim) S.W.Arnell, Hepat. South Africa: 401, 1963 (
** Phaeoceros brevicapsulus (Steph.) Hässel, Candollea 44 (2): 725, 1989 (
*** Phaeoceros carolinianus (Michx.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
*** Phaeoceros delicatus E.O.Campb. et Outred, New Zealand J. Bot. 33 (3): 285, 1995 (
*** Phaeoceros dendroceroides (Steph.) Hässel, Beih. Nova Hedwigia 134: 487, 2009 (
*** Phaeoceros engelii Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 248, 2008 (
** Phaeoceros erectus Udar et D.K.Singh, Geophytology 11 (2): 257, 1981 (
*** Phaeoceros evanidus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 245, 2008 (
*** Phaeoceros exiguus (Steph.) J.Haseg., J. Hattori Bot. Lab. 60: 387, 1986 (
*** Phaeoceros flexivalvis (Gottsche et Nees) Hässel, Candollea 44 (2): 728, 1989 (
*** Phaeoceros fulvisporus (Steph.) J.Haseg., Trop. Bryol. 8: 52, 1993 (
*** Phaeoceros gemmifer (Horik.) J.Haseg., J. Hattori Bot. Lab. 57: 252, 1984 (
** Phaeoceros gualaquizanus (Steph.) Gradst., J. Hattori Bot. Lab. 45: 123, 1979 (
*** Phaeoceros himalayensis (Kashyap) Prosk. ex Bapna et G.G.Vyas, J. Hattori Bot. Lab. 25: 88, 1962 (
*** Phaeoceros huebschmannii Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 301, 1986 (
*** Phaeoceros inflatus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 246, 2008 (
*** Phaeoceros kashyapii A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 129, 1991 (
*** Phaeoceros laevis (L.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
** Phaeoceros maranguensis (Steph.) Bapna, Univ. Udaipur Res. Stud. 3: 138, 1966 (
* Phaeoceros microsporus (Steph.) Hässel, Candollea 44 (2): 721, 1989 (
*** Phaeoceros minutus (Mitt.) S.W.Arnell, Hepat. South Africa: 403, 1963 (
*** Phaeoceros mohrii (Austin) Hässel, Candollea 44 (2): 721, 1989 (
*** Phaeoceros oreganus (Austin) Hässel, Candollea 44 (2): 718, 1989 (
*** Phaeoceros parvulus (Schiffn.) J.Haseg., J. Hattori Bot. Lab. 57: 248, 1984 (
*** Phaeoceros perpusillus Chantanaorr., Acta Bot. Hung. 51 (1/2): 30, 2009 (
** Phaeoceros pichinchensis (Spruce) Hässel, Candollea 44 (2): 725, 1989 (
** Phaeoceros propagulifer (Steph.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (
* Phaeoceros striatisporus J.Haseg., J. Hattori Bot. Lab. 75, 268 (
*** Phaeoceros tenuis (Spruce) Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 303, 1986 (
** Phaeoceros tigrinus (Gola) J.C.Villarreal, Phytotaxa 208 (1): 95, 2015 (
*** Phaeoceros tuberosus (Taylor) Prosk., J. Indian Bot. Soc. 42A: 185, 1964 (
*** Phaeoceros udarii A.K.Asthana et V.Nath, Proc. Natl. Acad. Sci. India, B 63 (4): 461, 1993 (
** Phaeoceros wrightii (Steph.) Hässel, Candollea 44 (2): 722, 1989 (
by J.C. Villarreal and D.C. Cargill
*** Leiosporoceros Hässel, J. Bryol. 14 (2): 255, 1986 (
*** Leiosporoceros dussii (Steph.) Hässel, J. Bryol. 14 (2): 255, 1986 (
by S. Bartholomew-Began
The treatment of Haplomitriaceae follows
*** Haplomitrium Nees, Naturgesch. Eur. Leberm. 1: 109, 1833 (
*** subg. Calobryum (Nees) R.M.Schust., Nova Hedwigia 13 (1/2): 40, 1967 (
*** Haplomitrium blumei (Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (
*** Haplomitrium mnioides (Lindb.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (
*** subg. Haplomitrium
** sect. Archibryum (R.M.Schust.) J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 675, 1981 (
*** Haplomitrium gibbsiae (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (
*** Haplomitrium intermedium Berrie, Proc. Linn. Soc. New South Wales (ser. 2) 87 (399): 191, 1963 (
** sect. Haplomitrium
*** Haplomitrium hookeri (Lyell ex Sm.) Nees, Naturgesch. Eur. Leberm. 1: 111, 1833 (
*** Haplomitrium hookeri var. minutum (E.O.Campb.) Barthol.-Began, Bryophyt. Biblioth. 41: 230, 1991 (
*** Haplomitrium monoicum J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 668, 1981 [1982] (
*** Haplomitrium ovalifolium R.M.Schust., Bryologist 74 (2): 136, 1971 (
by R. Stotler and B.J. Crandall-Stotler
*** Apotreubia S.Hatt. et Mizut., Bryologist 69 (4): 491, 1966 [1967] (
*** Apotreubia hortoniae Konstant., Phytotaxa 76 (3): 33, 2013 (
*** Apotreubia nana (S.Hatt. et Inoue) S.Hatt. et Mizut., Bryologist 69 (4): 492, 1966 [1967] (
** Apotreubia pusilla (R.M.Schust.) Grolle, Acta Bot. Fenn. 125: 63, 1984 (
*** Apotreubia yunnanensis Higuchi, Cryptog. Bryol. Lichénol. 19 (4): 321, 1998 (
*** Treubia K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (
*** Treubia insignis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (
** Treubia insignis subsp. bracteata (Steph.) R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 241, 1969 (
** Treubia insignis subsp. caledonica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 243, 1969 (
** Treubia insignis subsp. vitiensis R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 242, 1969 (
*** Treubia lacunosa (Colenso) Prosk., Bryologist 58 (3): 199, 1955 (
** Treubia lacunosoides T.Pfeiff., W.Frey et M.Stech, Nova Hedwigia 75 (1/2): 249, 2002 (
*** Treubia pygmaea R.M.Schust., Phytologia 56 (7): 460, 1985 (
*** Treubia scapanioides R.M.Schust., J. Hattori Bot. Lab. 32: 246, 1969 (
*** Treubia tahitensis (Nadeaud) Besch., J. Bot. (Morot) 12: 147, 1898 (
*** Treubia tasmanica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 248, 1969 (
by K. Feldberg, J. Váňa and J. Heinrichs
The subfamily Jamesonielloideae was excluded from Lophoziaceae/Scapaniaceae by
*** Adelanthoideae K.Feldberg, Heinrichs et Váňa
*** Adelanthus Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (
** sect. Adelanthus
*** Adelanthus falcatus (Hook.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (
*** Adelanthus occlusus (Hook.f. et Taylor) Carrington, Trans. Bot. Soc. Edinburgh 10: 381, 1870 (
** sect. Calyptrocolea (R.M.Schust.) Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (
*** Adelanthus aureomarginatus R.M.Schust., Phytologia 39 (4): 250, 1978 (
*** Adelanthus gemmiparus (R.M.Schust.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Biol. Sci. 11 (18): 241, 1970 (
** Adelanthus lingulatus J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 26, 2013 (
*** Adelanthus tenuis J.J.Engel et Grolle, J. Hattori Bot. Lab. 35: 333, 1972 (
** sect. Lindenbergiani Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (
*** Adelanthus carabayensis (Mont.) Grolle, J. Hattori Bot. Lab. 35: 348, 1972 (
*** Adelanthus integerrimus Grolle, J. Hattori Bot. Lab. 35: 340, 1972 (
*** Adelanthus lindenbergianus (Lehm.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 244, 1864 (
** sect. Pittieri Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (
*** Adelanthus pittieri (Steph.) Grolle, J. Hattori Bot. Lab. 35: 337, 1972 (
*** Adelanthus squarrosus Grolle, J. Hattori Bot. Lab. 67: 243, 1989 (
*** Pseudomarsupidium Herzog, Svensk Bot. Tidskr. 47 (1): 42, 1953 (
*** Pseudomarsupidium aureocinctum (R.M.Schust.) J.J.Engel, Novon 17 (3): 312, 2007 (
*** Pseudomarsupidium borneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 60, 2012 (
*** Pseudomarsupidium decipiens (Hook.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (
*** Pseudomarsupidium piliferum (Steph.) Herzog ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (
*** Wettsteinia Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 44, 1898 (
*** Wettsteinia densiretis (Herzog) Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (
*** Wettsteinia inversa (Sande Lac.) Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 45, 1898 (
*** Wettsteinia rotundifolia (Horik.) Grolle, J. Hattori Bot. Lab. 28: 100, 1965 (
*** Wettsteinia schusteriana Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (
** Jamesonielloideae Inoue
*** Cuspidatula Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1141 (124), 1901 (
*** Cuspidatula contracta (Reinw., Blume et Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1141 (124), 1901 (
*** Cuspidatula flaccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (
*** Cuspidatula flexicaulis (Nees) Váňa et L.Söderstr., Phytotaxa 76 (3): 35, 2013 (
*** Cuspidatula kirkii (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (
*** Cuspidatula monodon (Taylor) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1143 (126), 1901 (
*** Cuspidatula orbicularis (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (
*** Cuspidatula robusta (Austin) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (
** Denotarisia Grolle, Feddes Repert. 82 (1): 6, 1971 (
*** Denotarisia linguifolia (De Not.) Grolle, Feddes Repert. 82 (1): 6, 1971 (
** Nothostrepta R.M.Schust., Phytologia 45 (5): 420, 1980 (
*** Nothostrepta bifida (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (
*** Nothostrepta longissima (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (
** Pisanoa Hässel, Lindbergia 14 (3): 179, 1988 [1989] (
*** Pisanoa chilensis Hässel, Lindbergia 14 (3): 179, 1988 [1989] (
** Protosyzygiella (Inoue) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (
*** Protosyzygiella pseudoconnexa (Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (
*** Syzygiella Spruce, J. Bot. 14: 234, 1876 (
*** subg. Anomalae (Inoue) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella anomala (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 471 (190), 1902 (
*** Syzygiella bilobata Inoue, J. Hattori Bot. Lab. 29: 186, 1966 (
** Syzygiella ciliata Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 115, 2014 (
*** Syzygiella concreta (Gottsche) Spruce, J. Bot. 14: 234, 1876 (
*** Syzygiella manca (Mont.) Steph., Hedwigia 31 (1): 14, 1892 (
*** Syzygiella pectiniformis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 501, 1885 (
*** Syzygiella tonduzana Steph., Sp. Hepat. (Stephani) 6: 118, 1917 (
*** Syzygiella trigonifolia (Steph.) Herzog, Hedwigia 74 (2): 87, 1934 (
*** subg. Cryptochila (R.M.Schust.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella acinacifolia (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella nigrescens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella paludosa (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella pseudocclusa (E.A.Hodgs.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella sonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (
*** Syzygiella spegazziniana (Spruce ex C.Massal.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** subg. Pseudoplagiochila Inoue, J. Hattori Bot. Lab. 29: 182, 1966 (
*** Syzygiella ovalifolia Inoue, J. Hattori Bot. Lab. 29: 191, 1966 (
* Syzygiella securifolia (Nees) Inoue, J. Hattori Bot. Lab. 46: 232, 1979 (
*** Syzygiella subintegerrima (Reinw., Blume et Nees) Spruce, J. Linn. Soc., Bot. 30 (210): 362, 1895 (
*** Syzygiella tasmanica (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** subg. Roivainenia (Perss.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** Syzygiella jacquinotii (Mont.) Hentschel, K.Feldberg, Váňa et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** subg. Syzygiella
*** Syzygiella autumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** Syzygiella campanulata Herzog, Rev. Bryol. Lichénol. 11 (1): 9, 1938 [1939] (
*** Syzygiella colorata (Lehm.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
** Syzygiella colorata var. collenchymata J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 102, 2013 (
*** Syzygiella contigua Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 470 (189), 1902 (
*** Syzygiella elongella (Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (
*** Syzygiella macrocalyx (Mont.) Spruce, J. Bot. 14: 234, 1876 (
*** Syzygiella nipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
** Syzygiella oenops (Lindenb. et Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
*** Syzygiella perfoliata (Sw.) Spruce, J. Bot. 14: 234, 1876 (
*** Syzygiella purpurascens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
*** Syzygiella rubricaulis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 468 (187), 1902 (
*** Syzygiella setulosa Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 469 (188), 1902 (
*** Syzygiella teres (Carrington et Pearson) Váňa, Phytotaxa 76 (3): 35, 2013 (
*** Syzygiella undata (Mont.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (
Incertae sedis
*** Syzygiella eatonii (Austin) Inoue, J. Jap. Bot. 37 (12): 359, 1962 (
*** Syzygiella uleana Steph., Hedwigia 44 (4): 224, 1905 (
** Vanaea (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (
*** Vanaea plagiochiloides (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (
by J. Váňa and L. Söderström
Anastrophyllaceae was described by
*** Anastrepta (Lindb.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (
*** Anastrepta orcadensis (Hook.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (
*** Anastrophyllum (Spruce) Steph., Hedwigia 32 (3): 139, 1893 (
*** Anastrophyllum alpinum Steph., Sp. Hepat. (Stephani) 6: 103, 1917 (
*** Anastrophyllum assimile (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum auritum (Lehm.) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1137 (120), 1901 (
*** Anastrophyllum ciliatum Steph., Hedwigia 32 (3): 139, 1893 (
*** Anastrophyllum donnianum (Hook.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum ellipticum Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (1): 13, 1978 (
** Anastrophyllum esenbeckii (Mont.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum fissum Steph., Bull. Herb. Boissier 5 (10): 845, 1897 (
*** Anastrophyllum joergensenii Schiffn., Hedwigia 49 (4): 396, 1910 (
* Anastrophyllum lignicola D.B.Schill et D.G.Long, Ann. Bot. Fenn. 39 (2): 130, 2002 (
*** Anastrophyllum michauxii (F.Weber) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 289, 1932 [1933] (
*** Anastrophyllum nigrescens (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum obtusum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 285, 1950 (
*** Anastrophyllum piligerum (Nees) Steph., Hedwigia 32 (3): 140, 1893 (
*** Anastrophyllum squarrosum Herzog, Ann. Bryol. 5: 72, 1932 (
*** Anastrophyllum stellatum R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Anastrophyllum tubulosum (Nees) Grolle, J. Hattori Bot. Lab. 28: 101, 1965 (
*** Barbilophozia Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
** subg. Barbilophozia
*** Barbilophozia barbata (Schmidel ex Schreb.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
*** Barbilophozia hatcheri (A.Evans) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
*** Barbilophozia lycopodioides (Wallr.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (
*** Barbilophozia rubescens (R.M.Schust. et Damsh.) Kartt. et L.Söderstr., Ann. Bot. Fenn. 29 (2): 120, 1992 (
** subg. Sudeticae (Schljakov) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Barbilophozia sudetica (Nees ex Huebener) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Biantheridion (Grolle) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Biantheridion undulifolium (Nees) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Chandonanthus Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
*** Chandonanthus squarrosus (Menzies) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
*** Crossocalyx Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (
*** Crossocalyx hellerianus (Nees ex Lindenb.) Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (
*** Crossocalyx tenuis (Harry Williams) Schljakov, Novosti Sist. Nizš. Rast. 15: 246, 1978 (
*** Gymnocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
*** Gymnocolea borealis (Frisvoll et Moen) R.M.Schust., Lindbergia 12 (1): 7, 1986 (
** Gymnocolea fascinifera Potemkin, Arctoa 2: 76, 1993 (
*** Gymnocolea inflata (Huds.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
* Gymnocolea inflata subsp. acutiloba (Schiffn.) R.M.Schust. et Damsh. ex L.Söderstr. et Váňa, Lindbergia 27 (1): 43, 2002 (
