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World checklist of hornworts and liverworts
expand article infoLars Söderström, Anders Hagborg§, Matt von Konrat§, Sharon Bartholomew-Began|, David Bell, Laura Briscoe§, Elizabeth Brown#, D. Christine Cargill¤, Denise Pinheiro da Costa«, Barbara J. Crandall-Stotler», Endymion Dante Cooper˄, Gregorio Dauphin˅, John Engel¦, Kathrin Feldbergˀ, David Glennyˁ, S. Robbert Gradstein, Xiaolan He, Jochen Heinrichs, Joern Hentschel, Anna Luiza Ilkiu-Borges, Tomoyuki Katagiri, Nadezhda A. Konstantinova, Juan Larraín¦, David Long, Martin Nebel, Tamás Pócs, Felisa Puche, Elena Reiner-Drehwald, Matt Renner#, Andrea Sass-Gyarmati, Alfons Schäfer-Verwimp, José Gabriel Segarra-Moragues‡‡, Raymond E. Stotler», Phiangphak Sukkharak§§, Barbara Thiers||, Jaime Uribe¶¶, Jiří Váňa##, Juan Villarreal¤¤, Martin Wigginton««, Li Zhang»», Rui-Liang Zhu˄˄
‡ NTNU, Trondheim, Norway
§ The Field Museum, Chicago, United States of America
| West Chester University, West Chester, United States of America
¶ University of British Columbia, Vancouver, Canada
# Royal Botanic Gardens and Domain Trust, Sydney, Australia
¤ Australian National Herbarium, Canberra, Australia
« Instituto de Pesquisas Jardim Botânico do Rio de Janeiro, Rio de Janeiro, Brazil
» Southern Illinois University, Carbondale, United States of America
˄ University of Maryland, Maryland, United States of America
˅ Unaffiliated, Acosta, Costa Rica
¦ Field Museum, Chicago, United States of America
ˀ Ludwig-Maximilians-Universität München, München, Germany
ˁ unknown, unknown, New Zealand
₵ Muséum National d’Histoire Naturelle, Paris, France
ℓ Finnish Museum of Natural History, University of Helsinki, Helsinki, Finland
₰ Ludwig-Maximilians-Universität Münchenen, München, Germany
₱ Friedrich-Schiller-University Jena, Jena, Germany
₳ Museu Paraense Emilio Goeldi, Belem, Brazil
₴ Hiroshima University, Hiroshima, Japan
₣ N.A. Avrorin Polar-Alpine Botanical Garden–Institute of Kola SC RAS, Kirovsk, Russia
₮ Royal Botanical Gardens, Edinburgh, United Kingdom
₦ Staatliches Museum für Naturkunde Stuttgart, Stuttgart, Germany
₭ Eszterházy Károly College, Eger, Hungary
₲ Universitat de València, Valencia, Spain
‽ University of Göttingen, Göttingen, Germany
₩ Royal Botanic Garden, Edinburgh, United Kingdom
₸ Unaffiliated, Herdwangen-Schönach, Germany
‡‡ Facultad de Ciencias Biológicas, Universidad de València, Burjassot (Valencia), Spain
§§ Burapha University, Mueang, Thailand
|| The New York Botanical Garden, Bronx, United States of America
¶¶ Universidad Nacional de Colombia, Bogotá, Colombia
## Charles University, Praha, Czech Republic
¤¤ Ludwig-Maximilians-Universität, München, Germany
«« Unaffiliated, Peterborough, United Kingdom
»» Fairylake Botanical Garden, Guangdong, China
˄˄ East China Normal University, Shanghai, China
† Deceased author
Open Access

Abstract

A working checklist of accepted taxa worldwide is vital in achieving the goal of developing an online flora of all known plants by 2020 as part of the Global Strategy for Plant Conservation. We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 7486 species in 398 genera representing 92 families from the two phyla. The checklist has far reaching implications and applications, including providing a valuable tool for taxonomists and systematists, analyzing phytogeographic and diversity patterns, aiding in the assessment of floristic and taxonomic knowledge, and identifying geographical gaps in our understanding of the global liverwort and hornwort flora. The checklist is derived from a working data set centralizing nomenclature, taxonomy and geography on a global scale. Prior to this effort a lack of centralization has been a major impediment for the study and analysis of species richness, conservation and systematic research at both regional and global scales. The success of this checklist, initiated in 2008, has been underpinned by its community approach involving taxonomic specialists working towards a consensus on taxonomy, nomenclature and distribution.

Keywords

Marchantiophyta , Anthocerophyta , nomenclature, taxonomy

This checklist is dedicated to Elizabeth Brown and Ray Stotler, who sadly left us during our work with it.

“……Charles Darwin wrote of his desire to provide financial support ‘for the formation of a perfect M.S. catalogue of all known plants’ (Darwin 1881, letter 13570). It is a personal embarrassment to me, and should be chastening to us all, that more than 120 years later we still have not delivered on that commitment.” (Crane 2004).

Introduction

The natural world is changing fast (Balmford et al. 2005). The Global Strategy for Plant Conservation (GSPC) was adopted by the Conference of Parties (COP) of the Convention on Biological Diversity (CBD) in April 2002 (http://www.cbd.int/decision/cop/?id=7183; accessed 2014.06.02). The Strategy set out 16 outcome-oriented targets that were to be achieved by 2010. The GSPC was designed as a framework for action to halt the loss of plant diversity. Target 1 of the Strategy was to complete “a widely accessible working list of all known plant species, as a step towards a complete world Flora” (Lughadha 2004). As early as 1881 Charles Darwin expressed a wish to have a catalogue of all known plants (Crane 2004). However, over 125 years later, this wish is not yet fulfilled. Without a working checklist, many of the other objectives in the GSPC cannot be met and botanists around the world cannot communicate about plants on a global basis (Crane 2004). A working list of known plant species is critical for (I) its underpinning role in the effective implementation of the other targets through provision of baseline information; (II) increasing the accessibility and use of accurate botanical name information for research, conservation and sustainable use; and (III) real-world politics and how taxonomists respond to the decisions of policy makers (Paton et al. 2008).

Version 1.0 of The Plant List (http://www.theplantlist.org/) was released in December 2010 aimed to be comprehensive for species of vascular plants (flowering plants, conifers, ferns and their allies) and of bryophytes (mosses, liverworts and hornworts), as a response to the 2010 Target 1 of the GSPC and to a clear global need for such data. The Plant List is a broad collaboration, coordinated by the Royal Botanic Gardens, Kew, and the Missouri Botanical Garden (MO), involving diverse partnerships. Paton (2013) noted The Plant List represents a work in progress with future versions planned to improve the quality of names and decrease the number of unresolved names. Target 1 has since been revised and agreed at the 10th COP of the CBD in Nagoya, Japan, with the goal of developing ‘‘an online flora of all known plants’’ by 2020 (http://www.cbd.int/decision/cop/default.shtml?id=12283; accessed 2014.06.02). Paton (2013) identified several factors that might assist in the implementation of the revised target, stressing that achieving a working list for plant taxa worldwide was a vital step to an online flora.

Earlier, Paton et al. (2008) assessed the progress made at that time and discussed prospects for the completion of Target 1. Paton noted that good progress had been made in bryophytes (mosses, liverworts and hornworts), ferns and gymnosperms with widely accessible working lists either complete or almost so for those groups. For bryophytes alone, he tabulated 13,370 accepted species noting that the data was largely derived from Tropicos. However, although Tropicos is an indispensable reference for anyone dealing with bryophyte names and the database is very strong for mosses, the nomenclatural and auxiliary data for liverworts is less complete, especially for larger genera (von Konrat et al. 2008a). The lack of a central source providing a synthesis of nomenclatural data and global distributional data was the impetus toward developing the current checklist of liverworts and hornworts (von Konrat et al. 2008a, 2010d, Söderström et al. 2012e). Checklists are powerful and important tools that can integrate the almost overwhelmingly scattered information concerning taxonomy, systematics, nomenclature, distribution, and even frequency (Söderström et al. 2008).

Ecological and biological significance of liverworts and hornworts

Liverworts and hornworts (Figure 1) are of critical biological, ecological, and phylogenetic significance (e.g. Asakawa 1998, Longton 1992, Hallingbäck and Hodgetts 2000, Gradstein et al. 2001b, Wellman et al. 2003, Qiu et al. 2007). Liverworts are found on soil, rocks, and trees throughout the world, from coastal Antarctica to the tundra of the Northern Hemisphere, and from semi-arid areas of Australia to the Amazon rainforests (Hallingbäck and Hodgetts 2000). Although there are xero-tolerant taxa, the majority of liverworts are found in relatively humid and shaded terrestrial ecosystems (Gradstein and Costa 2003). Liverworts and hornworts are an important component of the vegetation in many regions of the world, constituting a major part of the biodiversity in moist forest, wetlands, mountain, and tundra ecosystems (Hallingbäck and Hodgetts 2000).

Figure 1. 

Habit images of selected lineages of Anthocerotophyta (hornworts) and Marchantiophyta (liverworts). APhaeomegacerossquamuliger, photo by J. Hollinger BHaplomitriummnioides, photo by L. Zhang CConocephalumjaponicum, photo by D. G. Long DColuravitiensis, photo by T. Pócs EPleuroziagigantea, photo by L. Söderström FRiccardiaspegazziniana, photo by Juan Larraín.

Liverworts, hornworts and mosses offer microhabitats that are critical to the survival of a tremendous diversity of organisms such as single-celled eukaryotes, protozoa, and numerous groups of invertebrates (Gerson 1982). Their structural contribution to levels of diversity might be as significant as that of vascular plants, albeit at a smaller scale (von Konrat et al. 2008a). These plants are also important environmental and ecological indicators (Rao 1982, Pitcairn et al. 1995, Gradstein et al. 2001b, Giordano et al. 2004). Liverworts, in concert with mosses and hornworts, play a significant role in the global carbon budget (O’Neill 2000) and CO2 exchange (De Lucia et al. 2003), plant succession (Cremer and Mount 1965), net production and phytomass (Frahm 1990), nutrient cycling (Coxson 1991), and water retention (Pócs 1980a, Gradstein et al. 2001b). These groups of land plants also have been used as indicators of past climate change, to validate climate models, and as potential indicators of global warming (Gignac 2001, Tuba et al. 2011).