*** Hamatostrepta Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (
*** Hamatostrepta concinna Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (
** Hattoria R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 69, 1961 (
*** Hattoria yakushimensis (Horik.) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 70, 1961 (
*** Isopaches H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 287, 1932 [1933] (
*** Isopaches alboviridis (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (
*** Isopaches bicrenatus (Schmidel ex Hoffm.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (
*** Isopaches decolorans (Limpr.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (
*** Isopaches pumicicola (Berggr.) Bakalin, Arctoa 17: 162, 2008 [2009] (
*** Neoorthocaulis L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (
*** Neoorthocaulis attenuatus (Mart.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (
*** Neoorthocaulis binsteadii (Kaal.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (
*** Neoorthocaulis floerkei (F.Weber et D.Mohr) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
** Neoorthocaulis hyperboreus (R.M.Schust.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Orthocaulis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 293, 1932 [1933] (
*** Orthocaulis atlanticus (Kaal.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 294, 1932 [1933] (
* Orthocaulis cavifolius H.Buch et S.W.Arnell, Memoranda Soc. Fauna Fl. Fennica 26: 71, 1951 (
** Plicanthus R.M.Schust., Nova Hedwigia 74 (3/4): 484, 2002 (
*** Plicanthus birmensis (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 223, 2002 (
** Plicanthus difficilis (Steph.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 30, 2013 (
** Plicanthus giganteus (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 485, 2002 (
*** Plicanthus hirtellus (F.Weber) R.M.Schust., Nova Hedwigia 74 (3/4): 492, 2002 (
*** Schizophyllopsis Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
** Schizophyllopsis aristata (Herzog ex N.Kitag.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
*** Schizophyllopsis bidens (Reinw., Blume et Nees) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
** Schizophyllopsis lanciloba (Steph.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
*** Schizophyllopsis papillosa (J.J.Engel et Braggins) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (
*** Schizophyllopsis sphenoloboides (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 152 (1): 49, 2013 (
*** Schljakovia Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Schljakovia kunzeana (Huebener) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Schljakovianthus Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Schljakovianthus quadrilobus (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Sphenolobopsis R.M.Schust. et N.Kitag., Nova Hedwigia 22: 152, 1971 [1972] (
*** Sphenolobopsis pearsonii (Spruce) R.M.Schust., Nova Hedwigia 22: 153, 1971 [1972] (
*** Sphenolobus (Lindb.) Berggr., New Zealand Hepat.: 22, 1898 (
*** Sphenolobus austroamericanus (Váňa) Váňa, Phytotaxa 81 (1): 30, 2013 (
*** Sphenolobus minutus (Schreb. ex D.Crantz) Berggr., New Zealand Hepat.: 22, 1898 (
*** Sphenolobus saxicola (Schrad.) Steph., Bull. Herb. Boissier (sér. 2) 2 (2): 168 (160), 1902 (
** Tetralophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 13: 227, 1976 (
*** Tetralophozia cavallii (Gola) Váňa, Trop. Bryol. 8: 102, 1993 (
*** Tetralophozia filiformis (Steph.) Urmi, J. Bryol. 12 (3): 394, 1983 (
** Tetralophozia pilifera (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 482, 2002 (
*** Tetralophozia setiformis (Ehrh.) Schljakov, Novosti Sist. Nizš. Rast. 13: 228, 1976 (
*** Zantenia (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (
*** Zantenia borneensis (Herzog) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (
*** Zantenia denticulata (Grolle) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (
*** Zantenia karstenii (Schiffn.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (
*** Zantenia prionophylla (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (
by J. Váňa with contributions by S.R. Gradstein (Odontoschisma)
Cephaloziaceae was recently studied by
** Alobiella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
*** Alobiella husnotii (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
** Alobiellopsis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (
*** Alobiellopsis acroscypha (Spruce) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (
*** Alobiellopsis dominicensis (Spruce) Fulford, Mem. New York Bot. Gard. 11 (3): 350, 1968 (
*** Alobiellopsis heteromorpha (Lehm.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 682, 1969 (
*** Alobiellopsis parvifolia (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 679, 1969 (
*** Alobiellopsis pillansii (Sim) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 683, 1969 (
*** Cephalozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
*** Cephalozia acuminata (Herzog) Váňa, Phytotaxa 112 (1): 11, 2013 (
** Cephalozia acutiloba (Inoue) Váňa, Phytotaxa 112 (1): 11, 2013 (
*** Cephalozia albula Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephalozia ambigua C.Massal., Malpighia 21 (7/8): 310, 1907 (
*** Cephalozia austrigena R.M.Schust. ex J.J.Engel, Novon 17 (3): 312, 2007 (
*** Cephalozia badia (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 483 (313), 1908 (
*** Cephalozia bicuspidata (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
* Cephalozia bicuspidata subsp. lammersiana (Huebener) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 730, 1974 (
*** Cephalozia chilensis (J.J.Engel et R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 29, 2002 (
*** Cephalozia conchata (Grolle et Váňa) Váňa, Syst. Bot. 40 (1): 38, 2015 (
*** Cephalozia crossii Spruce, Cephalozia: 46, 1882 (
*** Cephalozia darjeelingensis Udar et D.Kumar, Geophytology 6 (1): 36, 1976 (
*** Cephalozia drucei (R.M.Schust.) Váňa, Phytotaxa 112 (1): 11, 2013 (
*** Cephalozia fuegiensis Váňa, Phytotaxa 112 (1): 12, 2013 (
*** Cephalozia hamatiloba Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 427 (303), 1908 (
*** Cephalozia hamatiloba subsp. siamensis (N.Kitag.) Váňa, Acta Bot. Fenn. 177: 16, 2004 (
*** Cephalozia lacinulata (J.B.Jack ex Gottsche et Rabenh.) Spruce, Cephalozia: 45, 1882 (
*** Cephalozia lucens (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908 (
*** Cephalozia macgregorii (Steph.) Váňa, Phytotaxa 112 (1): 12, 2013 (
*** Cephalozia macounii (Austin) Austin, Hepat. bor.-amer.: 14, 1873 (
*** Cephalozia maxima Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (
*** Cephalozia mollusca (De Not.) Váňa, Phytotaxa 112 (1): 12, 2013 (
* Cephalozia neesiana Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 429 (305), 1908 (
*** Cephalozia nishimurae (N.Kitag.) Váňa, Phytotaxa 112 (1): 12, 2013 (
** Cephalozia pachygyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (
** Cephalozia physocaula (Hook.f. et Taylor) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 485 (315), 1908 (
** Cephalozia schusteriana J.J.Engel, Novon 17 (3): 313, 2007 (
*** Cephalozia stolonacea (Herzog) Váňa, Phytotaxa 112 (1): 12, 2013 (
*** Cephalozia tubulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (
Excluded from the genus
* Cephalozia hians Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (
* Cephalozia indica Udar et D.Kumar, Geophytology 6 (1): 42, 1976 (
* Cephalozia kodaikanalensis G.Asthana et Saumya Srivast., Geophytology 43 (1): 63, 2013 (
* Cephalozia parvifolia Arnell, Rev. Bryol. 25 (1): 1, 1898 (
* Cephalozia tricuspidata (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (
*** Fuscocephaloziopsis Fulford, Mem. New York Bot. Gard. 11 (3): 353, 1968 (
** Fuscocephaloziopsis affinis (Lindb. ex Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis africana (Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis albescens (Hook.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
* Fuscocephaloziopsis albescens var. islandica (Nees) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis baldwinii (C.M.Cooke) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
** Fuscocephaloziopsis biloba (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (
*** Fuscocephaloziopsis catenulata (Huebener) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis catenulata subsp. nipponica (S.Hatt.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis connivens (Dicks.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis connivens subsp. fissa (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis connivens subsp. sandwicensis (Mont.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (
*** Fuscocephaloziopsis crassifolia (Lindenb. et Gottsche) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis gollanii (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis leucantha (Spruce) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis loitlesbergeri (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis lunulifolia (Dumort.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis macrostachya (Kaal.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis macrostachya subsp. australis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis macrostachya subsp. macrostachya var. spiniflora (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (
*** Fuscocephaloziopsis monticola (J.D.Godfrey) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Fuscocephaloziopsis pachycaulis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Fuscocephaloziopsis pleniceps (Austin) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
** Fuscocephaloziopsis pleniceps var. caroliniana (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Fuscocephaloziopsis pulvinata (Steph.) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (
* Fuscocephaloziopsis schusteri (Sushil K.Singh et D.K.Singh) Váňa, Phytotaxa 183 (4): 291, 2014 (
*** Fuscocephaloziopsis subintegra Gradst. et Váňa, Cryptog. Bryol. 25 (3): 274, 2004 (
*** Fuscocephaloziopsis zoopsioides (Horik.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (
*** Nowellia Mitt., Nat. hist. Azores: 321, 1870 (
** sect. Acronowellia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (
*** Nowellia reedii H.Rob., Bryologist 73 (1): 150, 1970 (
*** Nowellia yunckeri Fulford, Mem. New York Bot. Gard. 11 (3): 329, 1968 (
** sect. Metanowellia (Grolle) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (
*** Nowellia borneensis (De Not.) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
*** Nowellia dominicensis Steph., Sp. Hepat. (Stephani) 6: 443, 1924 (
*** Nowellia evansii Grolle, J. Hattori Bot. Lab. 31: 33, 1968 (
*** Nowellia langii Pearson, J. Linn. Soc., Bot. 46 (305): 25, 1922 (
*** Nowellia pusilla Grolle, J. Hattori Bot. Lab. 31: 37, 1968 (
*** Nowellia wrightii (Gottsche ex Spruce) Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 21, 1903 (
** sect. Nowellia
*** Nowellia aciliata (P.C.Chen et P.C.Wu) Mizut., Hikobia 11: 469, 1994 (
*** Nowellia curvifolia (Dicks.) Mitt., Nat. hist. Azores: 321, 1870 (
*** Odontoschisma (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** sect. Cladopodiella (H.Buch) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (
*** Odontoschisma francisci (Hook.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (
** sect. Denudata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 833, 1974 (
*** Odontoschisma brasiliense Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 585 (369), 1908 (
** Odontoschisma cleefii Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (
*** Odontoschisma denudatum (Mart.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** Odontoschisma denudatum subsp. naviculare (Steph.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (
** Odontoschisma denudatum subsp. sandvicense (Ångstr.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (
*** Odontoschisma elongatum (Lindb.) A.Evans, Rhodora 14 (157): 13, 1912 (
*** Odontoschisma engelii Gradst. et Burghardt, Fieldiana, Bot. (n.ser.) 47: 194, 2008 (
*** Odontoschisma longiflorum (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (
*** Odontoschisma macounii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 92, 1884 (
*** Odontoschisma portoricense (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 296, 1888 (
** Odontoschisma pseudogrosseverrucosum Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (
*** Odontoschisma purpuratum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 297, 1950 (
*** Odontoschisma soratamum Fulford, Mem. New York Bot. Gard. 11 (3): 338, 1968 (
*** Odontoschisma variabile (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (
** Odontoschisma zhui Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1020, 2014 (
** sect. Iwatsukia (N.Kitag.) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (
*** Odontoschisma bifidum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (
*** Odontoschisma jishibae (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 13, 2013 (
*** Odontoschisma spinosum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (
** sect. Neesia Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (
*** Odontoschisma fluitans (Nees) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (
** sect. Odontoschisma
*** Odontoschisma grosseverrucosum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 593 (377), 1908 (
*** Odontoschisma sphagni (Dicks.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (
** Odontoschisma steyermarkii Gradst. et Ilk.-Borg., Nova Hedwigia 100 (1/2): 38, 2015 [2014 online] (
Incertae sedis
* Odontoschisma obcordatum (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 586 (370), 1908 (
** Schiffneria Steph., Österr. Bot. Z. 44 (1): 1, 1894 (
*** Schiffneria hyalina Steph., Österr. Bot. Z. 44 (1): 1, 1894 (
** Haesselia Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (
*** Haesselia acuminata Gradst., Trop. Bryol. 1: 30, 1989 (
*** Haesselia roraimensis Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (
** Trabacellula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (
*** Trabacellula tumidula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (
by J. Váňa
The treatment of Cephaloziellaceae follows what was outlined in
*** Allisoniella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 80, 1965 (
** sect. Allisoniella
*** Allisoniella nigra (Rodway) R.M.Schust., Nova Hedwigia 22: 137, 1971 [1972] (
** Allisoniella nigra var. acutiloba J.J.Engel, Novon 17 (3): 313, 2007 (
** Allisoniella nigra subsp. novaezelandiae R.M.Schust., Nova Hedwigia 22: 143, 1971 [1972] (
*** Allisoniella recurva R.M.Schust., Nova Hedwigia 22: 146, 1971 [1972] (
*** Allisoniella subbipartita (C.Massal.) R.M.Schust. et J.J.Engel, Nova Hedwigia 22: 147, 1971 [1972] (
*** Allisoniella tasmanica R.M.Schust., Nova Hedwigia 22: 145, 1971 [1972] (
** sect. Protomarsupella (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 150, 1971 (
*** Allisoniella scottii (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 151, 1971 [1972] (
*** Amphicephalozia R.M.Schust., Nova Hedwigia 22: 131, 1971 [1972] (
*** Amphicephalozia africana Váňa et M.Wigginton, J. Bryol. 30 (1): 55, 2008 (
*** Amphicephalozia amplexicaulis R.M.Schust., Nova Hedwigia 22: 133, 1971 [1972] (
*** Amphicephalozia geisslerae Pócs et Váňa, Polish Bot. J. 46 (2): 145, 2001 (
*** Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (
*** Anastrophyllopsis involutifolia (Mont. ex Gottsche, Lindenb. et Nees) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (
*** Anastrophyllopsis revoluta (Steph.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (
*** Anastrophyllopsis subcomplicata (Lehm. et Lindenb.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (
** Cephalojonesia Grolle, Rev. Bryol. Lichénol. 37 (4): 763, 1970 [1971] (
*** Cephalojonesia incuba Grolle et Vanden Berghen, Rev. Bryol. Lichénol. 37 (4): 764, 1970 [1971] (
*** Cephalojonesia incuba subsp. mexicana Burghardt, Gradst. et Váňa, J. Hattori Bot. Lab. 100: 35, 2006 (
** Cephalomitrion R.M.Schust., Nova Hedwigia 61 (3/4): 550, 1995 (
*** Cephalomitrion aterrimum (Steph.) R.M.Schust., Nova Hedwigia 61 (3/4): 554, 1995 (
*** Cephaloziella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (
** subg. Cephaloziella
*** Cephaloziella aenigmatica R.M.Schust., Nova Hedwigia 63 (1/2): 20, 1996 (
*** Cephaloziella anthelioides S.W.Arnell, Bot. Not. 105: 322, 1952 (
*** Cephaloziella arctogena (R.M.Schust.) Konstant., Arctoa 3: 126, 1994 (
** Cephaloziella arenaria (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (
*** Cephaloziella aspericaulis Jørg., Bergens Mus. Skr. (n.ser.) 16: 197, 1934 (
*** Cephaloziella baumgartneri Schiffn., Verh. K.K. Zool.-Bot. Ges. Wien 56 (3): 273, 1906 (
** Cephaloziella breviperianthia C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 208, 1981 (
** Cephaloziella brinkmannii Douin, Mém. Soc. Bot. France 29: 75, 1920 (
*** Cephaloziella capensis (Sim) S.W.Arnell, Bot. Not. 105: 326, 1952 (
*** Cephaloziella capillaris (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (
*** Cephaloziella crassigyna (R.M.Schust.) R.M.Schust., Nova Hedwigia 61 (3/4): 556, 1995 (
*** Cephaloziella crispata N.Kitag., J. Hattori Bot. Lab. 32: 301, 1969 (
*** Cephaloziella densifolia R.M.Schust., Nova Hedwigia 22: 199, 1971 [1972] (
** Cephaloziella densifolia var. dubia R.M.Schust., Nova Hedwigia 22: 202, 1971 [1972] (
*** Cephaloziella divaricata (Sm.) Schiffn., Hepat. (Engl.-Prantl): 99, 1893 (
** Cephaloziella divaricata var. scabra (M.Howe) Haynes, Bryologist 12 (4): 68, 1909 (
** Cephaloziella dusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 49, 1900 (
*** Cephaloziella elachista (J.B.Jack ex Gottsche et Rabenh.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (
** Cephaloziella elachista var. spinophylla (C.Gao) C.Gao, Fl. Bryoph. Sin. 9: 181, 2003 (
** Cephaloziella elegans (Heeg) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (
*** Cephaloziella exigua R.M.Schust., Nova Hedwigia 63 (1/2): 49, 1996 (
*** Cephaloziella exiliflora (Taylor) Douin, Mém. Soc. Bot. France 29: 72, 1920 (
** Cephaloziella fragillima (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 409, 1976 (
*** Cephaloziella garsidei S.W.Arnell, Rev. Bryol. Lichénol. 23 (1/2): 173, 1954 (
*** Cephaloziella grandiretis (R.M.Schust.) R.M.Schust., Nova Hedwigia 63 (1/2): 35, 1996 (
*** Cephaloziella grimsulana (J.B.Jack ex Gottsche et Rabenh.) Lacout., Hépat. France: 52, 1905 (
*** Cephaloziella hampeana (Nees) Schiffn. ex Loeske, Moosfl. Harz.: 92, 1903 (
** Cephaloziella hebridensis Steph., Hedwigia 32 (5): 316, 1893 (
*** Cephaloziella herzogiana (Pandé et K.P.Srivast.) Udar et D.Kumar, Geophytology 6 (1): 45, 1976 (
** Cephaloziella heteroica (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 76, 1920 (
** Cephaloziella hyalina Douin, Mém. Soc. Bot. France 29: 77, 1920 (
* Cephaloziella hyalina var. rappii (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 56, 1980 (
*** Cephaloziella inaequalis R.M.Schust., Nova Hedwigia 22: 186, 1971 [1972] (
*** Cephaloziella invisa R.M.Schust., Nova Hedwigia 63 (1/2): 30, 1996 (
** Cephaloziella kilohanensis (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 85, 1920 (
*** Cephaloziella longii Váňa, Folia Geobot. Phytotax. 27 (2): 193, 1992 (
*** Cephaloziella lycopodioides (Sim) S.W.Arnell, Bot. Not. 105: 321, 1952 (
** Cephaloziella mammillifera R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (
*** Cephaloziella massalongi (Spruce) Müll.Frib., Lebermoose 2 (17): 191, 1913 (
*** Cephaloziella muelleriana R.M.Schust., Nova Hedwigia 63 (1/2): 24, 1996 (
*** Cephaloziella natalensis (Sim) S.W.Arnell, Hepat. South Africa: 338, 1963 (
** Cephaloziella nicholsonii Douin, Rev. Bryol. 40 (6): 81, 1913 (
*** Cephaloziella nothogena R.M.Schust., Nova Hedwigia 63 (1/2): 37, 1996 (
** Cephaloziella obtusilobula R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 108, 1980 (
** Cephaloziella patulifolia (Steph.) Douin, Mém. Soc. Bot. France 29: 70, 1920 (
*** Cephaloziella phyllacantha (C.Massal. et Carestia) Müll.Frib., Lebermoose 2 (17): 194, 1913 (
*** Cephaloziella polystratosa (R.M.Schust. et Damsh.) Konstant., Bot. Zhurn. (Moscow & Leningrad) 85 (10): 127, 2000 (
*** Cephaloziella pseudocrassigyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (
*** Cephaloziella pulcherrima R.M.Schust., Nova Hedwigia 22: 203, 1971 [1972] (
** Cephaloziella pulcherrima subsp. sphagnicola R.M.Schust., Nova Hedwigia 22: 205, 1971 [1972] (
** Cephaloziella pungens Steph. ex Fulford, Mem. New York Bot. Gard. 11 (4): 410, 1976 (
*** Cephaloziella rubella (Nees) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 231, 1902 (
*** Cephaloziella schelpei S.W.Arnell, Bot. Not. 110 (1): 18, 1957 (
*** Cephaloziella spegazziniana (C.Massal.) Douin, Mém. Soc. Bot. France 29: 69, 1920 (
*** Cephaloziella spinicaulis Douin, Mém. Soc. Bot. France 29: 62, 1920 (
*** Cephaloziella spinigera (Lindb.) Jørg., Bergens Mus. Skr. (n.ser.) 16: 189, 1934 (
*** Cephaloziella stellulifera (Taylor ex Carrington et Pearson) Croz., Rev. Bryol. 30 (2): 31, 1903 (
*** Cephaloziella stephanii Schiffn. ex Douin, Mém. Soc. Bot. France 29: 85, 1920 (
** Cephaloziella stolonifera R.M.Schust., Phytologia 39 (6): 426, 1978 (
** Cephaloziella subtilis (Lindenb. et Gottsche) Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephaloziella sumatrana Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (
*** Cephaloziella tabularis S.W.Arnell, Bot. Not. 105: 318, 1952 (
*** Cephaloziella transvaalensis S.W.Arnell, Bot. Not. 110 (1): 19, 1957 (
*** Cephaloziella triplicata S.W.Arnell, Bot. Not. 115: 203, 1962 (
*** Cephaloziella umtaliensis S.W.Arnell, Bot. Not. 110 (1): 20, 1957 (
*** Cephaloziella uncinata R.M.Schust., Meddel. Grønland 199 (1): 316, 1974 (
** Cephaloziella uncinata var. brevigyna R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (
** Cephaloziella uncinata var. sphagnicola R.M.Schust., Meddel. Grønland 199 (1): 323, 1974 (
*** Cephaloziella vaginans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 119, 1911 (
*** Cephaloziella varians (Gottsche) Steph., Wiss. Ergebn. Schwed. Südpolar-Exped. [1901–1903] 4 (1): 4, 1905 (
*** Cephaloziella verrucosa Steph., Hedwigia 32 (5): 318, 1893 (
* Cephaloziella villaumei (Steph.) Váňa, Phytotaxa 112 (1): 3, 2013 (
** Cephaloziella violacea Schljakov, Novosti Sist. Nizš. Rast. 15: 242, 1978 (
** Cephaloziella welwitschii (Steph.) Douin, Mém. Soc. Bot. France 29: 58, 1920 (
** subg. Dichiton (Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (
*** Cephaloziella calyculata (Durieu et Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (
*** Cephaloziella integerrima (Lindb.) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 232, 1902 (
** subg. Evansia (Douin et Schiffn.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (
*** Cephaloziella antillana (Besch. et Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 405, 1976 (
*** Cephaloziella dentata (Raddi) Steph., Bull. Herb. Boissier 5 (2): 78, 1897 (
*** Cephaloziella hirta (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (
** Cephaloziella squarrosula (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephaloziella subspinosa R.M.Schust., Nova Hedwigia 22: 210, 1971 [1972] (
** subg. Prionolobus (Spruce) Müll.Frib., Lebermoose 2 (16): 110, 1912 (
*** Cephaloziella acanthophora (S.Hatt.) Horik., Hikobia 1 (2): 79, 1951 (
*** Cephaloziella biokoensis Váňa et Frank Müll., Trop. Bryol. 24: 1, 2003 (
*** Cephaloziella granatensis (J.B.Jack ex Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 411, 1976 (
*** Cephaloziella grisea R.M.Schust., Phytologia 39 (6): 425, 1978 (
* Cephaloziella meghalayensis Udar et Ad.Kumar, Lindbergia 8 (1): 34, 1982 (
*** Cephaloziella microphylla (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (
*** Cephaloziella tenuissima (Lehm.) Steph., Hedwigia 32 (5): 318, 1893 (
*** Cephaloziella turneri (Hook.) Müll.Frib., Lebermoose 2 (17): 202, 1913 (
Incertae sedis
** Cephaloziella dentifolia Udar et Ad.Kumar, Lindbergia 8 (1): 30, 1982 (
** Cephaloziella filum (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (
* Cephaloziella flexuosa C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 88, 1984 (
** Cephaloziella intricata Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (
** Cephaloziella levieri Schiffn. ex Douin, Mém. Soc. Bot. France 29: 80, 1920 (
*** Cephaloziella pellucida R.M.Schust., Nova Hedwigia 63 (1/2): 57, 1996 (
* Cephaloziella pygmaea (Spruce) Váňa, Phytotaxa 183 (4): 290, 2014 (
* Cephaloziella secundifolia Pearson, Ann. Bryol. 4: 106, 1931 (
** Cephaloziella sinensis Douin, Rev. Bryol. 41 (1): 8, 1914 (
** Cephaloziopsis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (
*** Cephaloziopsis intertexta (Gottsche) R.M.Schust., Nova Hedwigia 22: 183, 1971 [1972] (
** Chaetophyllopsis R.M.Schust., J. Hattori Bot. Lab. 23: 69, 1960 [1961] (
*** Chaetophyllopsis whiteleggei (Carrington et Pearson) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 255, 1972 (
** Cylindrocolea R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
** subg. Cylindrocolea
*** Cylindrocolea kiaeri (Austin) Váňa, Phytotaxa 112 (1): 2, 2013 (
*** Cylindrocolea sanctae-helenae M.Wigginton, Polish Bot. J. 58 (1): 107, 2013 (
** sect. Cylindrocolea
*** Cylindrocolea andersonii R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 33, 1980 (
*** Cylindrocolea brasiliensis D.P.Costa, N.D.Santos et Váňa, Bryologist 111 (4): 667, 2008 (
*** Cylindrocolea chevalieri (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
*** Cylindrocolea gittinsii (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (
*** Cylindrocolea madagascariensis (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (
*** Cylindrocolea nigerica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (
*** Cylindrocolea novae-caledoniae (Grolle) R.M.Schust., Nova Hedwigia 22: 161, 1971 [1972] (
*** Cylindrocolea planifolia (Steph.) R.M.Schust., Nova Hedwigia 22: 164, 1971 [1972] (
*** Cylindrocolea sprucei R.M.Schust., Nova Hedwigia 22: 163, 1971 [1972] (
*** Cylindrocolea ugandica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 171, 1971 [1972] (
** sect. Platycaulis R.M.Schust., Nova Hedwigia 22: 173, 1971 (
*** Cylindrocolea recurvifolia (Steph.) Inoue, J. Jap. Bot. 47 (11): 348, 1972 (
*** Cylindrocolea tagawae (N.Kitag.) R.M.Schust., Nova Hedwigia 22: 174, 1971 [1972] (
** subg. Cylindroscyphus (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 20, 1980 (
*** Cylindrocolea abyssinica (Gola) Váňa, Phytotaxa 112 (1): 2, 2013 (
** Cylindrocolea obtusifolia Fulford, Mem. New York Bot. Gard. 11 (4): 401, 1976 (
** Cylindrocolea reticulata Udar et Ad.Kumar, Lindbergia 8 (3): 181, 1982 [1983] (
*** Cylindrocolea rhizantha (Mont.) R.M.Schust., Nova Hedwigia 22: 175, 1971 [1972] (
*** Gottschelia Grolle, J. Hattori Bot. Lab. 31: 13, 1968 (
*** Gottschelia maxima (Steph.) Grolle, J. Bryol. 25 (1): 6, 2003 (
*** Gottschelia schizopleura (Spruce) Grolle, J. Hattori Bot. Lab. 31: 16, 1968 (
** Gymnocoleopsis (R.M.Schust.) R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Gymnocoleopsis capensis (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 78: 123, 1995 (
*** Gymnocoleopsis cylindriformis (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 78: 126, 1995 (
*** Herzogobryum Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (
*** Herzogobryum atrocapillum (Hook.f. et Taylor) Grolle, Österr. Bot. Z. 113 (2): 228, 1966 (
*** Herzogobryum filiforme R.M.Schust., Phytologia 45 (5): 422, 1980 (
*** Herzogobryum molle Grolle, Österr. Bot. Z. 113 (2): 226, 1966 (
*** Herzogobryum vermiculare (Schiffn.) Grolle, J. Hattori Bot. Lab. 28: 103, 1965 (
** Kymatocalyx Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 56, 1950 (
*** Kymatocalyx africanus Váňa et M.Wigginton, Haussknechtia, Beih. 9: 158, 1999 (
*** Kymatocalyx dominicensis (Spruce) Váňa, Österr. Bot. Z. 118 (5): 575, 1970 (
*** Kymatocalyx madagascariensis (Steph.) Gradst. et Váňa, Haussknechtia, Beih. 9: 164, 1999 (
*** Kymatocalyx rhizomaticus (Herzog) Gradst. et Váňa, Haussknechtia, Beih. 9: 166, 1999 (
*** Lophonardia R.M.Schust., Phytologia 39 (4): 244, 1978 (
*** Lophonardia jamesonii (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 19, 2013 (
*** Lophonardia laxifolia (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (
*** Lophonardia tristaniana (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (
*** Nothogymnomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 43, 1996 (
*** Nothogymnomitrion erosum (Carrington et Pearson) R.M.Schust., J. Hattori Bot. Lab. 80: 44, 1996 (
*** Obtusifolium S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 309, 1956 (
*** Obtusifolium obtusum (Lindb.) S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 133, 1956 (
*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (
*** Oleolophozia perssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (
** Phycolepidozia R.M.Schust., Bull. Torrey Bot. Club 93 (6): 438, 1966 [1967] (
** subg. Metaphycolepidozia Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 506, 2014 (
** Phycolepidozia indica Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 499, 2014 (
** subg. Phycolepidozia
*** Phycolepidozia exigua R.M.Schust., Bull. Torrey Bot. Club 93 (6): 440, 1966 [1967] (
** Protolophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (
** Protolophozia androgyna R.M.Schust. ex Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
** Protolophozia autoica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
** Protolophozia crispata (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia druceae (Grolle et E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia elongata (Steph.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (
*** Protolophozia herzogiana (E.A.Hodgs. et Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia lancistipa (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia leucorhiza (Mitt.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (
*** Protolophozia longiflora (Herzog) L.Söderstr. et Váňa, Phytotaxa 76 (3): 51, 2013 (
** Protolophozia monoica (E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia multicuspidata (Hook.f. et Taylor) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
** Protolophozia nivicola (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia perssoniana (H.A.Mill.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
** Protolophozia subalpina (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia tasmanica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
*** Protolophozia verruculosa (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (
by L. Söderström and J. Váňa
Lophoziaceae was broadly defined by
*** Andrewsianthus R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 66, 1961 (
** subg. Andrewsianthus
*** Andrewsianthus aberrans (Nees et Mont.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 440, 1963 (
*** Andrewsianthus bidens (Mitt. ex Steph.) R.M.Schust., Nova Hedwigia 15: 492, 1968 (
*** Andrewsianthus bilobus (Mitt.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 437, 1963 (
*** Andrewsianthus carinatus Grolle, Marion Prince Edw. Is: 229, 1971 (
*** Andrewsianthus cavifolius Grolle et Váňa, J. Hattori Bot. Lab. 38: 640, 1974 (
*** Andrewsianthus chimbuensis R.M.Schust., Nova Hedwigia 15: 491, 1968 (
*** Andrewsianthus kinabaluensis N.Kitag., J. Hattori Bot. Lab. 33: 205, 1970 (
*** Andrewsianthus koponenii Váňa et Piippo, Ann. Bot. Fenn. 26 (3): 284, 1989 (
*** Andrewsianthus mizutanii N.Kitag., J. Hattori Bot. Lab. 32: 307, 1969 (
*** Andrewsianthus papillosus N.Kitag., J. Hattori Bot. Lab. 33: 207, 1970 (
*** Andrewsianthus perigonialis (Hook.f. et Taylor) R.M.Schust., Beih. Nova Hedwigia 119: 336, 2002 (
*** Andrewsianthus puniceus (Nees) R.M.Schust. ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 439, 1963 (
*** Andrewsianthus recurvifolius (Nees) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (
*** Andrewsianthus sundaicus (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (
*** Andrewsianthus zantenii Váňa, J. Hattori Bot. Lab. 38: 645, 1974 (
** subg. Cephalolobus (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 326, 2002 (
*** Andrewsianthus hodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
*** Andrewsianthus marionensis (S.W.Arnell) Grolle, Marion Prince Edw. Is: 232, 1971 (
*** Andrewsianthus scabrellus (C.Massal.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
** Andrewsianthus sphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
** Gerhildiella Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (
*** Gerhildiella rossneriana Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (
*** Heterogemma (Jørg.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Heterogemma capitata (Hook.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Heterogemma laxa (Lindb.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
*** Heterogemma patagonica (Herzog et Grolle) L.Söderstr. et Váňa, Phytotaxa 97 (2): 29, 2013 (
*** Lophozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
*** Lophozia ascendens (Warnst.) R.M.Schust., Bryologist 55 (3): 180, 1952 (
* Lophozia austrosibirica Bakalin, Ann. Bot. Fenn. 40 (1): 49, 2003 (
** Lophozia ciliata Damsh., L.Söderstr. et H.Weibull, Lindbergia 25 (1): 3, 2000 (
*** Lophozia guttulata (Lindb. et Arnell) A.Evans, Proc. Wash. Acad. Sci. 2: 302, 1900 (
* Lophozia jamaicensis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 128 (155), 1901 (
** Lophozia lacerata N.Kitag., Hikobia 3 (3): 172, 1963 (
** Lophozia lantratoviae Bakalin, Ann. Bot. Fenn. 40 (1): 47, 2003 (
** Lophozia murmanica Kaal., Rep. Second Norweg. Arctic Exped. 11: 34, 1906 (
** Lophozia pacifica Bakalin, Bryologist 114 (2): 302, 2011 (
* Lophozia pallida (Steph.) Grolle, J. Jap. Bot. 39 (6): 174, 1964 (
** Lophozia savicziae Schljakov, Novosti Sist. Nizš. Rast. 10: 299, 1973 (
* Lophozia schusterana Schljakov, Novosti Sist. Nizš. Rast. 12: 320, 1975 (
*** Lophozia silvicola H.Buch, Beret. 18 Skand. Naturforskarmøde: 228, 1929 (
*** Lophozia silvicoloides N.Kitag., J. Hattori Bot. Lab. 28: 276, 1965 (
* Lophozia subapiculata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 104, 1974 (
** Lophozia udarii S.Srivast., S.C.Srivast. et K.K.Rawat, Nelumbo 55: 130, 2013 (
*** Lophozia ventricosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
* Lophozia wenzelii (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 35 (135), 1902 (
Excluded from the genus
* Lophozia serpens (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (
** Lophoziopsis Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Lophoziopsis excisa (Dicks.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis excisa var. elegans (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis excisa var. infuscata (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis excisa var. succulenta (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Lophoziopsis longidens (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Lophoziopsis longidens subsp. arctica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 97 (2): 28, 2013 (
*** Lophoziopsis pellucida (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Lophoziopsis pellucida var. minor (R.M.Schust.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 28, 2013 (
*** Lophoziopsis polaris (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Lophoziopsis polaris var. sphagnorum (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
* Lophoziopsis propagulifera (Gottsche) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Lophoziopsis rubrigemma (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (
** Pseudocephaloziella R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Pseudocephaloziella epiphytica R.M.Schust., Phytologia 39 (4): 243, 1978 (
*** Trilophozia (R.M.Schust.) Bakalin, Monogr. Lophozia: 34, 2005 (
*** Trilophozia quinquedentata (Huds.) Bakalin, Monogr. Lophozia: 34, 2005 (
** Trilophozia quinquedentata var. asymmetrica (Horik.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 32, 2013 (
*** Tritomaria Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (
*** Tritomaria exsecta (Schmidel) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (
** Tritomaria exsecta subsp. novaezelandiae J.J.Engel, Bryologist 109 (1): 61, 2006 (
*** Tritomaria exsectiformis (Breidl.) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (
** Tritomaria exsectiformis subsp. arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 661, 1969 (
** Tritomaria exsectiformis subsp. camerunensis S.W.Arnell ex Váňa, Phytotaxa 167 (2): 216, 2014 (
*** Tritomaria ferruginea (Grolle) Váňa, Phytotaxa 81 (1): 24, 2013 (
* Tritomaria koreana Bakalin, S.S.Choi et B.Y.Sun, Arctoa 18: 163, 2009 [2010] (
* Tritomaria mexicana Bakalin, Arctoa 17: 162, 2008 [2009] (
*** Tritomaria scitula (Taylor) Jørg., Bergens Mus. Aarbok 1919/20 (7): 3, 1922 (
by J. Váňa
Scapaniaceae, in its classical concept, was recognized as monophyletic and nested within Lophoziaceae s. lat. (cf.
*** Diplophyllum (Dumort.) Dumort., Recueil Observ. Jungerm.: 15, 1835 (
*** subg. Austrodiplophyllum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 18, 1968 (
** Diplophyllum recurvifolium C.Massal., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 221, 1928 (
*** Diplophyllum squarrosum Steph., Sp. Hepat. (Stephani) 4: 116, 1910 (
*** Diplophyllum verrucosum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 19, 1968 (
*** subg. Diplophyllum
*** sect. Diplophyllum
*** Diplophyllum albicans (L.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (
*** sect. Protodiplophyllum (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 29, 2013 (
*** Diplophyllum africanum S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 527, 1956 (
*** Diplophyllum andicola R.M.Schust., Phytologia 39 (4): 248, 1978 (
*** Diplophyllum andrewsii A.Evans, Bryologist 25 (2): 28, 1922 (
* Diplophyllum androgynum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 277, 1998 (
* Diplophyllum angustifolium J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 262, 1998 (
*** Diplophyllum apiculatum (A.Evans) Steph., Sp. Hepat. (Stephani) 4: 110, 1910 (
** Diplophyllum dioicum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 20, 1968 (
** Diplophyllum exiguum Steph., Sp. Hepat. (Stephani) 6: 500, 1924 (
* Diplophyllum gemmiparum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 255, 1998 (
* Diplophyllum incrassatum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 265, 1998 (
*** Diplophyllum nanum Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 48, 1950 [1951] (
* Diplophyllum novum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 274, 1998 (
*** Diplophyllum obtusatum (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 215, 1974 (
*** Diplophyllum obtusifolium (Hook.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (
** Diplophyllum obtusifolium subsp. domesticum (Gottsche) Váňa, Phytotaxa 76 (3): 29, 2013 (
*** Diplophyllum serrulatum (Müll.Frib.) Steph., Sp. Hepat. (Stephani) 4: 112, 1910 (
*** Diplophyllum taxifolium (Wahlenb.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (
** Diplophyllum taxifolium var. mucronatum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 203, 1974 (
*** Diplophyllum trollii Grolle, Khumbu Himal 1 (4): 273, 1966 (
*** Douinia (C.E.O.Jensen) H.Buch, Scapan. N.-Eur. Sib.: 13, 1928 (
*** Douinia imbricata (M.Howe) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (
*** Douinia ovata (Dicks.) H.Buch, Scapan. N.-Eur. Sib.: 14, 1928 (
*** Douinia plicata (Lindb.) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (
** Pseudotritomaria Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
*** Pseudotritomaria heterophylla (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (
** Saccobasis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 291, 1932 [1933] (
*** Saccobasis polita (Nees) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 292, 1932 [1933] (
** Saccobasis polymorpha (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (
*** Scapania (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** subg. Ascapania Grolle, Khumbu Himal 1 (4): 268, 1966 (
*** Scapania contorta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (
** subg. Gracilidae (H.Buch) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (
*** Scapania ampliata Steph., Bull. Herb. Boissier 5 (2): 106, 1897 (
** Scapania ampliata subsp. queenslandica M.L.Hicks, J. Hattori Bot. Lab. 69: 130, 1991 (
*** Scapania bolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 218, 1869 (
*** Scapania gracilis Lindb., Morgonbladet (Helsinki) 1873 (286, 9 Dec): 2, 1873 (
** Scapania macroparaphyllia T.Cao, C.Gao et J.Sun, Acta Phytotax. Sin. 42 (2): 180, 2004 (
*** Scapania maxima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 223, 1934 (
*** Scapania nipponica (Amakawa et S.Hatt.) Amakawa, J. Hattori Bot. Lab. 30: 319, 1967 (
** Scapania paraphyllia T.Cao, C.Gao, J.Sun et B.R.Zuo, Acta Phytotax. Sin. 45 (3): 311, 2007 (
*** Scapania subnimbosa Steph., Sp. Hepat. (Stephani) 4: 150, 1910 (
** subg. Macroscapania R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 248, 1974 (
* Scapania geppii Steph., Hedwigia 44 (1): 14, 1904 (
*** Scapania portoricensis Hampe et Gottsche, Linnaea 25 (3): 342, 1852 [1853] (
** Scapania portoricensis var. boliviensis (Steph.) Herzog, Ann. Bryol. 1: 110, 1928 (
** Scapania portoricensis var. organensis (Herzog) Herzog, Ann. Bryol. 1: 110, 1928 (
** Scapania portoricensis var. roraimensis Warnst., Hedwigia 63 (2): 109, 1921 (
** subg. Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903 (
** sect. Planifoliae (Müll.Frib.) Potemkin, J. Hattori Bot. Lab. 85: 56, 1998 (
*** Scapania davidii Potemkin, Ann. Bot. Fenn. 38 (2): 83, 2001 (
** Scapania ferruginaeoides T.Cao, C.Gao et J.Sun, Guihaia 24 (1): 23, 2004 (
** Scapania gaochii X.Fu ex T.Cao, Phytotaxa 97 (2): 26, 2013 (
*** Scapania harae Amakawa, J. Hattori Bot. Lab. 27: 5, 1964 (
*** Scapania nimbosa Taylor, Nov. Stirp. Pug. 8: 6, 1844 (
*** Scapania ornithopodioides (With.) Waddell, Moss Exch. Club Cat. Hepat.: 4, 1897 (
*** Scapania rotundifolia W.E.Nicholson, Symb. Sin. 5: 31, 1930 (
*** Scapania secunda Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 226, 1894 (
*** Scapania zhukovae Potemkin, Arctoa 9: 121, 2000 (
** sect. Plicaticalyx (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 326, 2002 (
*** Scapania ciliatospinosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 222, 1934 (
*** Scapania ferruginea (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 1: 72, 1844 (
*** Scapania hians Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 223, 1905 (
** Scapania hians subsp. salishensis J.D.Godfrey et G.Godfrey, Bryologist 81 (3): 362, 1978 (
*** Scapania orientalis Steph. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 606, 1901 (
*** Scapania pseudocontorta Potemkin, Arctoa 9: 115, 2000 (
*** Scapania sinikkae Potemkin, Ann. Bot. Fenn. 38 (2): 85, 2001 (
*** Scapania spiniloba Potemkin, Arctoa 9: 117, 2000 (
** subg. Pseudomacrodiplophyllum Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (
*** Scapania microdonta (Mitt.) Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 262, 1905 (
*** subg. Scapania
** sect. Aequilobae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 110, 1928 (
*** Scapania aequiloba (Schwägr.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Scapania aspera M.Bernet et Bernet, Cat. hép. Suisse: 42, 1888 (
** sect. Americanae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (
*** Scapania americana Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 44, 1903 (
** sect. Apiculatae H.Buch, Scapan. N.-Eur. Sib.: 53, 1928 (
*** Scapania apiculata Spruce, Hep. Pyr. Exsic.: no. 15, 1847 (
*** Scapania carinthiaca J.B.Jack ex Lindb., Rev. Bryol. 7 (4): 77, 1880 (
*** Scapania carinthiaca var. massalongi Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 598, 1901 (
*** Scapania umbrosa (Schrad.