Species numbers

There are estimated to exist only 215 hornwort species. By comparison estimates of liverwort species richness have varied considerably, by as much as 50%, with estimates ranging from 4,500 to 9,000 (e.g. Pearson 1995, Forrest et al. 2006). In the last decade or so estimates in the range of 5,000 to 6,000 liverwort species have been widely accepted (e.g. Gradstein and Costa 2003, Gradstein and Ilkiu-Borges 2009, Heinrichs et al. 2007). von Konrat et al. (2010a) provided a mean estimate of 7,500 for the number of liverwort species based on estimating rates of synonymy in a sample of recently monographed and revised taxa. A standardized global worldwide liverwort checklist with strong community participation, as presented here, coupled with the critical need for ongoing monographs and revisions, will aid in arriving at clearer estimates of liverwort and hornwort diversity. Significantly, a list of standardized names reviewed by taxonomic experts will enable more meaningful geographical and biogeographical species comparative studies. The global checklist of accepted liverwort and hornwort taxa has vast potential, not only in aiding our understanding of liverwort diversity, patterns, and processes, but also to the broader biological community (von Konrat et al. 2008a).

Below we provide context to the worldwide checklist, including a brief historical account and current informational resources, structure and layout, systematic and classification concepts, and methodology, including a brief overview of the underlying data set and nomenclatural elements.

Major historical works

Schuster (1966d) has provided a detailed account on the history of hepaticology, up to the first half of the 20th century. Earlier, Verdoorn (1934c) also outlined a valuable historical review of important figures in hepaticology. In the early 19th century, possibly the two most influential persons who provided an early framework for taxonomic concepts were W.J. Hooker and G. Raddi. During the middle of the 19th century C.G. Nees von Esenbeck, C.M. Gottsche and J.B.W. Lindenberg provided vastly significant contributions towards developing a classification framework. Frahm and Eggers (2001) provided a brief biography of these workers. This was later followed by R. Spruce and the Austrian-born H. Leitgeb in the second half 19th century. Leitgeb in particular was instrumental in liverwort plant morphology and anatomy including a six part treatise, “Untersuchungen ueber die Lebermoose”, from 1874 to 1881. Towards the end of 19th century and early 20th century V. Schiffner, F. Stephani, and A. Evans were prominent hepaticologists. In the first half of the 20th century A. Evans, H. Buch, F. Verdoorn, and K. Müller (Frib.) were leading influential hepaticologists. In the second half of the 20th century, R. Grolle and R.M. Schuster were of extraordinary influence in working towards contemporary classification schemes. Many of these groundbreaking concepts still serve the foundation from which our current classification scheme is derived. R.M. Schuster’s major contribution to botany and hepaticology lies in the astounding new diversity of liverworts he added to our knowledge. He described a staggering 463 species, 83 genera and 15 families new to science. It is difficult to name another contemporary botanist who discovered this much new diversity of a major clade of land plants (Qiu et al. 2013). R.M. Schuster’s contribution to botany went beyond hepatics. He was one of the first botanists who recognized the importance of Wallace’s Line in plant biogeography, separating Australia of Gondwanaland from Southeast Asia of Laurasia (Cronquist 1988).

Figure 2 depicts the number of novel liverwort species, excluding new combinations that have been described over the last 250 years. The first major peak corresponds to the works of several early 19th century botanists, including Synopsis Hepaticarum by Gottsche, Lindenberg, and Nees (1844–1847). The three decades leading into, but prior to the highest peak, between 1860 and 1890, correspond to publications by a number of prominent bryologists including W. Mitten, J.D. Hooker, T. Taylor, and V. Schiffner. The second and highest peak of almost 1,200 names, in the early 1900s corresponds largely to the plethora of taxa described by F. Stephani (1898–1924) in his monumental work Species Hepaticarum. The periods of highest rates of new species described in the 1830s and around 1900 are the same for seed plants (Mutke and Barthlott 2005). The third peak over the four decades between 1950 and 1980 can be attributed mainly to the works of R.M. Schuster, H. Inoue, and S. Hattori. H. Inoue and S. Hattori were influential on large species-rich genera such as Plagiochila, Frullania, and Porella. The decline in newly described species since 1970 does not necessarily translate to the conclusion that taxonomists are closer to discovering all known species. The almost 200 novel liverwort species that have been described in the past six years alone still represent a significant number, considering the relatively few liverwort taxonomists and monographers. Moreover, bryological exploration has been very uneven in many parts of the world. For example, many areas of the Neotropics still remain without a single bryophyte record (Gradstein et al. 2001a). Paradoxically, scores of new species are still being discovered and described in relatively well-studied areas such as New Zealand, e.g. over 50 new taxa since 2000 alone (e.g. Engel and Schuster 2001, Engel and Glenny 2008a, Renner and de Lange 2011, von Konrat et al. 2012b). Recent attention to cryptic speciation in bryophytes is also revealing novel liverwort species (e.g. Szweykowski et al. 2005). The combination of collecting in yet-to-be explored areas, the continued discovery of species in well-studied regions, and an increased understanding of the biology of liverworts (including cryptic speciation), will lead to a significant number of newly discovered species in the foreseeable future. The corollary of this, coupled with increased monographic and revisional work, will be the increased discovery and the unravelling of synonymy.

Figure 2. 

Number of novel liverwort species, excluding new combinations, which have been described over the last 250 years, with an inset of the number described from 2001–2012.

Contemporary resources

Data availability and information needs associated with liverworts and hornworts have been reviewed extensively by von Konrat et al. (2008a, 2010a). A fundamental problem common to all nomenclatural indexing projects is the dispersed nature of the biological literature, some of which may date back over 250 years (Lughadha 2004). Our own data records, from 1990 to the present alone, indicate there are over 190 periodical titles and non-serials in which liverwort nomenclatural novelties were published. The most useful and successful web-accessible bryophyte nomenclatural database is Tropicos (http://www.tropicos.org), which offers name data with references and type information as well as links to specimen data of its holdings. Tropicos has therefore become an indispensable reference for anyone dealing with bryophyte names. However, as stated above, the database is particularly strong for mosses and far less comprehensive for liverworts. Moreover, Tropicos does not prescribe taxonomic disposition of names, i.e., there is no attempt to adopt a single consistent view on the status of any particular name (Lughadha 2004, von Konrat et al. 2008a).

A major nomenclatural work is the Index Hepaticarum (http://www.ville-ge.ch/musinfo/bd/cjb/hepatic/index.php), which includes all effectively published liverwort epithets spanning 12 volumes with the closing date of 1973. The indices were prepared as a purely nomenclatural resource and did not claim to express any particular taxonomic concept. Recently, Crosby and Engel (2006) provided an equally valuable nomenclatural resource and catalog of names at all ranks for liverworts and hornworts published during 1974 to 2000. Subsequently, indices of the citations for names published for bryophytes have been compiled for the years 2001-2004 (Crosby and Magill 2005) and 2005 (Crosby and Magill 2006). The Early Land Plants Today initiative has continued this series (von Konrat et al. 2010c, Söderström et al. 2012b, 2014).

The application of molecular phylogenetics continues to generate new insights into the evolutionary history of liverworts and hornworts. In recent years, inferences made from these phylogenies have especially revolutionized liverwort classification (Crandall-Stotler et al. 2009). Yet, the reconstruction of the phylogenetic history of the Marchantiophyta and Anthocerotophyta remains an ongoing effort and the classification is fluid (von Konrat et al. 2010a). The current checklist largely follows the comprehensive phylogenetic liverwort classification scheme provided by Crandall- Stotler et al. (2009) and the hornwort classification scheme of Stotler and Crandall-Stotler (2005). Thus the checklist largely adheres to their higher classification namely from class to suborder, and the majority of family and genera arrangements provided therein. However, there are some notable exceptions, especially in the systematic treatment of liverworts, which largely either reflect publications subsequent to 2009, or are a slight departure in taxonomic opinion based on earlier works, e.g. the concepts of Saccogynaceae and Stephaniellaceae depart from Crandall-Stotler et al. (2009). Since 2010 there have been a number of realignments and novel hypotheses proposed that were generated from conclusions based on molecular evidence which have influenced the systematic treatment presented here. These include, for example, Adelanthaceae (e.g. Konstantinova and Vilnet 2009, Feldberg et al. 2010b, 2010a, Söderström et al. 2010b, Vilnet et al. 2010), Anastrophyllaceae (e.g. Söderström et. al 2010b), Gymnomitriaceae (e.g. Vilnet et al 2007a, 2010, Váňa et al. 2010b, Shaw et al. 2015), Lepidoziaceae (Cooper 2013), Lophocoleaceae (Söderström et al. 2013b), and Scapaniaceae (e.g. Vilnet et al. 2010, Heinrichs et al. 2012a). In all cases, any such departures, new families, or realignments are noted and very briefly discussed throughout the checklist. As noted by Crandall-Stotler et al. (2009) many small families still remain to be investigated using molecular data, and many of the large families and genera that have been sampled appear to be either polyphyletic or paraphyletic.

The systematic treatment as outlined here also includes the addition of subfamily and infrageneric classification; ranks not presented by Crandall-Stotler et al. (2008b, 2009). Although there are arguably a number of ambiguous placements, especially at the infrageneric rank, and there remain many areas of contention, particularly at the familial and generic level, the systematic treatment itself serves as an excellent synopsis of contemporary taxonomic opinion, including infrageneric ranks. The infrageneric treatments will also aid future efforts in providing a taxonomic framework that will be particularly useful for extremely large genera. In large part, the infrageneric treatments only deal with those taxa that represent phylogenetic clades with strong support. Care has been taken to reduce ambiguity and those taxa where infrageneric placement is weak or without evidence are simply listed under the genus and not placed systematically.

Structure and format of the checklist

The checklist is presented in a taxonomic framework as outlined above with four main sections. The first is arranged systematically with taxa ordered alphabetically within the nearest higher rank. A brief discussion for selected families is provided in the main body of the text under the respective families. This does not appear uniformly for all families, but typically for larger more complicated families. Rather, a brief description is provided mainly where concepts depart from Crandall-Stotler et al. (2009) or if new information has come to light since that publication. Subfamilies and infrageneric ranks have been incorporated where appropriate. However, if the subgeneric placement is unclear the species are listed under “Incertae sedis”. From the rank of genus and below, all entries include a three star confidence ranking (described below), accepted taxon name, authority, abbreviated publication title, page number and date, which is cross- referenced to the full citation in the reference list, as well as the basionym if appropriate. A typical entry appears as follows:

*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b).

*** Oleolophoziaperssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (Söderström et al. 2010b). Bas.: Lophoziaperssonii H.Buch et S.W.Arnell, Bot. Not. 97: 382, 1944 (Buch 1944).