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** sect. Ciliatae Grolle, Khumbu Himal 1 (4): 272, 1966 (
*** Scapania bhutanensis Amakawa, Fl. E. Himalaya 2: 230, 1971 (
*** Scapania ciliata Sande Lac., Prolus. fl. jap.: 209, 1867 (
*** Scapania ciliata subsp. hawaiica (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 321, 2002 (
*** Scapania hirosakiensis Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 120, 1905 (
*** Scapania hollandiae W.S.Hong, Bryologist 83 (1): 56, 1980 (
*** Scapania koponenii Potemkin, Ann. Bot. Fenn. 37 (1): 41, 2000 (
*** Scapania lepida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (
*** Scapania sandei Schiffn. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 612, 1901 (
** sect. Compactae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 101, 1928 (
*** Scapania compacta (Roth) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Scapania kaurinii Ryan, Bot. Not. 42: 210, 1889 (
*** Scapania spitsbergensis (Lindb.) Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 607, 1901 (
** sect. Curtae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 55, 1928 (
*** Scapania curta (Mart.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** Scapania curta var. grandiretis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 393, 1974 (
** Scapania curta var. isoloba R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 396, 1974 (
*** Scapania diplophylloides Amakawa et S.Hatt., J. Hattori Bot. Lab. 9: 59, 1953 (
*** Scapania esterhuyseniae S.W.Arnell, Bot. Not. 110 (1): 26, 1957 (
*** Scapania fulfordiae W.S.Hong, Bryologist 83 (1): 46, 1980 (
*** Scapania gamundiae R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 14, 1968 (
*** Scapania helvetica Gottsche, Hepat. Eur., Leberm. 42-44: no. 426, 1868 (
*** Scapania irrigua (Nees) Nees, Syn. Hepat. 1: 67, 1844 (
** Scapania irrigua subsp. rufescens (Loeske) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 471, 1974 (
*** Scapania lingulata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 92, 1916 (
** Scapania lingulata var. microphylla (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 415, 1974 (
** Scapania magadanica S.S.Choi, Bakalin et B.Y.Sun, Bot. Pacifica 1: 46, 2012 (
*** Scapania mucronata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 91, 1916 (
*** Scapania obcordata (Berggr.) S.W.Arnell, Ark. Bot. (n.ser.) 4 (6): 117, 1959 (
** Scapania parvifolia Warnst., Hedwigia 63 (2): 78, 1921 (
*** Scapania praetervisa Meyl., Jahresber. Naturf. Ges. Graubündens (n.f.) 64: 364, 1926 (
*** Scapania scandica (Arnell et H.Buch) Macvicar, Stud. handb. Brit. hepat. (ed. 2): 394, 1926 (
** Scapania scandica var. argutedentata H.Buch, Scapan. N.-Eur. Sib.: 75, 1928 (
** Scapania scandica var. dimorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 453, 1974 (
** Scapania scandica var. grandiretis (Schljakov) Schljakov, Pečen. Mchi Sev. SSSR 4: 152, 1981 (
*** Scapania uliginosa (Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
*** Scapania valdonii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 126, 2009 (
*** Scapania zemliae S.W.Arnell, Svensk Bot. Tidskr. 41: 215, 1947 (
** sect. Cuspiduligerae H.Buch, Scapan. N.-Eur. Sib.: 125, 1928 (
*** Scapania cuspiduligera (Nees) Müll.Frib., Lebermoose 2 (22): 472, 1915 (
** Scapania cuspiduligera var. diplophyllopsis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 361, 1974 (
** sect. Grolleoscapania Potemkin, Ann. Bot. Fenn. 39 (4): 329, 2002 (
*** Scapania karl-muelleri Grolle, Khumbu Himal 1 (4): 270, 1966 (
** sect. Hyperboreae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (
*** Scapania hyperborea Jørg., Forh. Vidensk.-Selsk. Kristiania 1894 (8): 56, 1894 (
*** Scapania paludicola Loeske et Müll.Frib., Lebermoose 2 (21): 425, 1915 (
** Scapania paludicola var. viridigemma R.M.Schust., Bull. Natl. Mus. Canada 122: 20, 1950 [1951] (
*** Scapania tundrae (Arnell) H.Buch, Scapan. N.-Eur. Sib.: 99, 1928 (
** sect. Kaalaasia (H.Buch) Jørg., Bergens Mus. Skr. (n.ser.) 16: 210, 1934 (
*** Scapania calcicola (Arnell et J.Perss.) Ingham, Naturalist (Hull) 564: 11, 1904 (
*** Scapania gymnostomophila Kaal., Bot. Not. 49: 21, 1896 (
*** Scapania ligulifolia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 306, 1974 (
*** Scapania pseudocalcicola R.M.Schust., Phytologia 63 (5): 327, 1987 (
* Scapania sect. Muelleria Potemkin, Ann. Bot. Fenn. 39 (4): 320, 2002 (
*** Scapania himalayica Müll.Frib. ex Herzog, Ann. Bryol. 12: 81, 1939 (
*** Scapania schljakovii Potemkin, Ann. Bot. Fenn. 38 (2): 87, 2001 (
** sect. Nemorosae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 152, 1928 (
*** Scapania brevicaulis Taylor, London J. Bot. 5: 272, 1846 (
*** Scapania crassiretis Bryhn, Rev. Bryol. 19 (1): 7, 1892 (
* Scapania degenii Schiffn. ex Müll.Frib., Lebermoose 2 (22): 497, 1915 (
* Scapania glaucoviridis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 221, 1934 (
** Scapania grossidens Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 146, 1905 (
*** Scapania hedbergii S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 556, 1956 (
*** Scapania integerrima Steph., Sp. Hepat. (Stephani) 4: 148, 1910 (
*** Scapania matveyevae Potemkin, Arctoa 9: 97, 2000 (
*** Scapania nemorea (L.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (
** Scapania parvidens Steph., Hedwigia 44 (1): 15, 1904 (
*** Scapania parvitexta Steph., Bull. Herb. Boissier 5 (2): 107, 1897 (
*** Scapania rigida Nees, Syn. Hepat. 1: 69, 1844 (
** sect. Scapania
*** Scapania gigantea Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 15, 1931 (
** Scapania grandiloba Steph., Sp. Hepat. (Stephani) 6: 502, 1924 (
*** Scapania komagadakensis Amakawa, J. Hattori Bot. Lab. 31: 96, 1968 (
*** Scapania obscura (Arnell et C.E.O.Jensen) Schiffn., Österr. Bot. Z. 58 (10): 377, 1908 (
** Scapania paludosa (Müll.Frib.) Müll.Frib., Mitt. Bad. Bot. Vereins 4 (182/183): 287, 1902 (
*** Scapania rufidula Warnst., Hedwigia 63 (2): 94, 1921 (
*** Scapania serrulata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 539, 1974 (
*** Scapania subalpina (Nees ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** Scapania subalpina var. haynesiae Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (4): 638, 1946 (
** Scapania subalpina var. muddiae C.D.Bird et W.S.Hong, Bryologist 83 (1): 51, 1980 (
*** Scapania undulata (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (
** sect. Scapaniella (H.Buch) Potemkin, J. Hattori Bot. Lab. 85: 43, 1998 (
*** Scapania glaucocephala (Taylor) Austin, Bull. Torrey Bot. Club 6 (16): 85, 1876 (
*** Scapania glaucocephala var. saxicola (R.M.Schust.) Potemkin, Bryologist 102 (1): 36, 1999 (
* Scapania scapanioides (C.Massal.) Grolle, Feddes Repert. 87 (3/4): 235, 1976 (
** sect. Simmonsiae (R.M.Schust.) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (
*** Scapania simmonsii Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 51, 1906 (
** sect. Sphaeriferae Konstant. et Potemkin, Ann. Bot. Fenn. 31 (2): 125, 1994 (
*** Scapania sphaerifera H.Buch et Tuom., Memoranda Soc. Fauna Fl. Fennica 11: 227, 1936 (
** sect. Stephaniae Potemkin, J. Hattori Bot. Lab. 85: 57, 1998 (
*** Scapania griffithii Schiffn., Österr. Bot. Z. 49 (6): 204, 1899 (
*** Scapania javanica Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 575, 1853 (
* Scapania javanica var. scabra Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955 (
*** Scapania ligulata Steph., Hedwigia 44 (1): 14, 1904 (
** Scapania ligulata subsp. stephanii (Müll.Frib.) Potemkin, Piippo et T.J.Kop., Ann. Bot. Fenn. 41 (6): 423, 2004 (
** sect. Verrucosae Potemkin, J. Hattori Bot. Lab. 85: 54, 1998 (
*** Scapania udarii S.C.Srivast. et A.Srivast., J. Indian Bot. Soc. 72: 237, 1993 (
*** Scapania verrucosa Heeg, Rev. Bryol. 20 (5): 81, 1893 (
*** Schistochilopsis (N.Kitag.) Konstant., Arctoa 3: 125, 1994 (
*** Schistochilopsis cornuta (Steph.) Konstant., Arctoa 3: 125, 1994 (
*** Schistochilopsis grandiretis (Lindb. ex Kaal.) Konstant., Arctoa 3: 125, 1994 (
** Schistochilopsis hyperarctica Konstant. et L.Söderstr., Phytotaxa 162 (4): 240, 2014 (
*** Schistochilopsis incisa (Schrad.) Konstant., Arctoa 3: 125, 1994 (
* Schistochilopsis nakanishii (Inoue) Konstant., Arctoa 3: 125, 1994 (
** Schistochilopsis opacifolia (Culm. ex Meyl.) Konstant., Arctoa 3: 125, 1994 (
*** Schistochilopsis setosa (Mitt.) Konstant., Arctoa 3: 125, 1994 (
by L. Briscoe and J.J. Engel
** Enigmella G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (
** Enigmella thallina G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (
*** Acrobolbus Nees, Syn. Hepat. 1: 5, 1844 (
** Acrobolbus africanus (Pearson) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus anisodontus (Hook.f. et Taylor) Briscoe, Phytotaxa 202 (1): 59, 2015 (
** Acrobolbus antillanus R.M.Schust., J. Hattori Bot. Lab. 90: 143, 2001 (
** Acrobolbus azoricus (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus caducifolius R.M.Schust., J. Hattori Bot. Lab. 90: 154, 2001 (
*** Acrobolbus ciliatus (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 86, 1893 (
*** Acrobolbus cinerascens (Lehm. et Lindenb.) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 242, 1888 (
*** Acrobolbus concinnus (Mitt.) Steph., Trans. & Proc. New Zealand Inst. 24: 399, 1892 (
** Acrobolbus cuneifolius (Steph.) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus diversifolius R.M.Schust., J. Hattori Bot. Lab. 90: 150, 2001 (
*** Acrobolbus epiphytus (Colenso) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus flavicans (J.J.Engel et Grolle) Briscoe et J.J.Engel, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus gradsteinii (Grolle) Briscoe, Phytotaxa 202 (1): 59, 2015 (
** Acrobolbus integrifolius (A.Evans) Briscoe, Phytotaxa 202 (1): 59, 2015 (
*** Acrobolbus knightii (Mitt.) Briscoe, Phytotaxa 202 (1): 59, 2015 (
** Acrobolbus kunkelii (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus laxus (Lehm. et Lindenb.) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus limbatus (Steph.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus lophocoleoides (Mitt.) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
** Acrobolbus madeirensis (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 60, 2015 (
** Acrobolbus mittenii Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902 (
*** Acrobolbus ochrophyllus (Hook.f. et Taylor) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 64, 1961 (
*** Acrobolbus papillosus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus perpusillus (Colenso) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus perpusillus var. denticulatus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus plagiochiloides (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus pseudosaccatus (Grolle) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus renifolius (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (
** Acrobolbus ruwenzorensis (S.W.Arnell) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus saccatus (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus setulosus (Mitt.) Briscoe, Phytotaxa 202 (1): 60, 2015 (
*** Acrobolbus spinifolius R.M.Schust., J. Hattori Bot. Lab. 90: 137, 2001 (
*** Acrobolbus sumatranus (Schiffn.) Briscoe, Phytotaxa 202 (1): 61, 2015 (
*** Acrobolbus surculosus (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus tenellus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus tenellus var. diversifolius (E.A.Hodgs.) Briscoe, Phytotaxa 202 (1): 61, 2015 (
*** Acrobolbus urvilleanus (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (
*** Acrobolbus viridis (Mitt.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 61, 2015 (
*** Acrobolbus wilsonii Nees, Syn. Hepat. 1: 5, 1844 (
** Acrobolbus wilsonii var. andinus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 522, 1885 (
Excluded from the genus
* Acrobolbus bispinosus (J.B.Jack et Steph.) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 459 (178), 1902 (
** Austrolophozia R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (
** Austrolophozia andina R.M.Schust., Nova Hedwigia 15: 495, 1968 (
*** Austrolophozia camensis (Steph.) Grolle ex Hässel et Solari, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 3 (6): 240, 1970 (
*** Austrolophozia paradoxa R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (
*** Goebelobryum Grolle, J. Hattori Bot. Lab. 25: 135, 1962 (
*** Goebelobryum grossitextum (Steph.) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (
*** Goebelobryum unguiculatum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (
*** Goebelobryum vermiculare J.J.Engel et Glenny, Nova Hedwigia 95 (3/4): 320, 2012 (
*** Lethocolea Mitt., Handb. N. Zeal. fl. 2: 751, 1867 (
** Lethocolea congesta (Lehm.) S.W.Arnell, Bot. Not. 108: 311, 1955 (
*** Lethocolea glossophylla (Spruce) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (
** Lethocolea indica G.Asthana et Maurya, Natl. Acad. Sci. Lett. 37 (6): 535, 2014 (
*** Lethocolea javanica (Schiffn.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (
** Lethocolea naruto-toganensis Furuki, Bryologist 104 (2): 306, 2001 (
*** Lethocolea pansa (Taylor) G.A.M.Scott et K.G.Beckm., Symp. Biol. Hung. 35: 212, 1987 (
*** Lethocolea radicosa (Lehm. et Lindenb.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (
* Lethocolea repens S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 67, 1969 (
*** Saccogynidium Grolle, J. Hattori Bot. Lab. 23: 43, 1960 [1961] (
** sect. Decurvum Grolle, J. Hattori Bot. Lab. 23: 59, 1960 (
*** Saccogynidium decurvum (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 59, 1960 [1961] (
** sect. Jugata Grolle, J. Hattori Bot. Lab. 23: 55, 1960 (
** Saccogynidium rigidulum (Nees) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (
** sect. Saccogynidium
*** Saccogynidium australe (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 49, 1960 [1961] (
** Saccogynidium caldense (Ångstr.) Grolle, J. Hattori Bot. Lab. 23: 44, 1960 [1961] (
** Saccogynidium goebelii (Herzog) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (
** Saccogynidium muricellum (De Not.) Grolle, J. Hattori Bot. Lab. 36: 80, 1972 [1973] (
*** Saccogynidium vasculosum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 23: 46, 1960 [1961] (
Incertae sedis
** Saccogynidium chiloscyphoides R.M.Schust., J. Hattori Bot. Lab. 26: 272, 1963 (
** Saccogynidium irregularospinum C.Gao, T.Cao et M.J.Lai, Bryologist 104 (1): 129, 2001 (
*** Anthelia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
*** Anthelia julacea (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
** Anthelia juratzkana (Limpr.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (
*** Arnellia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (
*** Arnellia fennica (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (
by J.J. Engel with contribution by J. Váňa (Neesioscyphus)
The placement of Pseudoisotachis in Balantiopsidaceae is only preliminary (cf.