There are comprehensive footnotes throughout the systematic section of the checklist. These are applied to selected names representing the rank of genus and below only. Footnotes without any literature citation represent observations from the authors of the respective family. Footnotes are wide ranging in their content. Generally, footnotes can be categorized into the following: i) general statements relating to taxonomic opinion, ii) comments on the type specimen, iii) comments about possible conspecificity, including citation of conflicting opinions, and iv) comment about species complexes.

This section is followed by a list of taxa in genera that we do not recognize. Those names represent taxa that have not been studied recently and have not been recognized in any recent treatment. However, many of those names are old and may gain priority over some recognized taxa once their identity is determined.

The next section is an alphabetical list of taxa of the ranks of species and below. The alphabetical list includes the confidence level, the taxon name and authority as well as a reference to the page of the corresponding entry in the systematic section. Thus, the alphabetical list provides a rapid gateway requiring no prior knowledge of higher taxonomy, but also serves as an index to the corresponding name in the systematic section that includes more detailed information.

The last section is a reference list where full bibliographic citations are given to all references for taxa included in the checklist. Titles and references have all been checked and verified, except in a few cases where we could not get hold of the publications.

Taxon confidence levels

Significantly, each accepted taxon is qualified using a three level ranking system that summarizes our knowledge about a taxon. The coding convention we are adopting largely follows that described by von Konrat et al. (2010d) using one to three stars, which has been applied to recent regional checklists produced by the Early Land Plants Today initiative, e.g. Söderström et al. (2010a, 2011a, 2011b, 2012g, 2013e). The conventions are briefly outlined below coupled with samples illustrating how these conventions are applied in practice. The application of a confidence level to a taxon’s status and whether it represents a genuine “species” that is reached through community consensus may go towards refining species estimates using an evidence-based approach (von Konrat et al. 2008a, 2008b). The confidence levels coupled with the detailed annotations in the form of footnotes may also provide a rapid assessment of taxa and help aid and drive future research into specific taxonomic or nomenclatural problems and issues.

The coding convention:

* Serious doubts. There are doubts about the value of the taxon. It can be that there are conflicting views without any substantial evidence in any direction, conflicting views with substantiating evidence supporting one or both positions, or evidence pointing towards synonymization but it is premature to do it. We have adhered to the principle that it is better to keep a taxon with one star (and preferably a note) than to synonymize it too quickly. Example: Bazzaniaasymmetrica is conspecific with Bazzaniamacgregorii in Grolle (1968a), but Kitagawa (1979a) kept them separate. Example: Nardiakamtschatica may be conspecific with Nardiaassamica (Váňa 1976c), but the type specimen could not be studied.

** Knowledge problem. The taxon is not well known by the person evaluating it. It may be a newly described species or a species originally not well described and not restudied recently. Example: Jungermanniaerectii Ajit P.Singh et V.Nath was recently described and has not been independently studied by someone with a global overview. It is therefore difficult to evaluate.

*** Accepted. A good taxon as currently understood based on personal experience or on taxonomic revisions that have been convincingly performed. Nomenclature and/or taxonomic position may, however, be questioned. Elements may be excluded from the taxon, but the taxon with the current type will still be accepted. Example: H. Bischler-Causse revised the genus Marchantia worldwide in a series of publications. Although she had a broad species concept, she also recognized infraspecific taxa. The taxa that she recognized without doubt should be accepted unless new evidence against it exists. Adopting a narrower species concept so that many of her subspecies are elevated to species does not change her view of what a good taxon is.

Methodology

The foundation of the checklist is in the underlying data set from which it is derived. It was briefly described by von Konrat et al. (2008a, 2010a). The working data sets now includes a bibliography of 25,000 publications; approximately 39,000 published liverwort names (including “accepted” taxa, synonyms, invalid and illegitimate names). The data quality and standards were outlined by von Konrat et al. (2010d). In summary, for authorities and for the citation abbreviations, we follow the standards set by the on-line version of Authors of Plant Names at the Royal Botanical Gardens, Kew Website (http://www.ipni.org), with whom we collaborate closely and provide with updated data records. Publications and journal abbreviations follow Taxonomic Literature: A selective guide to botanical publications and collections with dates, commentaries and types (Stafleu and Cowan 1976–2009) (http://www.sil.si.edu/digitalcollections/tl-2/search.cfm) and Botanico Periodicum HuntianumLawrence et al. (1968) (http://huntbot.andrew.cmu.edu/HIBD/Departments/Databases.shtml).

There has been an intense systematic effort focusing on data quality. In all but a few cases nomenclatural data has been verified against original publications. Söderström and Hagborg have checked and confirmed almost every original publication (three publications have not been available) for correct author, title and journal/book citation, date of publication as far as possible, page number for the protologue, and if the name is validly and legitimately published according to the International Code of Nomenclature for algae, fungi and plants (Melbourne Code; McNeill et al. 2012). Most significantly, the checklist has been community driven and collaborative. Broader participation by taxonomic specialists and regional experts has lead to the checklist containing high quality data (cf. Söderström et al. 2008). Development of the checklist has included three international meetings (the first one in 2009) and generated 74 published notes on taxonomy and nomenclature under the auspices of the Early Land Plants Today initiative from 2012 to 2015. The series of meetings were instrumental in providing a framework for direct interaction with taxonomic experts, workshops reviewing names, and helping identify potential participants who were taxonomic experts in specific taxonomic groups, whether these were individual genera or entire families.

The notes, published in the journal Phytotaxa, have included 38 authors from 13 countries. These are the results of several years of revisions of taxonomic groups by taxonomic experts. Following these revisions and working with participants identifying nomenclatural and taxonomic problems that would impact the worldwide checklist, Söderström, Hagborg and von Konrat led editorial efforts in compiling manuscripts with baseline information and nomenclatural or taxonomic issues that required resolving. The work was then coordinated with the taxonomic experts who drove and led the process and often identified further issues. The series of notes on taxonomy and nomenclature of liverworts and hornworts provided updated nomenclature, corrected invalid and illegitimate names and described new taxa based on studies (mainly molecular) that did not draw nomenclatural conclusions. Those notes are all open access and any changes were effective immediately as the Melbourne Code of Botanical Nomenclature allows publishing on internet from 1 Jan. 2012 (Knapp et al. 2011).

This checklist builds on all published taxonomic and nomenclatural papers, that have come to our attention until June 30, 2015.

Summary statistics

We here present the first-ever worldwide checklist for liverworts (Marchantiophyta) and hornworts (Anthocerotophyta) that includes 8,078 taxa (species and below) in 7,486 species representing 398 genera, 92 families, 20 orders and 7 classes from the two phyla. The list includes 3,533 species with three stars, 2,988 species with two stars and 915 species with one star. The checklist also has extra utility in that it contains 3,106 references in the bibliography that serve as a powerful bibliographic resource for liverwort and hornwort systematic and taxonomic research.

Concluding remarks

The marked-up publication form of the current checklist by PhytoKeys provides a virtual instrument with a linked environment both internally (e.g. within an article) and externally (GBIF, IPNI, Tropicos, Wikispecies, etc.) that will undoubtedly help accelerate taxonomic research. The published checklist was the first phase in providing a worldwide list of accepted names. The next phase is to establish a generally recognized online repository to augment the huge underlying informational auxiliary data of over 25,000 publications, almost 39,000 published names, and the over 700,000 geographical observations. Several features of Web-based technology make it an essential tool, including 1) a vehicle to facilitate data access, 2) to unify the vastly scattered data on distributional information and nomenclature, 3) offering dynamic rather than static data in a searchable forum. The broader accessibility to the wealth of auxiliary data will help augment monographic and revisional work for many taxonomic groups, aid in identifying the need for increased floristic and survey work in many regions throughout the world, and have broad implications and applications beyond taxonomic research such as conservation science. The current Early Land Plants Today model has strong participation from taxonomic experts and an online resource will provide an opportunity to expand stakeholders to include ecologists, conservationists, scientists from other disciplines and general interest groups. However, such an effort can only be successful if it comes with sustained funding and infrastructure rather than depending on an ad hoc commitment by a few individuals, however dedicated.

The project will help augment monographic and revisional work for many taxonomic groups and aid in identifying the need for increased floristic and survey work in many regions throughout the world. Although there are many challenges ahead to obtain high quality data, quantifying global liverwort diversity is a tractable, multi-faceted and scientifically important goal, and everyone stands to gain by fostering this endeavour. The success of the project will lie on strong collaboration between institutions and the bryological community in general.

Taxonomic list

Anthocerotophyta

Anthocerotopsida de Bary ex Jancz

Anthocerotidae Rosenv

Anthocerotales Limpr
*** Anthocerotaceae Dumort

by J.C. Villarreal and D.C. Cargill

Notes on nomenclature and taxonomy can also be found in Cargill et al. (2013b) and Villarreal et al. (2015)

*** Anthoceros L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753). 1

*** Anthocerosadscendens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 24, 1832 (Lehmann 1832).

*** Anthocerosagrestis Paton, J. Bryol. 10 (3): 257, 1979 (Paton 1979a), nom. conserv. 2

*** Anthocerosalpinus Steph., Sp. Hepat. (Stephani) 6: 425, 1923 (Stephani 1923).

* Anthocerosangustifolius Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 582, 1846 (Gottsche et al. 1846). 3

*** Anthocerosangustus Steph., Sp. Hepat. (Stephani) 5: 1001, 1916 (Stephani 1916b).

*** Anthocerosbharadwajii Udar et A.K.Asthana, Proc. Indian Natl. Sci. Acad., B 51 (4): 484, 1985 (Udar and Asthana 1985b).

** Anthocerosbuettneri Steph., Sp. Hepat. (Stephani) 5: 997, 1916 (Stephani 1916b).

*** Anthoceroscapricornii Cargill et G.A.M.Scott, J. Hattori Bot. Lab. 82: 55, 1997 (Cargill and Scott 1997).

*** Anthoceroscaucasicus Steph., Izv. Kavkazsk. Muz. 8: 87, 1914 (Voronov’ 1914).

** Anthoceroscavernosus Steph., Sp. Hepat. (Stephani) 5: 998, 1916 (Stephani 1916b).

* Anthoceroschambensis Kashyap, J. Bombay Nat. Hist. Soc. 25 (2): 281, 1917 (Kashyap 1917).