*** Acroscyphella N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
*** Acroscyphella iwatsukii (N.Kitag.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
*** Acroscyphella phoenicorhiza (Grolle) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
*** Acroscyphella tjiwideiensis (Sande Lac.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (
** Pseudoisotachis Váňa, Polish Bot. J. 58 (1): 55, 2013 (
** Pseudoisotachis pocsii Váňa, Polish Bot. J. 58 (1): 55, 2013 (
*** Balantiopsis Mitt., Handb. N. Zeal. fl. 2: 751, 1867 (
*** Balantiopsis asymmetrica (Herzog) J.J.Engel, Nova Hedwigia 16: 93, 1968 (
*** Balantiopsis bisbifida (Steph.) Steph., Sp. Hepat. (Stephani) 4: 101, 1910 (
*** Balantiopsis brasiliensis Steph., Sp. Hepat. (Stephani) 4: 104, 1910 (
*** Balantiopsis cancellata (Nees) Steph., Sp. Hepat. (Stephani) 4: 103, 1910 (
*** Balantiopsis ciliaris S.Hatt., J. Jap. Bot. 41 (5): 129, 1966 (
** Balantiopsis ciliaris subsp. novoguineensis S.Hatt., J. Jap. Bot. 41 (5): 131, 1966 (
*** Balantiopsis convexiuscula Berggr., New Zealand Hepat.: 44, 1898 (
*** Balantiopsis crocea Herzog, Beih. Bot. Centralbl. 60B (1/2): 12, 1939 (
*** Balantiopsis diplophylla (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
** Balantiopsis diplophylla var. hockenii (Berggr.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 11, 1997 (
*** Balantiopsis erinacea (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (
*** Balantiopsis lingulata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 26, 1968 (
*** Balantiopsis montana (Colenso) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 12, 1997 (
** Balantiopsis neocaledonica Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922 (
** Balantiopsis paucidens Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 29, 1901 (
*** Balantiopsis purpurata Mitt., Rep. Challenger, Bot. 1 (3, 1): 86, 1884 (
*** Balantiopsis rosea Berggr., New Zealand Hepat.: 43, 1898 (
*** Balantiopsis splendens (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 55, 1997 (
*** Balantiopsis tumida Berggr., New Zealand Hepat.: 45, 1898 (
*** Balantiopsis verrucosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 16, 1997 (
*** Isotachis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 148, 1854 (
*** Isotachis armata (Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (
*** Isotachis aubertii (Schwägr.) Mitt., J. Linn. Soc., Bot. 22 (146): 322, 1886 (
* Isotachis boliviensis Gottsche, Sp. Hepat. (Stephani) 3: 670, 1909 (
** Isotachis chinensis C.Gao, T.Cao et J.Sun, Bryologist 105 (4): 694, 2002 [2003] (
** Isotachis erythrorhiza (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 338, 1885 (
*** Isotachis fragilis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 67, 1911 (
** Isotachis grandis Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1041, 1888 (
*** Isotachis grossidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 69, 1911 (
** Isotachis hastatistipula (Steph.) J.J.Engel, Phytotaxa 183 (4): 299, 2014 (
* Isotachis hians Steph., Sp. Hepat. (Stephani) 3: 665, 1909 (
*** Isotachis humectata (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 3: 654, 1909 (
** Isotachis inflata Steph., Arch. Mus. Nac. Rio de Janeiro 13: 113, 1905 (
*** Isotachis intortifolia (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (
** Isotachis japonica Steph., Sp. Hepat. (Stephani) 3: 652, 1909 (
* Isotachis lacustris Herzog, Hedwigia 74 (2): 94, 1934 (
*** Isotachis lopezii (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 84: 66, 1999 (
*** Isotachis lyallii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 149, 1854 (
*** Isotachis minima Pearson, Univ. Calif. Publ. Bot. 10 (4): 322, 1923 (
*** Isotachis montana Colenso, Trans. & Proc. New Zealand Inst. 21: 68, 1889 (
*** Isotachis multiceps (Lindenb. et Gottsche) Gottsche, Mexik. Leverm.: 105, 1863 (
** Isotachis multiceps var. fendleri Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 125, 1864 (
** Isotachis nigella Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 107, 1952 (
*** Isotachis obtusa Steph., Sp. Hepat. (Stephani) 6: 354, 1922 (
** Isotachis olivacea R.M.Schust., J. Hattori Bot. Lab. 83: 207, 1997 (
*** Isotachis plicata J.J.Engel, J. Hattori Bot. Lab. 83: 210, 1997 (
** Isotachis pusilla Steph., Sp. Hepat. (Stephani) 3: 655, 1909 (
** Isotachis riparia Rodway, Tasm. Bryoph.: 63, 1917 (
*** Isotachis serrulata (Sw.) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (
*** Isotachis spegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 220, 1885 (
* Isotachis sprucei Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (
*** Isotachis stephanii E.S.Salmon, Rev. Bryol. 28 (4): 75, 1901 (
* Isotachis vexans Steph., Sp. Hepat. (Stephani) 3: 662, 1909 (
*** Isotachis westlandica (E.A.Hodgs.) R.M.Schust., Nova Hedwigia 15: 455, 1968 (
*** Neesioscyphus Grolle, Österr. Bot. Z. 111 (1): 19, 1964 (
*** Neesioscyphus allionii (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 185, 1966 (
*** Neesioscyphus argillaceus (Nees) Grolle, Österr. Bot. Z. 111 (1): 24, 1964 (
*** Neesioscyphus bicuspidatus (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 182, 1966 (
*** Neesioscyphus carneus (Nees) Grolle, Österr. Bot. Z. 111 (1): 20, 1964 (
*** Neesioscyphus homophyllus (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 188, 1964 (
*** Ruizanthus R.M.Schust., Phytologia 39 (4): 240, 1978 (
*** Ruizanthus venezuelanus R.M.Schust., Phytologia 39 (4): 240, 1978 (
by J.J. Engel
*** Blepharidophyllum Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 151, 1873 (
*** Blepharidophyllum densifolium (Hook.) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (
*** Blepharidophyllum vertebrale (Gottsche) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (
*** Clandarium (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (
*** Clandarium clandestinum (Mont.) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (
*** Clandarium gottscheanum (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (
*** Clandarium xiphophyllum (Grolle) R.M.Schust., Phytologia 56 (2): 68, 1984 (
by M.A.M. Renner
Calypogeiaceae is shown to be monophyletic (
*** Calypogeia Raddi, Jungermanniogr. Etrusca: 31, 1818 (
** subg. Asperifoliae (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 115, 1969 (
*** Calypogeia arguta Nees et Mont., Naturgesch. Eur. Leberm. 3: 24, 1838 (
** Calypogeia sullivantii Austin, Hepat. bor.-amer.: 19, 1873 (
** subg. Calypogeia
*** Calypogeia andicola Bischl., Candollea 18: 79, 1962 (
** Calypogeia annabonensis Steph., Sp. Hepat. (Stephani) 6: 447, 1924 (
** Calypogeia azorica Bischl., Rev. Bryol. Lichénol. 37 (1): 116, 1970 (
*** Calypogeia azurea Stotler et Crotz, Taxon 32 (1): 74, 1983 (
*** Calypogeia bidentula (F.Weber) Nees, Syn. Hepat. 2: 199, 1845 (
** Calypogeia falcata Bischl., Candollea 18: 112, 1962 (
*** Calypogeia fissa (L.) Raddi, Jungermanniogr. Etrusca: 33, 1818 (
** Calypogeia fissa subsp. neogaea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 169, 1969 (
** Calypogeia goebelii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (
** Calypogeia goebelii var. siamensis N.Kitag., Beih. Nova Hedwigia 90: 165, 1988 (
*** Calypogeia grandistipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (
*** Calypogeia integristipula Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 662 (394), 1908 (
*** Calypogeia laxa Gottsche et Lindenb., Syn. Hepat. 5: 713, 1847 (
*** Calypogeia lechleri (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (
** Calypogeia lechleri var. densifolia (Steph.) Bischl., Candollea 18: 101, 1962 (
** Calypogeia longifolia Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (
** Calypogeia lophocoleoides Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (
** Calypogeia mascarenensis Bischl., Rev. Bryol. Lichénol. 37 (1): 89, 1970 (
** Calypogeia microstipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 670 (402), 1908 (
*** Calypogeia miquelii Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 200, 1845 (
*** Calypogeia muelleriana (Schiffn.) Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 217, 1901 (
** Calypogeia muelleriana subsp. blomquistii R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 187, 1969 (
*** Calypogeia neesiana (C.Massal. et Carestia) Müll.Frib., Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (
** Calypogeia neesiana subsp. subalpina (Inoue) Inoue, Mem. Natl. Sci. Mus. (Tokyo) 4: 58, 1971 (
*** Calypogeia oblata Herzog, Svensk Bot. Tidskr. 51 (1): 189, 1957 (
*** Calypogeia peruviana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (
*** Calypogeia rhombifolia (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 667 (399), 1908 (
** Calypogeia rhombifolia var. colombiana Bischl., Candollea 18: 104, 1962 (
*** Calypogeia sphagnicola (Arnell et J.Perss.) Warnst. et Loeske, Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (
*** Calypogeia subintegra (Gottsche, Lindenb. et Nees) Bischl., Candollea 18: 75, 1962 (
** Calypogeia subintegra var. dussiana (Steph.) Bischl., Candollea 18: 77, 1962 (
*** Calypogeia suecica (Arnell et J.Perss.) Müll.Frib., Beih. Bot. Centralbl. 17 (2): 224, 1904 (
*** Calypogeia tenax (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 664 (396), 1908 (
** Calypogeia uncinulatula Herzog, Hedwigia 67 (6): 250, 1927 (
Incertae sedis
*** Calypogeia aeruginosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (
* Calypogeia amazonica (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (
** Calypogeia angusta Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908 (
** Calypogeia apiculata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 668 (400), 1908 (
** Calypogeia asakawana S.Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107, 1964 (
** Calypogeia ceylanica S.Hatt. et Mizut., Candollea 23: 288, 1968 (
** Calypogeia contracta Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 139, 1975 (
** Calypogeia cuspidata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (
* Calypogeia decurrens (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 675 (407), 1908 (
** Calypogeia formosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 186, 1934 (
** Calypogeia fujisana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 135, 1975 (
** Calypogeia granulata Inoue, J. Jap. Bot. 43 (10/11): 468, 1968 (
** Calypogeia japonica Steph., Sp. Hepat. (Stephani) 6: 448, 1924 (
** Calypogeia khasiana Ajit P.Singh et V.Nath, Taiwania 52 (4): 320, 2007 (
** Calypogeia latissima Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (
*** Calypogeia lunata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (
** Calypogeia marginella Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 106, 1860 [1861] (
** Calypogeia obovata R.M.Schust., Phytologia 39 (4): 242, 1978 (
* Calypogeia steyermarkii Fulford, Mem. New York Bot. Gard. 11 (3): 305, 1968 (
** Calypogeia tosana (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 678 (410), 1908 (
** Calypogeia udarii Sudipa Das et D.K.Singh, Nelumbo 53: 194, 2011 (
** Eocalypogeia (R.M.Schust.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (
** Eocalypogeia quelpaertensis (S.Hatt. et Inoue) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (
** Eocalypogeia schusterana (S.Hatt. et Mizut.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (
** Metacalypogeia (S.Hatt.) Inoue, J. Hattori Bot. Lab. 21: 231, 1959 (
*** Metacalypogeia alternifolia (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 185, 1964 (
** Metacalypogeia cordifolia (Steph.) Inoue, J. Hattori Bot. Lab. 21: 233, 1959 (
** Mizutania Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (
*** Mizutania riccardioides Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (
*** Mnioloma Herzog, Ann. Bryol. 3: 115, 1930 (
** subg. Caracoma (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 833, 1995 (
** Mnioloma bolivianum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (
*** Mnioloma caespitosum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (
*** Mnioloma cellulosum (Spreng.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 836, 1995 (
*** Mnioloma crenulatum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (
*** Mnioloma cyclostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 843, 1995 (
** Mnioloma elliottii (Steph.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 841, 1995 (
** Mnioloma fissistipulum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (
*** Mnioloma fuscum (Lehm.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 848, 1995 (
*** Mnioloma nephrostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (
** Mnioloma novaezelandiae J.J.Engel, Cryptog. Bryol. 27 (1): 111, 2006 (
*** Mnioloma parallelogramum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (
** Mnioloma retusum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (
** Mnioloma stamatotonum M.A.M.Renner et E.A.Br., Fieldiana, Bot. (n.ser.) 47: 173, 2008 (
*** Mnioloma venezuelanum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (
** subg. Mnioloma, Fragm. Florist. Geobot. 40 (2): 833, 1995 (
*** Mnioloma rhynchophyllum Herzog, Ann. Bryol. 3: 120, 1930 (
by N.A. Konstantinova
*** Endogemma Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (
*** Endogemma caespiticia (Lindenb.) Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (
Placement of Geocalycaceae in Jungermanniinae follows
*** Geocalyx Nees, Naturgesch. Eur. Leberm. 1: 97, 1833 (
*** Geocalyx caledonicus Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 205 (265), 1908 (
*** Geocalyx graveolens (Schrad.) Nees, Naturgesch. Eur. Leberm. 2: 397, 1836 (
** Geocalyx lancistipulus (Steph.) S.Hatt., J. Jap. Bot. 28 (8): 234, 1953 (
** Geocalyx orientalis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxix, 1889 [1890] (
by J. Váňa
The treatment of the family follows
*** Acrolophozia R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 259, 1966 (
*** Acrolophozia fuegiana R.M.Schust., Nova Hedwigia 15: 499, 1968 (
*** Acrolophozia pectinata R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 261, 1966 (
*** Acrolophozia sulcata Hässel, J. Bryol. 11 (1): 108, 1980 (
** Nanomarsupella R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (
*** Nanomarsupella xenophylla (R.M.Schust.) R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (
** Paramomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 134, 1996 (
*** Paramomitrion paradoxum R.M.Schust., J. Hattori Bot. Lab. 80: 135, 1996 (
** Cryptocoleopsis Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (
*** Cryptocoleopsis imbricata Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (
*** Gymnomitrion Corda, Gen. hepat.: 651, 1829 (
*** Gymnomitrion adustum Nees, Naturgesch. Eur. Leberm. 1: 120, 1833 (
*** Gymnomitrion africanum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (
*** Gymnomitrion alpinum (Gottsche ex Husn.) Schiffn., Österr. Bot. Z. 53 (7): 280, 1903 (
*** Gymnomitrion asperulatum R.M.Schust., Acta Acad. Ped. Agr., Sect. Biol. 24: 114, 2003 (
*** Gymnomitrion atrofilum Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (
*** Gymnomitrion bolivianum (Steph.) Váňa, Novon 20 (2): 225, 2010 (
*** Gymnomitrion brevissimum (Dumort.) Warnst., Hedwigia 53 (3): 196, 1913 (
*** Gymnomitrion commutatum (Limpr.) Schiffn., Magyar Bot. Lapok 13: 304, 1914 [1915] (
*** Gymnomitrion concinnatum (Lightf.) Corda, Gen. hepat.: 651, 1829 (
*** Gymnomitrion corallioides Nees, Naturgesch. Eur. Leberm. 1: 118, 1833 (
*** Gymnomitrion crenatilobum Grolle, Khumbu Himal 1 (4): 278, 1966 (
*** Gymnomitrion crenulatum Gottsche ex Carrington, Trans. Bot. Soc. Edinburgh 7 (3): 444, 1863 (
*** Gymnomitrion crystallocaulon (Grolle) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
** Gymnomitrion incompletum (Gottsche) R.M.Schust. ex Váňa, J. Hattori Bot. Lab. 40: 186, 1976 (
*** Gymnomitrion laceratum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (
*** Gymnomitrion miniatum Lindenb. et Gottsche, Syn. Hepat. 4: 617, 1846 (
*** Gymnomitrion minutulum (Hässel) Váňa, Novon 20 (2): 225, 2010 (
** Gymnomitrion moralesae Váňa, J. Hattori Bot. Lab. 48: 230, 1980 (
*** Gymnomitrion mucronulatum (N.Kitag.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 53, 1962 (
** Gymnomitrion mucrophorum R.M.Schust., Bryologist 98 (2): 243, 1995 (
*** Gymnomitrion nigrum (Grolle et Váňa) Váňa, Novon 20 (2): 225, 2010 (
*** Gymnomitrion noguchianum S.Hatt., J. Jap. Bot. 27 (2): 55, 1952 (
*** Gymnomitrion obtusilobum N.Kitag., Bull. Univ. Mus. Univ. Tokyo 8: 229, 1975 (
*** Gymnomitrion obtusum Lindb., Morgonbladet (Helsinki) 1877 (30, 6 Feb): 2, 1877 (
*** Gymnomitrion pacificum Grolle, Trans. Brit. Bryol. Soc. 5 (1): 92, 1966 (
*** Gymnomitrion revolutum (Nees) H.Philib., Rev. Bryol. 17 (3): 34, 1890 (
** Gymnomitrion revolutum subsp. novoguineanensis (R.M.Schust.