** Anthoceroschungii Khanna, J. Indian Bot. Soc. 17 (5/6): 316, 1938 (Khanna 1938).

** Anthoceroscrispatus Griff., Not. pl. asiat. 2: 349, 1849 (Griffith 1849).

** Anthocerosdimorphus Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926 (Sim 1926).

*** Anthoceroserectus Kashyap, New Phytol. 14 (1): 9, 1915 (Kashyap 1915).

** Anthocerosexpansus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 92, 2015 (Villarreal et al. 2015). Bas.: Aspiromitusexpansus Steph., Sp. Hepat. (Stephani) 5: 961, 1916 (Stephani 1916b).

* Anthocerosferdinandi-muelleri Steph., Sp. Hepat. (Stephani) 5: 1007, 1916 (Stephani 1916b). 4

*** Anthocerosfragilis Steph., Sp. Hepat. (Stephani) 5: 1006, 1916 (Stephani 1916b).

*** Anthocerosfusiformis Austin, Bull. Torrey Bot. Club 6 (4): 28, 1875 [1876] (Austin 1875a).

*** Anthocerosfusiformisvar.taiwanensis J.Haseg., Acta Phytotax. Geobot. 44 (2): 100, 1993 (Hasegawa 1993b).

** Anthocerosgasongorii Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (Gola 1916).

** Anthocerosgranulatus Gottsche, Mexik. Leverm.: 275, 1863 (Gottsche 1863).

** Anthocerosharrisanus (Steph.) Parihar, Univ. Allahabad Stud., Bot. 1961-2: 31, 1962 (Parihar 1962). Bas.: Aspiromitusharrisanus Steph., Sp. Hepat. (Stephani) 5: 965, 1916 (Stephani 1916b).

* Anthoceroshelmsii Steph., Hedwigia 32 (3): 142, 1893 (Stephani 1893b). 5

** Anthocerosjamesonii Taylor ex Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 171, 1855 (Mitten 1855).

* Anthocerosjavanicoides H.A.Mill., Phytologia 47 (4): 319, 1981 (Miller 1981). Nom. nov. pro Anthoceros javanicus Steph., Sp. Hepat. (Stephani) 5: 988, 1916 (Stephani 1916b), nom. illeg. 6

** Anthocerosjungermannioides Schwein., Spec. Fl. Amer. Crypt.: 25, 1821 (Schweinitz 1821).

* Anthoceroskajumas (K.I.Goebel) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Aspiromituskajumas K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 69, 1928 (Goebel 1928).

* Anthoceroskoshii Khanna, J. Indian Bot. Soc. 15 (4): 237, 1936 (Khanna 1936).

*** Anthoceroslamellatus Steph., Sp. Hepat. (Stephani) 5: 1000, 1916 (Stephani 1916b).

*** Anthoceroslaminifer Steph., J. Linn. Soc., Bot. 29 (201): 266, 1892 (Stephani 1892b).

*** Anthocerosmacounii M.Howe, Bull. Torrey Bot. Club 25 (1): 19, 1898 (Howe 1898a).

*** Anthocerosmacrosporus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (Stephani 1916b).

** Anthocerosmaritimus Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (Stephani 1916b).

* Anthocerosmegasporus Meijer, J. Hattori Bot. Lab. 18: 6, 1957 (Meijer 1957).

** Anthocerosmuscoides Colenso, Trans. & Proc. New Zealand Inst. 16: 361, 1884 (Colenso 1884).

*** Anthocerosmyriandroecius Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 134, 1911 (Stephani 1911a).

** Anthocerosnatalensis Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (Stephani 1913b).

*** Anthocerosneesii Prosk., Leberm. Eur. 2 (9): 1312, 1958 (Proskauer 1958). Nom. nov. pro Anthocerospunctatus α* monocarpus Nees, Naturgesch. Eur. Leberm. 4: 339, 1838 (Nees 1838a).

** Anthocerosniger Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (Stephani 1916b).

** Anthocerosorizabensis (Steph.) Hässel, Candollea 45 (1): 211, 1990 (Hässel 1990b). Bas.: Aspiromitusorizabensis Steph., Sp. Hepat. (Stephani) 5: 965, 1916 (Stephani 1916b).

*** Anthocerospandei Udar et A.K.Asthana, J. Indian Bot. Soc. 64: 305, 1985 (Udar and Asthana 1985a).

*** Anthocerospatagonicus Hässel, Candollea 45 (1): 207, 1990 (Hässel 1990b).

*** Anthocerospatagonicussubsp.gremmenii J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 32, 2013 (Váňa and Engel 2013).

** Anthocerosperuvianus Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (Stephani 1916b).

** Anthocerospinnatus Steph., Bol. Soc. Brot. 4: 154 [182], 1885 [1886] (Stephani 1885c).

*** Anthocerospunctatus L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753). 7

* Anthocerospusillus Colenso, Trans. & Proc. New Zealand Inst. 18: 255, 1886 (Colenso 1886b).

*** Anthocerosrosulans J.Haseg., J. Hattori Bot. Lab. 60: 379, 1986 (Hasegawa 1986b).

*** Anthocerossambesianus Steph., Sp. Hepat. (Stephani) 5: 996, 1916 (Stephani 1916b).

*** Anthocerosscariosus Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 230, 1869 (Austin 1869).

** Anthocerosschroederi Steph., 52 (5): 306, 1912 (Stephani 1912a).

** Anthocerosserratus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911 (Stephani 1911b).

** Anthocerossimulans M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 204, 1934 (Howe 1934).

** Anthocerosspongiosus Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (Stephani 1916b).

*** Anthocerossubtilis Steph., Sp. Hepat. (Stephani) 5: 1003, 1916 (Stephani 1916b).

*** Anthocerostelaganus Steph., Sp. Hepat. (Stephani) 5: 1005, 1916 (Stephani 1916b).

*** Anthocerostristanianus J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 33, 2013 (Váňa and Engel 2013).

*** Anthocerostuberculatus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832 (Lehmann 1832).

** Anthocerosvenosus Lindenb. et Gottsche, Syn. Hepat. 4: 584, 1846 (Gottsche et al. 1846).

Excluded from the genus

* Anthocerosaethyopicus Gola, Ann. Bot. (Rome) 13 (1): 73, 1914 (Gola 1914a). 8

* Anthocerosbrunnthaleri Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 88: 732, 1913 (Stephani 1913b). 9

* Anthocerosfloribundus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (Stephani 1916b). 10

* Anthocerosmildbraedii Steph., Sp. Hepat. (Stephani) 6: 428, 1923 (Stephani 1923). 11

* Anthocerosparvifrons Steph., 52 (5): 307, 1912 (Stephani 1912a). 12

* Anthocerospseudocostus Steph., Sp. Hepat. (Stephani) 5: 977, 1916 (Stephani 1916b). 13

* Anthocerosrossoi Gola, Mem. Reale Accad. Sci. Torino (ser. 2) 65 (1): 10, 1916 (Gola 1916). 14

*** Folioceros D.C.Bharadwaj, Geophytology 1 (1): 9, 1971 (Bharadwaj 1971).

*** Foliocerosamboinensis (Schiffn.) Piippo, Acta Bot. Fenn. 148: 36, 1993 (Piippo 1993b). Bas.: Anthocerosamboinensis Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 45, 1890 (Schiffner 1890).

** Foliocerosapiahynus (Steph.) Hässel, Candollea 45 (1): 215, 1990 (Hässel 1990b). Bas.: Anthocerosapiahynus Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (Stephani 1916b).

** Foliocerosargillaceus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (Villarreal et al. 2015). Bas.: Aspiromitusargillaceus Steph., Sp. Hepat. (Stephani) 5: 970, 1916 (Stephani 1916b).

*** Foliocerosassamicus D.C.Bharadwaj, Geophytology 1 (1): 10, 1971 (Bharadwaj 1971).

** Foliocerosdilatatus (Steph.) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 93, 2015 (Villarreal et al. 2015). Bas.: Anthocerosdilatatus Steph., Bot. Jahrb. Syst. 8 (2): 95, 1886 (Stephani 1886d).

*** Foliocerosdixitianus (Mahab.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Aspiromitusdixitianus Mahab., Curr. Sci. 10 (12): 532, 1941 (Mahabale 1941).

*** Foliocerosfuciformis (Mont.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthocerosfuciformis Mont., Ann. Sci. Nat. Bot. (sér. 2) 20: 296, 1844 (Montagne 1844b).

*** Foliocerosglandulosus (Lehm. et Lindenb.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthocerosglandulosus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 26, 1832 (Lehmann 1832).

*** Foliocerosincurvus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthocerosincurvus Steph., Hedwigia 32 (3): 143, 1893 (Stephani 1893b).

*** Foliocerosindicus D.C.Bharadwaj, Geophytology 8 (1): 114, 1978 (Bharadwaj 1978).

*** Folioceroskashyapii S.C.Srivast. et A.K.Asthana, Bryologist 92 (2): 219, 1989 (Srivastava and Asthana 1989).

*** Foliocerosmangaloreus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Aspiromitusmangaloreus Steph., Sp. Hepat. (Stephani) 5: 967, 1916 (Stephani 1916b).

*** Foliocerospaliformis D.K.Singh, Bull. Bot. Surv. India 29: 176, 1987 [1989] (Singh 1987b).

*** Foliocerosphysocladus D.C.Bharadwaj, Geophytology 8 (1): 115, 1978 (Bharadwaj 1978). Based on: Anthocerosphysocladus Schiffn. ex Pandé, Proc. Indian Sci. Congr. Assoc. 47 (2): 98, 1960 (Pandé 1960), nom. inval.

*** Foliocerospinnilobus (Steph.) D.C.Bharadwaj, Geophytology 5 (2): 227, 1975 (Bharadwaj 1975). Bas.: Anthocerospinnilobus Steph., Akad. Wiss. Wien, Math.-Naturwiss. Kl., Denkschr. 81: 299, 1907 (Stephani 1907a).

*** Foliocerossatpurensis D.C.Bharadwaj et K.P.Srivast., Geophytology 8 (1): 112, 1978 (Bharadwaj 1978). Based on: Anthocerossatpurensis K.P.Srivast., Proc. Indian Sci. Congr. Assoc. 47 (3): 337, 1960 (Srivastava 1960), nom. inval.

*** Foliocerosudarii A.K.Asthana et S.C.Srivast., Cryptog. Bryol. Lichénol. 7 (2): 151, 1986 (Asthana and Srivastava 1986).

*** Foliocerosverruculosus (J.Haseg.) R.L.Zhu et M.J.Lai, Ann. Bot. Fenn. 48 (5): 383, 2011 (Wang et al. 2011). Bas.: Anthocerosverruculosus J.Haseg., Acta Phytotax. Geobot. 44 (2): 103, 1993 (Hasegawa 1993b).