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Gymnomitrion rubidum (Mitt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Gymnomitrion setaceum Grolle et Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (
*** Gymnomitrion sinense Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 176, 1951 (
** Gymnomitrion strictum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (
** Gymnomitrion strictum var. inaequale R.M.Schust., J. Hattori Bot. Lab. 80: 118, 1996 (
*** Gymnomitrion subintegrum (S.W.Arnell) Váňa, Novon 20 (2): 225, 2010 (
*** Gymnomitrion truncatoapiculatum Herzog, Hedwigia 74 (2): 81, 1934 (
*** Gymnomitrion verrucosum W.E.Nicholson, Symb. Sin. 5: 10, 1930 (
*** Marsupella Dumort., Commentat. Bot. (Dumortier): 114, 1822 (
*** Marsupella alata S.Hatt. et N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 79, 1960 (
*** Marsupella andreaeoides (Lindb.) Müll.Frib., Feddes Repert. Spec. Nov. Regni Veg. 54 (2/3): 214, 1951 (
*** Marsupella apiculata Schiffn., Österr. Bot. Z. 53 (6): 249, 1903 (
*** Marsupella aquatica (Lindenb.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 44 (8): 267, 1896 [1897] (
*** Marsupella arctica (Berggr.) Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 26, 1906 (
*** Marsupella boeckii (Austin) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 409, 1893 (
*** Marsupella bolanderi (Austin) Underw., Zoe 1 (12): 365, 1891 (
*** Marsupella condensata (Ångstr. ex C.Hartm.) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 420, 1893 (
*** Marsupella disticha Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 164 (25), 1901 (
*** Marsupella emarginata (Ehrh.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (
** Marsupella emarginata subsp. tubulosa (Steph.) N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 76, 1960 (
** Marsupella emarginata subsp. tubulosa var. apertifolia (Steph.) N.Kitag., J. Hattori Bot. Lab. 26: 89, 1963 (
** Marsupella emarginata subsp. tubulosa var. patens N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 77, 1960 (
** Marsupella emarginata subsp. tubulosa var. tubulosa (Steph.) N.Kitag. ex Váňa et L.Söderstr., Phytotaxa 183 (4): 288, 2014 (
*** Marsupella funckii (F.Weber et D.Mohr) Dumort., Recueil Observ. Jungerm.: 24, 1835 (
*** Marsupella microphylla R.M.Schust., Phytologia 39 (4): 249, 1978 (
** Marsupella minutissima N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 81, 1960 (
*** Marsupella neesii Sande Lac. ex Schiffn., Consp. Hepat. Arch. Ind.: 70, 1898 (
*** Marsupella paroica R.M.Schust., Bryologist 60 (2): 145, 1957 (
*** Marsupella profunda Lindb., Rev. Bryol. 14 (2): 19, 1887 (
*** Marsupella pseudofunckii S.Hatt., J. Hattori Bot. Lab. 4: 63, 1950 (
*** Marsupella sparsifolia (Lindb.) Dumort., Bull. Soc. Roy. Bot. Belgique 13: 128, 1874 (
** Marsupella sparsifolia subsp. childii R.M.Schust., Phytotaxa 183 (4): 288, 2014 (
*** Marsupella sphacelata (Giesecke ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (
*** Marsupella spiniloba R.M.Schust. et Damsh., Phytologia 63 (5): 326, 1987 (
*** Marsupella sprucei (Limpr.) Bernet, Cat. hép. Suisse: 33, 1888 (
** Marsupella stableri Spruce, Rev. Bryol. 8 (6): 96, 1881 (
*** Marsupella stoloniformis N.Kitag., J. Hattori Bot. Lab. 30: 201, 1967 (
** Marsupella stoloniformis subsp. vermiformis R.M.Schust., J. Hattori Bot. Lab. 80: 72, 1996 (
*** Marsupella yakushimensis (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 80, 1944 (
*** Poeltia Grolle, Khumbu Himal 1 (4): 280, 1966 (
*** Poeltia campylata Grolle, Khumbu Himal 1 (4): 280, 1966 (
*** Prasanthus Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889 (
** Prasanthus jamalicus Potemkin, Ann. Bot. Fenn. 29 (4): 319, 1992 (
*** Prasanthus suecicus (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889 (
*** Nardia Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (
*** Nardia arnelliana Grolle, Bot. Mag. (Tokyo) 77 (914): 297, 1964 (
*** Nardia assamica (Mitt.) Amakawa, J. Hattori Bot. Lab. 26: 23, 1963 (
*** Nardia breidleri (Limpr.) Lindb., Helsingf. Dagbl. 1880 (311, 15 Nov.): 2, 1880 (
*** Nardia compressa (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (
*** Nardia flagelliformis Inoue, J. Jap. Bot. 46 (1): 1, 1971 (
*** Nardia geoscyphus (De Not.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 371, 1874 (
* Nardia geoscyphus var. dioica Bakalin, Arctoa 18: 87, 2009 [2010] (
* Nardia geoscyphus var. suberecta (Lindb. ex Kaal.) Váňa, Phytotaxa 76 (3): 37, 2013 (
*** Nardia grollei Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 491, 2009 (
*** Nardia insecta Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878 (
*** Nardia japonica Steph., Bull. Herb. Boissier 5 (2): 101, 1897 (
* Nardia kamtschatica Arnell et C.E.O.Jensen, Hedwigia 67 (1/2): 111, 1927 (
* Nardia leptocaulis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (
*** Nardia lescurii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 115, 1884 (
** Nardia minutifolia Furuki, Bryol. Res. 9 (3): 73, 2006 (
** Nardia nuda (Lindenb. et Gottsche) Váňa, Folia Geobot. Phytotax. 8 (2): 193, 1973 (
*** Nardia poeltii Váňa, J. Hattori Bot. Lab. 36: 73, 1972 [1973] (
*** Nardia scalaris Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (
* Nardia scalaris var. botryoidea (R.M.Schust.) Váňa, Phytotaxa 76 (3): 38, 2013 (
* Nardia scalaris var. harae (Amakawa) Váňa, Phytotaxa 76 (3): 38, 2013 (
*** Nardia subclavata (Steph.) Amakawa, J. Jap. Bot. 32 (2): 40, 1957 (
*** Nardia succulenta (A.Rich.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 519, 1885 (
*** Nardia unispiralis Amakawa, J. Jap. Bot. 32 (6): 167, 1957 (
Placement of Gyrothyraceae in Jungermanniinae follows
** Gyrothyra M.Howe, Bull. Torrey Bot. Club 24 (4): 201, 1897 (
*** Gyrothyra underwoodiana M.Howe, Bull. Torrey Bot. Club 24 (4): 202, 1897 (
*** Harpanthus Nees, Naturgesch. Eur. Leberm. 2: 351, 1836 (
*** Harpanthus drummondii (Taylor) Grolle, Österr. Bot. Z. 112 (3): 274, 1965 (
*** Harpanthus flotovianus (Nees) Nees, Naturgesch. Eur. Leberm. 2: 353, 1836 (
*** Harpanthus scutatus (F.Weber et D.Mohr) Spruce, Trans. Bot. Soc. Edinburgh 3 (1/4): 209, 1850 (
by N. Konstantinova
The family Hygrobiellaceae was validated by
*** Hygrobiella Spruce, Cephalozia: 73, 1882 (
** Hygrobiella intermedia Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (
*** Hygrobiella laxifolia (Hook.) Spruce, Cephalozia: 74, 1882 (
** Hygrobiella squamosa Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (
by J. Váňa
The monogeneric status of Jackiella is supported by a molecular study based on three loci by
*** Jackiella Schiffn., Hep. Fl. Buitenzorg: 211, 1900 (
** Jackiella angustifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 296, 1950 (
** Jackiella ceylanica Schiffn. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 212 (272), 1908 (
*** Jackiella curvata E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand 85 (4): 571, 1958 (
*** Jackiella javanica Schiffn., Hep. Fl. Buitenzorg: 212, 1900 (
** Jackiella javanica var. cavifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900 (
** Jackiella javanica var. cordifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900 (
** Jackiella renifolia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (
** Jackiella sinensis (W.E.Nicholson) Grolle, Österr. Bot. Z. 111 (2/3): 186, 1964 (
** Jackiella singapurensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (
** Jackiella singapurensis var. philippinensis N.Kitag., Misc. Bryol. Lichenol. 9 (1): 9, 1981 (
Excluded from the genus
* Jackiella unica Steph., Sp. Hepat. (Stephani) 6: 318, 1922 (
by J. Váňa
The treatment of Jungermanniaceae follows
** Delavayella Steph., Hedwigia 33 (1): 4, 1894 (
** Delavayella serrata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 211, 1894 (
** Delavayella serrata var. purpurea P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 38, 1955 (
** Liochlaena Nees, Syn. Hepat. 2: 150, 1845 (
*** Liochlaena lanceolata Nees, Syn. Hepat. 2: 150, 1845 (
*** Liochlaena subulata (A.Evans) Schljakov, Pečen. Mchi Sev. SSSR 4: 71, 1981 (
** Eremonotus Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 200, 1900 (
*** Eremonotus myriocarpus (Carrington) Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 201, 1900 (
*** Jungermannia L., Sp. Pl. 1: 1131, 1753 (
*** Jungermannia atrovirens Dumort., Syll. Jungerm. Europ.: 51, 1831 (
*** Jungermannia borealis Damsh. et Váňa, Lindbergia 4 (1/2): 5, 1977 (
* Jungermannia erectii Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 117, 2007 (
*** Jungermannia exsertifolia Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (
** Jungermannia exsertifolia subsp. cordifolia (Dumort.) Váňa, Folia Geobot. Phytotax. 8 (3): 268, 1973 (
*** Jungermannia gollanii Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (
** Jungermannia konstantinovae Bakalin et Vilnet, Arctoa 18: 161, 2009 [2010] (
*** Jungermannia ovatotrigona (Steph.) Grolle, Feddes Repert. 82 (1): 90, 1971 (
*** Jungermannia polaris Lindb., Öfvers. Kongl. Vetensk.-Akad. Förh. 23 (10): 560, 1866 [1867] (
*** Jungermannia pumila With., Arr. Brit. Pl., ed. 3, 3: 883, 1796 (
Taxa doubtfully belonging to the genus
* Jungermannia amentacea Bertol., Mem. Reale Accad. Sci. Ist. Bologna (ser. 2) 1: 19, 1862 (
* Jungermannia brasiliensis Raddi, Critt. Brasil.: 15, 1822 (
* Jungermannia chinensis Osbeck, Dagb. Ostind. Resa: 221, 1757 (
* Jungermannia cordata Vill., Hist. Pl. Dauphiné (Villars) 3: 923, 1789 (
* Jungermannia crenulata Schmidel, Jungerm. Char.: 20, 1760 (
* Jungermannia creutzeri Kremer, Monogr. hépat. Moselle: 26, 1837 (
* Jungermannia digitata C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 326, 1827 (
* Jungermannia dubioides H.A.Mill., Phytologia 47 (4): 322, 1981 (
* Jungermannia fernandeziana Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884 (
* Jungermannia hexagona Schwägr., Hist. Musc. Hepat. Prodr.: 18, 1814 (
* Jungermannia holandriana Kremer, Monogr. hépat. Moselle: 25, 1837 (
* Jungermannia incerta Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882 (
* Jungermannia lateriflora Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (
* Jungermannia lescuriana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 67, 1866 (
* Jungermannia longiretis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxv, 1889 [1890] (
* Jungermannia mastigophora Spreng. Neue Entdeck. Pflanzenk. 2: 99, 1821 (
* Jungermannia michelii Mérat, Nouv. fl. env. Paris (ed. 2) 1: 219, 1821 (
* Jungermannia minima Scop., Fl. Carniol. (ed. 2) 2: 350, 1772 (
* Jungermannia odorata With., Bot. arr. veg. Gr. Brit. 2: 693, 1776 (
* Jungermannia peltata Schmidel, Jungerm. Char.: 14, 1760 (
* Jungermannia quadridigitata Griff., Not. pl. asiat. 2: 314, 1849 (
* Jungermannia sauteri De Not. ex Rabenh., Hedwigia 1 (20): 121, 1857 (
* Jungermannia secunda Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (
* Jungermannia stereocaulis Bory, Voy. Uranie, Bot. 4: 130, 1827 (
* Jungermannia submersa Kremer, Monogr. hépat. Moselle: 36, 1837 (
* Jungermannia sullivantiana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 66, 1866 (
* Jungermannia supina Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 86, 1795 [1796] (
* Jungermannia tenuis Ehrh., Beitr. Naturk. (Ehrhart) 4: 45, 1789 (
* Jungermannia uncifolia Steph., Hedwigia 34 (2): 51, 1895 (
* Jungermannia vernicosa Cass. ex Mérat, Nouv. fl. env. Paris (ed. 2) 1: 221, 1821 (
*** Mesoptychia (Lindb.) A.Evans, Ottawa Naturalist 17: 15, 1903 (
*** Mesoptychia badensis (Gottsche ex Rabenh.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (
*** Mesoptychia bantriensis (Hook.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (
** Mesoptychia bantriensis subsp. wallfischii (Ştefănuţ) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (
* Mesoptychia chichibuensis (Inoue) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
* Mesoptychia collaris (Nees) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
* Mesoptychia fitzgeraldiae (Paton et A.R.Perry) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
*** Mesoptychia gillmanii (Austin) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
*** Mesoptychia heterocolpos (Thed. ex Hartm.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
** Mesoptychia heterocolpos var. arctica (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
** Mesoptychia heterocolpos var. harpanthoides (Bryhn et Kaal.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (
*** Mesoptychia igiana (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
* Mesoptychia mamatkulovii (Duda) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia mayebarae (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia morrisoncola (Horik.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
* Mesoptychia polymorpha Stotler, Crand.-Stotl. et Bakalin, Polish Bot. J. 58 (1): 82, 2013 (
*** Mesoptychia rutheana (Limpr.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
** Mesoptychia rutheana var. laxa (Schiffn. ex Burrell) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia sahlbergii (Lindb. et Arnell) A.Evans, Ottawa Naturalist 17: 15, 1903 (
*** Mesoptychia subcrispa (Herzog) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (
*** Mesoptychia turbinata (Raddi) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (
* Mesoptychia ussuriensis (Bakalin) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (
** Rivulariella D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (
** Rivulariella gemmipara (A.Evans) D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (
*** Notoscyphus Mitt., Fl. vit.: 407, 1871 [1873] (
*** Notoscyphus lutescens (Lehm. et Lindenb.) Mitt., Fl. vit.: 407, 1871 [1873] (
*** Saccogyna Dumort., Commentat. Bot. (Dumortier): 113, 1822 (
* Saccogyna ligulata Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 207 (267), 1908 (
** Saccogyna subacuta Steph., Sp. Hepat. (Stephani) 6: 316, 1922 (
*** Saccogyna viticulosa (L.) Dumort., Syll. Jungerm. Europ.: 74, 1831 (
Excluded from the genus
* Saccogyna tridens Steph., Sp. Hepat. (Stephani) 6: 317, 1922 (
by J. Váňa
Solenostomataceae was first defined by
** Aponardia (R.M.Schust.) Váňa, Phytotaxa 65: 46, 2012 (
*** Aponardia huerlimannii (Váňa et Grolle) Váňa, Phytotaxa 65: 46, 2012 (
** Arctoscyphus Hässel, Lindbergia 16 (4): 133, 1990 [1992] (
** Arctoscyphus fuegiensis (C.Massal.) Hässel, Cryptog. Bryol. Lichénol. 17 (3): 164, 1996 (
** Arctoscyphus ronsmithii Hässel, Lindbergia 16 (4): 133, 1990 [1992] (
** Cryptocolea R.M.Schust., Amer. Midl. Naturalist 49 (2): 414, 1953 (
*** Cryptocolea imbricata R.M.Schust., Amer. Midl. Naturalist 49 (2): 417, 1953 (
** Diplocolea Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (
*** Diplocolea sikkimensis Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (
*** Solenostoma Mitt., J. Proc. Linn. Soc., Bot. 8 (29): 51, 1864 [1865] (
** subg. Eucalyx (Lindb.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (
*** Solenostoma bilobum (S.Hatt. ex Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 286, 2009 (
*** Solenostoma emarginatum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma flagellatum (S.Hatt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma hokkaidense (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma obovatum (Nees) C.Massal., Epat. erb. critt. ital.: 17, 1903 (
*** Solenostoma obscurum (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1013, 1969 (
*** Solenostoma schusteranum (J.D.Godfrey et G.Godfrey) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma subtilissimum (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (
** subg. Metasolenostoma Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (
*** Solenostoma fusiforme (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 944, 1969 (
*** Solenostoma gracillimum (Sm.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 972, 1969 (
*** Solenostoma handelii (Schiffn.) Müll.Frib., Beitr. Kryptogamenfl. Schweiz 10 (2): 38, 1947 (
** Solenostoma lignicola (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma limbatifolium (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (
** Solenostoma philippinense Váňa, Phytotaxa 152 (1): 43, 2013 (
*** Solenostoma rubrum (Gottsche) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 975, 1969 (
*** Solenostoma suborbiculatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
** subg. Plectocolea Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 (
*** Solenostoma balfourii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma borneense (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma callithrix (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 486 (48), 1901 (
*** Solenostoma champawatense (S.N.Srivast. et Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma comatum (Nees) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 73, 1981 (
*** Solenostoma comatum var. novae-guineae (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma crenuliforme (Austin) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (
*** Solenostoma decolor (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma dusenii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma erectum (Amakawa) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 66, 1981 (
* Solenostoma flagellalioides C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (
*** Solenostoma flavialbicans (Amakawa et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma fossombronioides (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (
*** Solenostoma glaucum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (
* Solenostoma gongshanense (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (
*** Solenostoma haskarlianum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (
*** Solenostoma hattorianum (Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 287, 2009 (
*** Solenostoma hirticalyx (Steph.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma horikawanum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma hyalinum (Lyell) Mitt., Nat. hist. Azores: 319, 1870 (
*** Solenostoma infuscum (Mitt.) Hentschel, Pl. Syst. Evol. 268 (1/4): 152, 2007 (
*** Solenostoma infuscum var. ovicalyx (Steph.) Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 288, 2009 (
** Solenostoma kurilense (Bakalin) Váňa, Phytotaxa 152 (1): 40, 2013 (
* Solenostoma lixingjiangii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (
*** Solenostoma marginatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (
*** Solenostoma micranthum (Mitt.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
** Solenostoma montanum (Steph.) Váňa, Phytotaxa 65: 44, 2012 (
* Solenostoma multicarpum (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
** Solenostoma nilgiriense (A.Alam, Ad.Kumar et S.C.Srivast.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma obliquifolium (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma onraedtii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
* Solenostoma orbicularifolium (Piippo ex C.Gao et Bai) Váňa, Phytotaxa 222 (2): 199, 2015 (
*** Solenostoma otianum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 984, 1969 (
*** Solenostoma ovalifolium (Amakawa) Váňa, Phytotaxa 152 (1): 40, 2013 (
*** Solenostoma paroicum (Schiffn.) R.M.Schust., Amer. Midl. Naturalist 49 (2): 402, 1953 (
*** Solenostoma plagiochilaceum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma polyrhizoides (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma radicellosum Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 [1865] (
*** Solenostoma renauldii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
*** Solenostoma rigidulum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (
*** Solenostoma rosulans (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma rotundatum Amakawa, J. Jap. Bot. 31 (2): 50, 1956 (
*** Solenostoma rubripunctatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 898, 1969 (
* Solenostoma rupicola (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma sikkimense (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
*** Solenostoma tetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (
* Solenostoma tetragonum var. kodaikanalense A.Alam, D.Sharma et So.Yadav, Phytotaxonomy 12: 70, 2012 (
*** Solenostoma torticalyx (Steph.) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 69, 1981 (
*** Solenostoma truncatum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (
** Solenostoma truncatum var. setulosum (Herzog) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
*** Solenostoma tuberculiferum (Herzog) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma unispire (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma virgatum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
*** Solenostoma vulcanicola (Schiffn.) Nyushko ex Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 289, 2009 (
* Solenostoma zangmuii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
** subg. Solenostoma
*** Solenostoma amoenum (Lindenb. et Gottsche) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma amplexifolium (Hampe) Váňa et Schäf.-Verw., Acta Bot. Hung. 51 (3/4): 408, 2009 (
** Solenostoma appalachianum R.M.Schust. ex Bakalin, Arctoa 23: 127, 2014 (
*** Solenostoma appressifolium (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 494, 2009 (
*** Solenostoma appressifolium var. minor (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma appressifolium var. nigricans (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma ariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma atrobrunneum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (
*** Solenostoma atrorevolutum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma atrovirens Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 493 (55), 1901 (
*** Solenostoma baueri (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 495 (57), 1901 (
*** Solenostoma bengalense (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
** Solenostoma breviflorum Kashyap et R.S.Chopra, Liverworts W. Himal. 2: 85, 1932 (
*** Solenostoma caeleste (Inoue et Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
* Solenostoma caoi (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma caucasicum (Váňa) Konstant., Arctoa 1: 123, 1992 (
*** Solenostoma chenianum (C.Gao, Y.H.Wu et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (
*** Solenostoma clavellatum Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 491 (53), 1901 (
*** Solenostoma confertissimum (Nees) Schljakov, Novosti Sist. Nizš. Rast. 17: 239, 1980 (
*** Solenostoma coniflorum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (
*** Solenostoma crassulum (Nees et Mont.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (
** Solenostoma cryptogynum R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (
*** Solenostoma cyclops (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (
*** Solenostoma diversiclavellatum (Amakawa et Grolle) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma dulongense Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 497, 2009 (
*** Solenostoma exsertum (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (
*** Solenostoma faurieanum (Beauverd) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (
*** Solenostoma flagellare (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma flavorevolutum (Váňa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (
*** Solenostoma grollei (D.G.Long et Váňa) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (
*** Solenostoma grosseverrucosum (Amakawa et S.Hatt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Solenostoma heterolimbatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma hewsoniae (Amakawa et Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (
*** Solenostoma hiugaense Amakawa, J. Jap. Bot. 31 (2): 47, 1956 (
*** Solenostoma indrodayanum (Sushil K.Singh et D.K.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma inundatum (Hook.f. et Taylor) Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (
*** Solenostoma javanicum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (
** Solenostoma kanaii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma kashyapii (S.C.Srivast., S.Srivast. et D.Sharma) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma lanigerum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (
*** Solenostoma macrocarpum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (
*** Solenostoma mamatkulovii (Váňa et Zerov) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
* Solenostoma microphyllum C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 206, 1981 (
* Solenostoma microrevolutum (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma mildbraedii (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 387, 2002 (
*** Solenostoma niveum (Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (
* Solenostoma novazelandiae R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (
*** Solenostoma ohbae (Amakawa) C.Gao, Bryofl. Xizang: 495, 1985 (
*** Solenostoma orbiculatum (Colenso) R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (
*** Solenostoma parvitextum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma patoniae (Grolle, D.B.Schill et D.G.Long) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (
*** Solenostoma pocsii (Váňa) Bakalin, Arctoa 18: 160, 2009 [2010] (
*** Solenostoma poeltii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (
*** Solenostoma pseudocyclops (Inoue) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (
** Solenostoma pseudopyriflorum Bakalin et Vilnet, Arctoa 18: 159, 2009 [2010] (
*** Solenostoma purpuratum (Mitt.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (
*** Solenostoma pyriflorum Steph., Sp. Hepat. (Stephani) 6: 83, 1917 (
*** Solenostoma pyriflorum var. gracillimum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (
** Solenostoma pyriflorum var. major (S.Hatt.) Bakalin, Arctoa 17: 230, 2008 [2009] (
*** Solenostoma pyriflorum var. minutissimum (Amakawa) Bakalin, Arctoa 16: 208, 2007 [2008] (
*** Solenostoma raujeanum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma riclefii Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (
*** Solenostoma sanguinolentum (Griff.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (
*** Solenostoma schaulianum (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
*** Solenostoma shimizuanum (S.Hatt. ex Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma speciosum (Horik.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (
** Solenostoma speciosum subsp. villosum (R.M.Schust.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 246, 2009 (
*** Solenostoma sphaerocarpum (Hook.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 499 (61), 1901 (
*** Solenostoma stephanii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 496 (58), 1901 (
*** Solenostoma stoloniferum (Steph.) S.W.Arnell, Hepat. South Africa: 316, 1963 (
*** Solenostoma strictum (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (
*** Solenostoma subacutum (Herzog) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (
*** Solenostoma subrubrum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (
* Solenostoma sunii Bakalin et Vilnet, Bot. Pacifica 3 (2): 15, 2014 (
* Solenostoma totopapillosum (E.A.Hodgs.) R.M.Schust., Bryologist 100 (3): 366, 1997 (
*** Solenostoma udarii (S.C.Srivast. et P.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
* Solenostoma ventroversum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
*** Solenostoma zantenii (Amakawa) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
* Solenostoma zengii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (
by J. Váňa
Southbyaceae has traditionally been placed in Arnelliaceae (e.g.
*** Gongylanthus Nees, Naturgesch. Eur. Leberm. 2: 405, 1836 (
*** Gongylanthus dusenii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (
*** Gongylanthus ericetorum (Raddi) Nees, Naturgesch. Eur. Leberm. 2: 407, 1836 (
*** Gongylanthus granatensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (
*** Gongylanthus himalayensis Grolle, Khumbu Himal 1 (4): 285, 1966 (
*** Gongylanthus javanicus Grolle, J. Jap. Bot. 40 (7): 206, 1965 (
*** Gongylanthus liebmanianus (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (
*** Gongylanthus limbatus (Herzog) Grolle et Váňa, Folia Geobot. Phytotax. 9 (2): 198, 1974 (
*** Gongylanthus muelleri (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (
** Gongylanthus oniscoides (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 386 (42), 1906 (
*** Gongylanthus richardsii E.W.Jones, Trans. Brit. Bryol. Soc. 4 (4): 650, 1964 (
*** Southbya Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (
*** Southbya gollanii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 381 (37), 1906 (
*** Southbya nigrella (De Not.) Henriq., Bol. Soc. Brot. 4: 244, 1886 [1887] (
*** Southbya organensis Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 53, 1950 (
*** Southbya tophacea (Spruce) Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (
by J. Váňa
In the absence of molecular data, the phylogenetic affinities of Stephaniella and Stephaniellidium remain equivocal, but there is no evidence to support their placement in either the Arnelliaceae or Southbyaceae where they have been placed recently. Thus the family as construed by
*** Stephaniella J.B.Jack, Hedwigia 33 (1): 11, 1894 (
* Stephaniella boliviensis Steph., Biblioth. Bot. 87 (2): 182, 1916 (
*** Stephaniella hamata Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1024 (87), 1901 (
*** Stephaniella paraphyllina J.B.Jack, Hedwigia 33 (1): 11, 1894 (
*** Stephaniella rostrata U.Schmitt, Österr. Bot. Z. 115 (2): 124, 1968 (
*** Stephaniella uncifolia S.Winkl., Österr. Bot. Z. 115 (2): 124, 1968 (
** Stephaniellidium S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (
*** Stephaniellidium sleumeri (Müll.Frib.) S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (
*** Trichotemnoma R.M.Schust., Nova Hedwigia 15: 440, 1968 (
*** Trichotemnoma corrugatum (Steph.) R.M.Schust., Nova Hedwigia 15: 440, 1968 (
The recognition of Blepharostomataceae follows
*** Blepharostoma (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
** Blepharostoma arachnoideum M.Howe, Mem. Torrey Bot. Club 7: 140, 1899 (
** Blepharostoma indicum G.Asthana, M.Saxena et Maurya, J. Bryol. 35 (4): 267, 2013 (
** Blepharostoma minor Horik., Hikobia 1 (2): 104, 1951 [1952] (
*** Blepharostoma trichophyllum (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (
** Blepharostoma trichophyllum subsp. brevirete (Bryhn et Kaal.) R.M.Schust., Bull. Natl. Mus. Canada 164: 16, 1959 (
by L. Söderström, J. Váňa, R. Stotler, B.J. Crandall-Stotler and J.J. Engel
The treatment of Brevianthaceae follows
*** Brevianthus J.J.Engel et R.M.Schust., Phytologia 47 (4): 317, 1981 (
*** Brevianthus flavus (Grolle) J.J.Engel et R.M.Schust., Phytologia 47 (4): 318, 1981 (
** Brevianthus flavus subsp. crenulatus J.J.Engel, Nova Hedwigia 93 (3/4): 406, 2011 (
* Brevianthus hypocanthidium M.A.M.Renner et J.J.Engel, PhytoKeys 50: 46, 2015 (
*** Tetracymbaliella Grolle, Nova Hedwigia 3 (1): 48, 1961 (
** Tetracymbaliella comptonii (Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 164, 1963 [1964] (
*** Tetracymbaliella cymbalifera (Hook.f. et Taylor) Grolle, Nova Hedwigia 3 (1): 50, 1961 (
*** Tetracymbaliella decipiens (Gottsche) Grolle, Nova Hedwigia 3 (1): 49, 1961 (
*** Tetracymbaliella subsimplex (Austin) J.J.Engel, Phytotaxa 207 (2): 185, 2015 (
** Chonecolea Grolle, Rev. Bryol. Lichénol. 25 (3/4): 294, 1956 [1957] (
** Chonecolea acutiloba (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 321, 1980 (
** Chonecolea andina Grolle et Váňa, J. Hattori Bot. Lab. 48: 229, 1980 (
** Chonecolea doellingeri (Nees) Grolle, Rev. Bryol. Lichénol. 25 (3/4): 295, 1956 [1957] (
** Chonecolea ruwenzorensis E.W.Jones, J. Bryol. 13 (4): 498, 1986 (
** Chonecolea schusteri Udar et Ad.Kumar, Bryologist 85 (3): 315, 1982 (
** Chonecolea verae Potemkin, Proc. int. meeting 90th anniv. Abramova: 165, 2005 (
** Grollea R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964 (
** Grollea antheliopsis R.M.Schust., Nova Hedwigia 8 (3/4): 288, 1964 (
by D. Bell
*** Herbertus Gray, Nat. Arr. Brit. Pl. 1: 705, 1821 (
*** Herbertus aduncus (Dicks.) Gray, Nat. Arr. Brit. Pl. 1: 705, 1821 (
** Herbertus arcticus (Inoue et Steere) Schljakov, Novosti Sist. Nizš. Rast. 19: 209, 1982 (
** Herbertus armitanus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 324, 1965 (
** Herbertus asparus Tixier, Bryophyt. Biblioth. 18: 64, 1979 (
*** Herbertus bivittatus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 343, 1885 (
** Herbertus borealis Crundw., Trans. Brit. Bryol. Soc. 6 (1): 41, 1970 (
** Herbertus buchii Juslén, Ann. Bot. Fenn. 43 (6): 416, 2006 (
** Herbertus ceylanicus (Steph.) Abeyw., Ceylon J. Sci., Biol. Sci. 2 (1): 43, 1959 (
** Herbertus circinatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 318, 1965 (
*** Herbertus delavayi (Steph.) Steph., Hedwigia 34 (2): 43, 1895 (
*** Herbertus dicranus (Gottsche, Lindenb. et Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (
** Herbertus durandii (Steph.) Herzog, Rev. Bryol. Lichénol. 11 (1): 25, 1938 [1939] (
* Herbertus evittatus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965 (
** Herbertus gaochienii X.Fu, Fl. Bryoph. Sin. 9: 38, 2003 (
** Herbertus gracilis (Mont.) Steph., Bull. Herb. Boissier 5 (10): 842, 1897 (
** Herbertus guangdongii P.J.Lin ex Piippo, Bryobrothera 1: 206, 1992 (
** Herbertus hawaiiensis H.A.Mill., J. Hattori Bot. Lab. 28: 317, 1965 (
** Herbertus helleri (Steph.) W.E.Nicholson, Rev. Bryol. Lichénol. 13: 143, 1942 (
** Herbertus herpocladioides E.B.Scott et H.A.Mill., Bryologist 62 (2): 116, 1959 (
*** Herbertus hutchinsiae (Gottsche et Rabenh.) A.Evans, Bull. Torrey Bot. Club 44 (4): 214, 1917 (
*** Herbertus juniperoideus (Sw.) Grolle, Rev. Bryol. Lichénol. 30 (1/2): 80, 1961 (
*** Herbertus juniperoideus subsp. acanthelius (Spruce) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 326, 2006 (
*** Herbertus juniperoideus subsp. pensilis (Taylor) K.Feldberg et Heinrichs, Bot. J. Linn. Soc. 151: 329, 2006 (
** Herbertus kurzii (Steph.) R.S.Chopra, J. Indian Bot. Soc. 22: 247, 1943 (
** Herbertus leratii (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 327, 1965 (
** Herbertus lonchobasis H.A.Mill., J. Hattori Bot. Lab. 28: 306, 1965 (
** Herbertus longifissus Steph., Hedwigia 34 (2): 44, 1895 (
** Herbertus longispinus J.B.Jack et Steph., Hedwigia 31 (1): 15, 1892 (
** Herbertus mauritianus N.G.Hodgetts, J. Bryol. 30 (4): 247, 2008 (
*** Herbertus norenus D.G.Long, D.Bell et H.H.Blom, Molec. Ecol. Res. 12: 44, 2012 (
** Herbertus oldfieldianus (Steph.) Rodway, Tasm. Bryoph.: 72, 1917 (
** Herbertus pilifer Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 252, 1893 (
** Herbertus pocsii N.G.Hodgetts, J. Bryol. 30 (4): 249, 2008 (
** Herbertus pumilus Steph., Hedwigia 34 (2): 44, 1895 (
** Herbertus ramosus (Steph.) H.A.Mill., J. Hattori Bot. Lab. 28: 314, 1965 (
** Herbertus runcinatus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 397, 1877 (