Dendrocerotidae R.J.Duff, J.C.Villarreal, Cargill et Renzaglia

Dendrocerotales Hässel
*** Dendrocerotaceae J.Haseg

by J.C. Villarreal and D.C. Cargill

*** Dendroceros Nees, Syn. Hepat. 4: 579, 1846 (Gottsche et al. 1846).

** subg.Apoceros R.M.Schust., Phytologia 63 (3): 200, 1987 (Schuster 1987b).

*** Dendroceroscavernosus J.Haseg., J. Hattori Bot. Lab. 47: 306, 1980 (Hasegawa 1980).

*** Dendrocerosdifficilis Steph., Sp. Hepat. (Stephani) 5: 1009, 1917 (Stephani 1917b).

*** Dendrocerosmuelleri Steph., Hedwigia 28 (2): 133, 1889 (Stephani 1889a).

*** Dendrocerosogeramnangus Piippo, Acta Bot. Fenn. 148: 40, 1993 (Piippo 1993b).

*** Dendrocerospedunculatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (Stephani 1909c).

** Dendrocerossubdifficilis S.Hatt., Bot. Mag. (Tokyo) 64 (755/756): 119, 1951 (Hattori 1951c).

** subg.Dendroceros

*** Dendrocerosacutilobus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (Stephani 1909c).

*** Dendrocerosborbonicus Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 32 (2): 31, 1893 [1894] (Stephani 1893e).

*** Dendroceroscrispus (Sw.) Nees, Syn. Hepat. 4: 581, 1846 (Gottsche et al. 1846). Bas.: Anthoceroscrispus Sw., Prodr. (Swartz): 146, 1788 (Swartz 1788).

*** Dendrocerosfoliicola J.Haseg., J. Hattori Bot. Lab. 47: 296, 1980 (Hasegawa 1980).

*** Dendrocerosjaponicus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 15, 1909 (Stephani 1909c).

*** Dendrocerosjavanicus (Nees) Nees, Syn. Hepat. 4: 582, 1846 (Gottsche et al. 1846). Bas.: Anthocerosjavanicus Nees, Enum. Pl. Crypt. Javae: 1, 1830 (Nees 1830).

*** Dendrocerossubplanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 20, 1909 (Stephani 1909c).

*** Dendrocerostubercularis S.Hatt., Bot. Mag. (Tokyo) 58 (685): 6, 1944 (Hattori 1944b).

*** Dendrocerosvalidus Steph., Sp. Hepat. (Stephani) 5: 1016, 1917 (Stephani 1917b).

** Dendrocerosvesconianus Gottsche, J. Bot. (Morot) 12: 150, 1898 (Bescherelle 1898).

** Dendroceroswattsianus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (Stephani 1909c).

Incertae sedis

* Dendrocerosadglutinatus (Hook.f. et Taylor) Gottsche, Lindenb. et Nees, Syn. Hepat. 4: 580, 1846 (Gottsche et al. 1846). Bas.: Monocleaadglutinata Hook.f. et Taylor, London J. Bot. 4: 96, 1845 (Hooker and Taylor 1845). 15

*** Dendrocerosafricanus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 18, 1909 (Stephani 1909c).

** Dendrocerosallionii Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (Stephani 1917b).

** Dendrocerosaustralis Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 17, 1909 (Stephani 1909c).

** Dendrocerosbreutelii Nees, Syn. Hepat. 4: 581, 1846 (Gottsche et al. 1846).

** Dendrocerosbreuteliivar.surinamensis Lindenb. et Gottsche, Linnaea 24 (6): 639, 1851 [1852] (Lindenberg and Gottsche 1851a).

*** Dendroceroscichoraceus (Mont.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 16, 1858 (Gottsche 1858). Bas.: Anthoceroscichoraceus Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 355, 1845 (Montagne 1845b).

* Dendroceroscrassicostatus Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (Stephani 1917b).

** Dendroceroscrassinervis (Nees) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 18, 1858 (Gottsche 1858). Bas.: Anthoceroscrassinervis Nees, Syn. Hepat. 4: 589, 1846 (Gottsche et al. 1846).

*** Dendroceroscrispatus (Hook.) Nees, Syn. Hepat. 4: 579, 1846 (Gottsche et al. 1846). Bas.: Monocleacrispata Hook., Bot. Misc. 1: 117, 1830 (Hooker 1830).

** Dendroceroscrispatusvar.simplicior Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 574, 1885 (Spruce 1885).

** Dendroceroscucullatus Steph., Sp. Hepat. (Stephani) 6: 429, 1923 (Stephani 1923). 16

* Dendrocerosexalatus Steph., Sitzungsber. Naturf. Ges. Leipzig 36: 14, 1909 (Stephani 1909c). 17

* Dendrocerosgracilis Steph., Sp. Hepat. (Stephani) 5: 1015, 1917 (Stephani 1917b).

*** Dendrocerosgranulatus Mitt., Fl. vit.: 419, 1871 [1873] (Mitten 1871).

** Dendrocerosherasii M.Infante, J. Bryol. 32 (4): 285, 2010 (Infante 2010).

** Dendroceroshumboldtensis Hürl., Bauhinia 1 (3): 254, 1960 (Hürlimann 1960).

** Dendrocerospaivae C.A.Garcia, Sérgio et J.C.Villarreal, Cryptog. Bryol. 33 (1): 5, 2012 (Garcia et al. 2012).

* Dendrocerosrarus Steph., Sp. Hepat. (Stephani) 5: 1014, 1917 (Stephani 1917b).

* Dendrocerosreticulus Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 37, 1950 [1951] (Herzog 1950b). 18

** Dendrocerosrigidus Steph., Sp. Hepat. (Stephani) 5: 1017, 1917 (Stephani 1917b).

*** Dendrocerosseramensis J.Haseg., Acta Phytotax. Geobot. 37 (1/3): 10, 1986 (Hasegawa 1986a).

** Dendrocerossubtropicus C.J.Wild, Trans. Nat. Hist. Soc. Queensland 1: 49, 1893 (Wild 1893).

** Dendrocerostahitensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 138, 1873 (Ångström 1873).

*** Megaceros Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (Campbell 1907).

** Megacerosaneuriformis Steph., Sp. Hepat. (Stephani) 5: 949, 1916 (Stephani 1916b).

*** Megacerosaustronesophilus Cargill et Seppelt, Austral. Syst. Bot. 26 (5): 372, 2013 (Cargill et al. 2013a).

** Megacerosciliatus K.I.Goebel, Ann. Jard. Bot. Buitenzorg 39: 74, 1928 (Goebel 1928).

*** Megacerosdenticulatus (Lehm.) Steph., Sp. Hepat. (Stephani) 5: 956, 1916 (Stephani 1916b). Bas.: Anthocerosdenticulatus Lehm., Nov. Stirp. Pug. 10: 25, 1857 (Lehmann 1857).

*** Megacerosflagellaris (Mitt.) Steph., Sp. Hepat. (Stephani) 5: 951, 1916 (Stephani 1916b). Bas.: Anthocerosflagellaris Mitt., Fl. vit.: 419, 1871 [1873] (Mitten 1871).

*** Megacerosgracilis (Reichardt) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (Stephani 1916b). Bas.: Anthocerosgracilis Reichardt, Verh. K.K. Zool.-Bot. Ges. Wien 16: 957, 1866 (Reichardt 1866).

** Megacerosleptohymenius (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 5: 955, 1916 (Stephani 1916b). Bas.: Monoclealeptohymenia Hook.f. et Taylor, London J. Bot. 3: 575, 1844 (Hooker and Taylor 1844d).

*** Megacerospellucidus (Colenso) E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 115, 1972 (Hodgson 1972). Bas.: Anthocerospellucidus Colenso, Trans. & Proc. New Zealand Inst. 17: 263, 1885 (Colenso 1885).

** Megacerostjibodensis Campb., Ann. Bot. (Oxford) 21 (4): 484, 1907 (Campbell 1907).

Excluded from the genus 19

* Megacerosflavens (Spruce) Campb., Ann. Bot. (Oxford) 21 (4): 483, 1907 (Campbell 1907). Bas.: Anthocerosflavens Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 575, 1885 (Spruce 1885).

* Megacerosjamesonii (Taylor) Steph., Biblioth. Bot. 87 (2): 268, 1916 (Stephani 1916a). Bas.: Dendrocerosjamesonii Taylor, London J. Bot. 7: 285, 1848 (Taylor 1848b).

*** Nothoceros (R.M.Schust.) J.Haseg., J. Hattori Bot. Lab. 76: 32, 1994 (Hasegawa 1994b). Bas.: Megacerossubg.Nothoceros R.M.Schust., Phytologia 63 (3): 200, 1987 (Schuster 1987b).

*** Nothocerosaenigmaticus J.C.Villarreal et K.D.McFarland, Bryologist 113 (1): 109, 2010 (Villarreal et al. 2010b). Based on: Megacerosaenigmaticus R.M.Schust., Hepat. Anthocerotae N. Amer. 6: 830, 1992 (Schuster 1992d), nom. inval.

*** Nothoceroscanaliculatus (Pagán) J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (Villarreal et al. 2007). Bas.: Dendroceroscanaliculatus Pagán, Bryologist 45 (4): 111, 1942 (Pagán 1942a).

*** Nothocerosendiviifolius (Mont.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (Villarreal et al. 2007). Bas.: Anthocerosendiviifolius Mont., Voy. Pole Sud, Bot. 1: 211, 1845 (Montagne 1845c).

** Nothocerosfuegiensis (Steph.) J.C.Villarreal, Bryologist 113 (1): 109, 2010 (Villarreal et al. 2010b). Bas.: Megacerosfuegiensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 91, 1911 (Stephani 1911b).

*** Nothocerosgiganteus (Lehm. et Lindenb.) J.Haseg. ex J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 283, 2007 (Villarreal et al. 2007). Bas.: Anthocerosgiganteus Lehm. et Lindenb., Nov. Stirp. Pug. 4: 25, 1832 (Lehmann 1832).

*** Nothocerosminarum (Nees) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (Villarreal and Renner 2014). Bas.: Anthocerosminarum Nees, Naturgesch. Eur. Leberm. 4: 340, 1838 (Nees 1838a).

** Nothocerosrenzagliensis J.C.Villarreal, L.V.Campos et Uribe, Syst. Bot. 37 (1): 32, 2012 (Villarreal et al. 2012).

*** Nothocerosschizophyllus (Steph.) J.C.Villarreal, Molec. Phylogen. Evol. 78: 34, 2014 (Villarreal and Renner 2014). Bas.: Megacerosschizophyllus Steph., Sp. Hepat. (Stephani) 5: 949, 1916 (Stephani 1916b).

*** Nothocerossuperbus J.C.Villarreal, Hässel et N.Salazar, Bryologist 110 (2): 280, 2007 (Villarreal et al. 2007).

*** Nothocerosvincentianus (Lehm. et Lindenb.) J.C.Villarreal, Bryologist 113 (1): 111, 2010 (Villarreal et al. 2010b). Bas.: Anthocerosvincentianus Lehm. et Lindenb., Nov. Stirp. Pug. 6: 16, 1834 (Lehmann 1834).

** Phaeomegaceros R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duff et al. 2007).

*** Phaeomegaceroscoriaceus (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duff et al. 2007). Bas.: Anthoceroscoriaceus Steph., Sp. Hepat. (Stephani) 5: 991, 1916 (Stephani 1916b).

*** Phaeomegacerosfimbriatus (Gottsche) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duff et al. 2007). Bas.: Anthocerosfimbriatus Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 187, 1864 (Gottsche 1864).

*** Phaeomegacerosfoveatus (J.Haseg.) J.C.Villarreal, Nova Hedwigia 91 (3/4): 352, 2010 (Villarreal et al. 2010a). Bas.: Phaeocerosfoveatus J.Haseg., Bryol. Res. 7 (12): 374, 2001 (Hasegawa 2001).

*** Phaeomegaceroshirticalyx (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duff et al. 2007). Bas.: Aspiromitushirticalyx Steph., Sp. Hepat. (Stephani) 5: 966, 1916 (Stephani 1916b).

*** Phaeomegacerosplicatus (Mitt.) J.C.Villarreal, J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 85, 2013 (Váňa and Engel 2013). Bas.: Anthocerosplicatus Mitt., Rep. Challenger, Bot. 1 (2): 178, 1884 (Mitten 1884a).

*** Phaeomegacerosskottsbergii (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 241, 2007 (Duff et al. 2007). Bas.: Anthocerosskottsbergii Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 90, 1911 (Stephani 1911b).

*** Phaeomegacerossquamuliger (Spruce) J.C.Villarreal, Nova Hedwigia 91 (3/4): 351, 2010 (Villarreal et al. 2010a). Bas.: Anthocerossquamuliger Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 576, 1885 (Spruce 1885).

** Phaeomegacerossquamuligersubsp.hasselii J.C.Villarreal, Cargill et Goffinet, Nova Hedwigia 91 (3/4): 352, 2010 (Villarreal et al. 2010a).

Phymatocerotales R.J.Duff, J.C.Villarreal, Cargill et Renzaglia
*** Phymatocerotaceae R.J.Duff, J.C.Villarreal, Cargill et Renzaglia

by J.C. Villarreal and D.C. Cargill

*** Phymatoceros Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 113, 2005 (Stotler et al. 2005).

*** Phymatocerosbulbiculosus (Brot.) Stotler, W.T.Doyle et Crand.-Stotl., Phytologia 87 (2): 114, 2005 (Stotler et al. 2005). Bas.: Anthocerosbulbiculosus Brot., Fl. lusit. 2: 430, 1804 [1805] (Brotero 1804).

*** Phymatocerosphymatodes (M.Howe) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia, Bryologist 110 (2): 240, 2007 (Duff et al. 2007). Bas.: Anthocerosphymatodes M.Howe, Bull. Torrey Bot. Club 25 (1): 12, 1898 (Howe 1898a).

Notothylatidae R.J.Duff, J.C.Villarreal, Cargill et Renzaglia

Notothyladales Hyvönen et Piippo
*** Notothyladaceae Müll.Frib. ex Prosk

by J.C. Villarreal and D.C. Cargill

** Notothyladoideae Grolle

*** Notothylas Sull. ex A.Gray, Amer. J. Sci. Arts (ser. 2) 1 (1): 74, 1846 (Gray 1846).

** subg.Notothylas

*** Notothylasanaporata Udar et D.K.Singh, Rev. Bryol. Lichénol. 45 (2): 202, 1979 (Udar and Singh 1979b).

*** Notothylasbreutelii (Gottsche) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 21, 1858 (Gottsche 1858). Bas.: Anthocerosbreutelii Gottsche, Syn. Hepat. 4: 583, 1846 (Gottsche et al. 1846).

*** Notothylasdepressispora J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 26, 1979 (Hasegawa 1979).

*** Notothylasdissecta Steph., Sp. Hepat. (Stephani) 5: 1020, 1917 (Stephani 1917b).

*** Notothylashimalayensis Udar et D.K.Singh, J. Bryol. 11 (3): 451, 1981 (Udar and Singh 1981a).

*** Notothylasindica Kashyap, Proc. Lahore Philos. Soc. 4: 54, 1925 (Kashyap and Dutt 1925).

*** Notothylasjavanica (Sande Lac.) Gottsche, Bot. Zeitung (Berlin) Beil. 16: 20, 1858 (Gottsche 1858). Bas.: Blasiajavanica Sande Lac., Syn. hepat. jav.: 94, 1856 [1857] (Sande Lacoste 1856b).

*** Notothylasorbicularis (Schwein.) Sull., Amer. J. Sci. Arts (ser. 2) 1 (1): 75, 1846 (Gray 1846). Bas.: Targioniaorbicularis Schwein., Spec. Fl. Amer. Crypt.: 23, 1821 (Schweinitz 1821).

*** Notothylaspandei Udar et V.Chandra, Geophytology 7 (2): 142, 1977 (Udar and Chandra 1977).

** subg.Notothyloides A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 106, 1991 (Asthana and Srivastava 1991).

*** Notothylaskhasiana Udar et D.K.Singh, J. Indian Bot. Soc. 60: 112, 1981 (Udar and Singh 1981b).

*** Notothylaspfleidereri Udar et D.K.Singh, Lindbergia 5 (1): 28, 1979 (Udar and Singh 1979a).

Incertae sedis

*** Notothylasdecurva (Mitt.) Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (Stephani 1901d). Bas.: Anthocerosdecurvus Mitt., Trans. Linn. Soc. London 23 (1): 58, 1860 (Mitten 1860a).

*** Notothylasflabellata Steph., Cat. Afr. Pl. (Hiern) 2 (2): 320, 1901 (Stephani 1901d).

*** Notothylasgalapagensis M.Howe, Proc. Calif. Acad. Sci. (ser. 4) 21 (17): 203, 1934 (Howe 1934).

** Notothylasirregularis Chantanaorr., Acta Bot. Hung. 56 (3/4): 270, 2014 (Chantanaorrapint 2014).

*** Notothylaskashyapii D.K.Singh, Indian J. Forest. 23 (4): 386, 2000 (Singh and Semwal 2000).

*** Notothylasnepalensis D.K.Singh, J. Bombay Nat. Hist. Soc. 84 (3): 650, 1987 [1988] (Singh 1987a).

*** Notothylastemperata J.Haseg., Acta Phytotax. Geobot. 30 (1/3): 20, 1979 (Hasegawa 1979).

*** Notothylasudarii D.K.Singh et Semwal, Phytotaxonomy 1: 35, 2001 (Singh and Semwal 2001).

* Notothylasverdoornii Khanna, Rev. Bryol. Lichénol. 6: 118, 1933 (Khanna 1933).

*** Notothylasvitalii Udar et D.K.Singh, Misc. Bryol. Lichenol. 8 (9): 173, 1980 (Udar and Singh 1980).

** Notothylasyunnanensis T.Peng et R.L.Zhu, Phytotaxa 156 (3): 157, 2014 (Peng and Zhu 2014).

** Phaeocerotoideae Hässel

* Mesoceros Piippo, Acta Bot. Fenn. 148: 30, 1993 (Piippo 1993b). 20

* Mesocerosmesophoros Piippo, Acta Bot. Fenn. 148: 30, 1993 (Piippo 1993b).

* Mesocerosporcatus Piippo, Haussknechtia, Beih. 9: 279, 1999 (Piippo 1999).

** Paraphymatoceros Hässel, Phytologia 88 (2): 208, 2006 (Hässel 2006a).

*** Paraphymatocerosdiadematus Hässel, Phytologia 88 (2): 209, 2006 (Hässel 2006a).

*** Paraphymatoceroshallii (Austin) Hässel, Phytologia 88 (2): 209, 2006 (Hässel 2006a). Bas.: Anthoceroshallii Austin, Bull. Torrey Bot. Club 6 (4): 26, 1875 [1876] (Austin 1875a).

*** Paraphymatocerospearsonii (M.Howe) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (Villarreal et al. 2015). Bas.: Anthocerospearsonii M.Howe, Bull. Torrey Bot. Club 25 (1): 8, 1898 (Howe 1898a).

*** Paraphymatocerosproskaueri (Stotler, Crand.-Stotl. et W.T.Doyle) J.C.Villarreal et Cargill, Phytotaxa 208 (1): 94, 2015 (Villarreal et al. 2015). Bas.: Phaeocerosproskaueri Stotler, Crand.-Stotl. et W.T.Doyle, Fieldiana, Bot. (n.ser.) 47: 232, 2008 (Crandall-Stotler et al. 2008a).

*** Phaeoceros Prosk., Bull. Torrey Bot. Club 78 (4): 346, 1951 (Proskauer 1951a).

** Phaeocerosaustroandinus Hässel, Candollea 44 (2): 721, 1989 (Hässel 1989a).

** Phaeocerosbolusii (Sim) S.W.Arnell, Hepat. South Africa: 401, 1963 (Arnell 1963b). Bas.: Anthocerosbolusii Sim, Trans. Roy. Soc. South Africa 15 (1): 114, 1926 (Sim 1926).

** Phaeocerosbrevicapsulus (Steph.) Hässel, Candollea 44 (2): 725, 1989 (Hässel 1989a). Bas.: Anthocerosbrevicapsulus Steph., Sp. Hepat. (Stephani) 5: 981, 1916 (Stephani 1916b).

*** Phaeoceroscarolinianus (Michx.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Anthoceroscarolinianus Michx., Fl. bor.-amer. (Michaux) 2: 280, 1803 (Michaux 1803).

*** Phaeocerosdelicatus E.O.Campb. et Outred, New Zealand J. Bot. 33 (3): 285, 1995 (Campbell and Outred 1995).

*** Phaeocerosdendroceroides (Steph.) Hässel, Beih. Nova Hedwigia 134: 487, 2009 (Hässel and Rubies 2009). Bas.: Anthocerosdendroceroides Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (Stephani 1916b). 21

*** Phaeocerosengelii Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 248, 2008 (Cargill and Fuhrer 2008).

** Phaeoceroserectus Udar et D.K.Singh, Geophytology 11 (2): 257, 1981 (Udar and Singh 1981c). Nom. nov. pro Anthoceros erectus Steph., Sp. Hepat. (Stephani) 5: 984, 1916 (Stephani 1916b), nom. illeg.

*** Phaeocerosevanidus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 245, 2008 (Cargill and Fuhrer 2008). Bas.: Anthocerosevanidus Steph., Sp. Hepat. (Stephani) 5: 990, 1916 (Stephani 1916b).

*** Phaeocerosexiguus (Steph.) J.Haseg., J. Hattori Bot. Lab. 60: 387, 1986 (Hasegawa 1986b). Bas.: Anthocerosexiguus Steph., Sp. Hepat. (Stephani) 5: 988, 1916 (Stephani 1916b).

*** Phaeocerosflexivalvis (Gottsche et Nees) Hässel, Candollea 44 (2): 728, 1989 (Hässel 1989a). Bas.: Anthocerosflexivalvis Gottsche et Nees, Syn. Hepat. 4: 586, 1846 (Gottsche et al. 1846).

*** Phaeocerosfulvisporus (Steph.) J.Haseg., Trop. Bryol. 8: 52, 1993 (Hasegawa 1993a). Bas.: Anthocerosfulvisporus Steph., 52 (5): 306, 1912 (Stephani 1912a).

*** Phaeocerosgemmifer (Horik.) J.Haseg., J. Hattori Bot. Lab. 57: 252, 1984 (Hasegawa 1984). Bas.: Anthocerosgemmifer Horik., Sci. Rep. Tôhoku Imp. Univ., Ser. 4, Biol. 4 (2): 426, 1929 (Horikawa 1929a).

** Phaeocerosgualaquizanus (Steph.) Gradst., J. Hattori Bot. Lab. 45: 123, 1979 (Gradstein and Hekking 1979). Bas.: Anthocerosgualaquizanus Steph., Sp. Hepat. (Stephani) 5: 979, 1916 (Stephani 1916b).

*** Phaeoceroshimalayensis (Kashyap) Prosk. ex Bapna et G.G.Vyas, J. Hattori Bot. Lab. 25: 88, 1962 (Bapna and Vyas 1962). Bas.: Anthoceroshimalayensis Kashyap, New Phytol. 14 (1): 8, 1915 (Kashyap 1915).

*** Phaeoceroshuebschmannii Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 301, 1986 (Hässel 1986b).

*** Phaeocerosinflatus (Steph.) Cargill et Fuhrer, Fieldiana, Bot. (n.ser.) 47: 246, 2008 (Cargill and Fuhrer 2008). Bas.: Anthocerosinflatus Steph., Sp. Hepat. (Stephani) 5: 990, 1916 (Stephani 1916b).

*** Phaeoceroskashyapii A.K.Asthana et S.C.Srivast., Bryophyt. Biblioth. 42: 129, 1991 (Asthana and Srivastava 1991).

*** Phaeoceroslaevis (L.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Anthoceroslaevis L., Sp. Pl. 1: 1139, 1753 (Linnaeus 1753).

** Phaeocerosmaranguensis (Steph.) Bapna, Univ. Udaipur Res. Stud. 3: 138, 1966 (Bapna 1966). Bas.: Aspiromitusmaranguensis Steph., Sp. Hepat. (Stephani) 5: 960, 1916 (Stephani 1916b).

* Phaeocerosmicrosporus (Steph.) Hässel, Candollea 44 (2): 721, 1989 (Hässel 1989a). Bas.: Aspiromitusmicrosporus Steph., Sp. Hepat. (Stephani) 5: 963, 1916 (Stephani 1916b). 22

*** Phaeocerosminutus (Mitt.) S.W.Arnell, Hepat. South Africa: 403, 1963 (Arnell 1963b). Bas.: Anthocerosminutus Mitt., J. Linn. Soc., Bot. 16 (91): 195, 1877 (Mitten 1877).

*** Phaeocerosmohrii (Austin) Hässel, Candollea 44 (2): 721, 1989 (Hässel 1989a). Bas.: Anthocerosmohrii Austin, Bull. Torrey Bot. Club 6 (52): 304, 1879 (Austin 1879).

*** Phaeocerosoreganus (Austin) Hässel, Candollea 44 (2): 718, 1989 (Hässel 1989a). Bas.: Anthocerosoreganus Austin, Bull. Torrey Bot. Club 6 (4): 26, 1875 [1876] (Austin 1875a).

*** Phaeocerosparvulus (Schiffn.) J.Haseg., J. Hattori Bot. Lab. 57: 248, 1984 (Hasegawa 1984). Bas.: Anthocerosparvulus Schiffn., Österr. Bot. Z. 49 (11): 391, 1899 (Schiffner 1899c).

*** Phaeocerosperpusillus Chantanaorr., Acta Bot. Hung. 51 (1/2): 30, 2009 (Chantanaorrapint 2009).

** Phaeocerospichinchensis (Spruce) Hässel, Candollea 44 (2): 725, 1989 (Hässel 1989a). Bas.: Anthocerospichinchensis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 577, 1885 (Spruce 1885).

** Phaeocerospropagulifer (Steph.) Prosk., Bull. Torrey Bot. Club 78 (4): 347, 1951 (Proskauer 1951a). Bas.: Anthocerospropagulifer Steph., Sp. Hepat. (Stephani) 5: 1001, 1916 (Stephani 1916b).

* Phaeocerosstriatisporus J.Haseg., J. Hattori Bot. Lab. 75, 268 (Hasegawa 1994a).23

*** Phaeocerostenuis (Spruce) Hässel, Veröff. Geobot. Inst. ETH Stiftung Rübel Zürich 91: 303, 1986 (Hässel 1986b). Bas.: Anthocerostenuis Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: cxcvi, 1889 [1890] (Spruce 1889).

** Phaeocerostigrinus (Gola) J.C.Villarreal, Phytotaxa 208 (1): 95, 2015 (Villarreal et al. 2015). Bas.: Anthocerostigrinus Gola, Ann. Bot. (Rome) 13 (1): 72, 1914 (Gola 1914a).

*** Phaeocerostuberosus (Taylor) Prosk., J. Indian Bot. Soc. 42A: 185, 1964 (Proskauer 1964). Bas.: Anthocerostuberosus Taylor, London J. Bot. 5: 412, 1846 (Taylor 1846b).

*** Phaeocerosudarii A.K.Asthana et V.Nath, Proc. Natl. Acad. Sci. India, B 63 (4): 461, 1993 (Asthana and Nath 1993).

** Phaeoceroswrightii (Steph.) Hässel, Candollea 44 (2): 722, 1989 (Hässel 1989a). Bas.: Anthoceroswrightii Steph., Sp. Hepat. (Stephani) 5: 999, 1916 (Stephani 1916b).

Leiosporocerotopsida Stotler et Crand.-Stotl

Leiosporocerotales Hässel

*** Leiosporocerotaceae Hässel ex Ochyra

by J.C. Villarreal and D.C. Cargill

*** Leiosporoceros Hässel, J. Bryol. 14 (2): 255, 1986 (Hässel 1986a).

*** Leiosporocerosdussii (Steph.) Hässel, J. Bryol. 14 (2): 255, 1986 (Hässel 1986a). Bas.: Anthocerosdussii Steph., Hedwigia 32 (3): 142, 1893 (Stephani 1893b).

Marchantiophyta

Haplomitriopsida Stotler et Crand.-Stotl

Haplomitriidae Stotler et Crand.-Stotl

Calobryales Hamlin
*** Haplomitriaceae Dědeček

by S. Bartholomew-Began

The treatment of Haplomitriaceae follows Bartholomew-Began (1991). Morphogenetic and molecular studies support the infrageneric ranks of Haplomitrium (Crandall-Stotler et al. 2009, Forrest et al. 2006).

*** Haplomitrium Nees, Naturgesch. Eur. Leberm. 1: 109, 1833 (Nees 1833c) nom. conserv.

*** subg.Calobryum (Nees) R.M.Schust., Nova Hedwigia 13 (1/2): 40, 1967 (Schuster 1967c). Bas.: Calobryum Nees, Syn. Hepat. 4: 507, 1846 (Gottsche et al. 1846).

*** Haplomitriumblumei (Nees) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (Schuster 1963b). Bas.: Monocleablumei Nees, Enum. Pl. Crypt. Javae: 2, 1830 (Nees 1830).

*** Haplomitriummnioides (Lindb.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (Schuster 1963b). Bas.: Rhopalanthusmnioides Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 391, 1874 (Lindberg 1874a).

*** subg.Haplomitrium

** sect.Archibryum (R.M.Schust.) J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 675, 1981 (Engel 1981). Bas.: Haplomitriumsubg.Archibryum R.M.Schust., Nova Hedwigia 13 (1/2): 28, 1967 (Schuster 1967c).

*** Haplomitriumgibbsiae (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 225, 1963 (Schuster 1963b). Bas.: Calobryumgibbsiae Steph., Sp. Hepat. (Stephani) 6: 76, 1917 (Stephani 1917a).

*** Haplomitriumintermedium Berrie, Proc. Linn. Soc. New South Wales (ser. 2) 87 (399): 191, 1963 (Berrie 1963).

** sect.Haplomitrium

*** Haplomitriumhookeri (Lyell ex Sm.) Nees, Naturgesch. Eur. Leberm. 1: 111, 1833 (Nees 1833c). Bas.: Jungermanniahookeri Lyell ex Sm., Engl. Bot. 35: tab. 2555, 1814 (Smith and Sowerby 1814).

*** Haplomitriumhookerivar.minutum (E.O.Campb.) Barthol.-Began, Bryophyt. Biblioth. 41: 230, 1991 (Bartholomew-Began 1991). Bas.: Steereomitriumminutum E.O.Campb., Mem. New York Bot. Gard. 45: 569, 1987 (Campbell 1987).

*** Haplomitriummonoicum J.J.Engel, Ann. Missouri Bot. Gard. 68 (4): 668, 1981 [1982] (Engel 1981).

*** Haplomitriumovalifolium R.M.Schust., Bryologist 74 (2): 136, 1971 (Schuster 1971d).

Treubiidae Stotler et Crand.-Stotl

Treubiales Schljakov
*** Treubiaceae Verd

by R. Stotler and B.J. Crandall-Stotler

Stech et al. (2002) showed a clear molecular distinction between Treubia and Apotreubia, which is in accordance with their morphological differences and supports their recognition as separate genera.

*** Apotreubia S.Hatt. et Mizut., Bryologist 69 (4): 491, 1966 [1967] (Hattori et al. 1966).

*** Apotreubiahortoniae Konstant., Phytotaxa 76 (3): 33, 2013 (Konstantinova et al. 2013b). Based on: Apotreubiahortoniae R.M.Schust. et Konstant., J. Hattori Bot. Lab. 78: 55, 1995 (Schuster and Konstantinova 1995), nom. inval.

*** Apotreubianana (S.Hatt. et Inoue) S.Hatt. et Mizut., Bryologist 69 (4): 492, 1966 [1967] (Hattori et al. 1966). Bas.: Treubianana S.Hatt. et Inoue, J. Hattori Bot. Lab. 11: 99, 1954 (Hattori and Inoue 1954).

** Apotreubiapusilla (R.M.Schust.) Grolle, Acta Bot. Fenn. 125: 63, 1984 (Grolle and Piippo 1984). Bas.: Treubiapusilla R.M.Schust., Nova Hedwigia 15: 515, 1968 (Schuster 1968b).

*** Apotreubiayunnanensis Higuchi, Cryptog. Bryol. Lichénol. 19 (4): 321, 1998 (Higuchi 1998).

*** Treubia K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (Goebel 1890) nom. conserv.

*** Treubiainsignis K.I.Goebel, Ann. Jard. Bot. Buitenzorg 9 (1): 1, 1890 [1891] (Goebel 1890).

** Treubiainsignissubsp.bracteata (Steph.) R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 241, 1969 (Schuster and Scott 1969). Bas.: Treubiabracteata Steph., Bot. Jahrb. Syst. 23 (1/2, 3): 302, 1896 (Stephani 1896a).

** Treubiainsignissubsp.caledonica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 243, 1969 (Schuster and Scott 1969).

** Treubiainsignissubsp.vitiensis R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 242, 1969 (Schuster and Scott 1969).

*** Treubialacunosa (Colenso) Prosk., Bryologist 58 (3): 199, 1955 (Proskauer 1955). Bas.: Noterocladalacunosa Colenso, Trans. & Proc. New Zealand Inst. 18: 248, 1886 (Colenso 1886b).

** Treubialacunosoides T.Pfeiff., W.Frey et M.Stech, Nova Hedwigia 75 (1/2): 249, 2002 (Pfeiffer et al. 2002).

*** Treubiapygmaea R.M.Schust., Phytologia 56 (7): 460, 1985 (Schuster 1985c).

*** Treubiascapanioides R.M.Schust., J. Hattori Bot. Lab. 32: 246, 1969 (Schuster and Scott 1969).

*** Treubiatahitensis (Nadeaud) Besch., J. Bot. (Morot) 12: 147, 1898 (Bescherelle 1898). Bas.: Gottscheatahitensis Nadeaud, Énum. Pl. Tahiti: 7, 1873 (Nadeaud 1873).

*** Treubiatasmanica R.M.Schust. et G.A.M.Scott, J. Hattori Bot. Lab. 32: 248, 1969 (Schuster and Scott 1969).

Jungermanniopsida Stotler et Crand.-Stotl

Jungermanniidae Engl

Jungermanniales H.Klinggr
Cephaloziineae Schljakov
*** Adelanthaceae Grolle

by K. Feldberg, J. Váňa and J. Heinrichs

The subfamily Jamesonielloideae was excluded from Lophoziaceae/Scapaniaceae by De Roo et al. (2007) and this status was confirmed by e.g. Vilnet et al. (2010). The family was described and defined by Feldberg et al. (2010a). Some taxonomic and nomenclatural notes can also be found in Feldberg et al. (2010b, 2011), Váňa et al. (2014d).

*** Adelanthoideae K.Feldberg, Heinrichs et Váňa

*** Adelanthus Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (Mitten 1864b) nom. conserv.

** sect.Adelanthus

*** Adelanthusfalcatus (Hook.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 243, 1864 (Mitten 1864b). Bas.: Jungermanniafalcata Hook., Musci Exot. 1: tab. 89, 1818 (Hooker 1818).

*** Adelanthusocclusus (Hook.f. et Taylor) Carrington, Trans. Bot. Soc. Edinburgh 10: 381, 1870 (Carrington 1870). Bas.: Jungermanniaocclusa Hook.f. et Taylor, London J. Bot. 3: 369, 1844 (Hooker and Taylor 1844a).

** sect.Calyptrocolea (R.M.Schust.) Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (Grolle 1972c). Bas.: Calyptrocolea R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 685, 1966 (Schuster 1966a).

*** Adelanthusaureomarginatus R.M.Schust., Phytologia 39 (4): 250, 1978 (Schuster 1978a).

*** Adelanthusgemmiparus (R.M.Schust.) E.A.Hodgs., Trans. Roy. Soc. New Zealand, Biol. Sci. 11 (18): 241, 1970 (Hodgson 1970). Bas.: Calyptrocoleagemmipara R.M.Schust., Rev. Bryol. Lichénol. 34 (3/4): 695, 1966 [1967] (Schuster 1966a).

** Adelanthuslingulatus J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 26, 2013 (Váňa and Engel 2013).

*** Adelanthustenuis J.J.Engel et Grolle, J. Hattori Bot. Lab. 35: 333, 1972 (Grolle 1972c).

** sect.Lindenbergiani Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (Grolle 1972c).

*** Adelanthuscarabayensis (Mont.) Grolle, J. Hattori Bot. Lab. 35: 348, 1972 (Grolle 1972c). Bas.: Plagiochilacarabayensis Mont., Ann. Sci. Nat. Bot. (sér. 4) 5: 348, 1856 (Montagne 1856c).

*** Adelanthusintegerrimus Grolle, J. Hattori Bot. Lab. 35: 340, 1972 (Grolle 1972c).

*** Adelanthuslindenbergianus (Lehm.) Mitt., J. Proc. Linn. Soc., Bot. 7 (28): 244, 1864 (Mitten 1864b). Bas.: Jungermannialindenbergiana Lehm., Linnaea 4: 367, 1829 (Lehmann 1829).

** sect.Pittieri Grolle, J. Hattori Bot. Lab. 35: 331, 1972 (Grolle 1972c).

*** Adelanthuspittieri (Steph.) Grolle, J. Hattori Bot. Lab. 35: 337, 1972 (Grolle 1972c). Bas.: Tylimanthuspittieri Steph., Sp. Hepat. (Stephani) 6: 250, 1922 (Stephani 1922).

*** Adelanthussquarrosus Grolle, J. Hattori Bot. Lab. 67: 243, 1989 (Grolle 1989d).

*** Pseudomarsupidium Herzog, Svensk Bot. Tidskr. 47 (1): 42, 1953 (Herzog 1953b).

*** Pseudomarsupidiumaureocinctum (R.M.Schust.) J.J.Engel, Novon 17 (3): 312, 2007 (Engel 2007). Bas.: Adelanthusdecipienssubsp.aureocinctus R.M.Schust., Phytologia 39 (4): 250, 1978 (Schuster 1978a).

*** Pseudomarsupidiumborneensis (Grolle) Váňa, L.Söderstr., A.Hagborg et von Konrat, Phytotaxa 65: 60, 2012 (Váňa et al. 2012f). Bas.: Adelanthusborneensis Grolle, J. Hattori Bot. Lab. 35: 362, 1972 (Grolle 1972c).

*** Pseudomarsupidiumdecipiens (Hook.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (Grolle 1963a). Bas.: Jungermanniadecipiens Hook., Brit. Jungermann.: tab. 50, 1813 (Hooker 1813).

*** Pseudomarsupidiumpiliferum (Steph.) Herzog ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 443, 1963 (Grolle 1963a). Bas.: Marsupidiumpiliferum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 602 (386), 1908 (Stephani 1908e).

*** Wettsteinia Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 44, 1898 (Schiffner 1898c).

*** Wettsteiniadensiretis (Herzog) Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (Grolle 1965f). Bas.: Tylimanthusdensiretis Herzog, Nat. Hist. Juan Fernandez (Botany) 2 (5): 712, 1942 (Herzog 1942a).

*** Wettsteiniainversa (Sande Lac.) Schiffn., Ann. Jard. Bot. Buitenzorg, suppl. 2: 45, 1898 (Schiffner 1898c). Bas.: Plagiochilainversa Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 289, 1864 (Sande Lacoste 1864).

*** Wettsteiniarotundifolia (Horik.) Grolle, J. Hattori Bot. Lab. 28: 100, 1965 (Grolle 1965f). Bas.: Adelanthusrotundifolius Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 181, 1934 (Horikawa 1934).

*** Wettsteiniaschusteriana Grolle, J. Hattori Bot. Lab. 28: 99, 1965 (Grolle 1965f).

** Jamesonielloideae Inoue

*** Cuspidatula Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1141 (124), 1901 (Stephani 1901c).

*** Cuspidatulacontracta (Reinw., Blume et Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1141 (124), 1901 (Stephani 1901c). Bas.: Jungermanniacontracta Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 233, 1824 [1825] (Reinwardt et al. 1824a).

*** Cuspidatulaflaccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (Feldberg et al. 2010b). Bas.: Anastrophyllumflaccidum Steph., Sp. Hepat. (Stephani) 6: 105, 1917 (Stephani 1917a).

*** Cuspidatulaflexicaulis (Nees) Váňa et L.Söderstr., Phytotaxa 76 (3): 35, 2013 (Váňa et al. 2013h). Bas.: Jungermanniaflexicaulis Nees, Linnaea 6 (4): 604, 1831 (Nees 1831).

*** Cuspidatulakirkii (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 142, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniellakirkii Steph., Hedwigia 34 (2): 47, 1895 (Stephani 1895c).

*** Cuspidatulamonodon (Taylor) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1143 (126), 1901 (Stephani 1901c). Bas.: Jungermanniamonodon Taylor, Nov. Stirp. Pug. 8: 7, 1844 (Lehmann 1844).

*** Cuspidatulaorbicularis (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (Váňa et al. 2013h). Bas.: Jamesoniellaorbicularis Grolle, Feddes Repert. 82 (1): 42, 1971 (Grolle 1971b).

*** Cuspidatularobusta (Austin) Váňa et L.Söderstr., Phytotaxa 76 (3): 36, 2013 (Váňa et al. 2013h). Bas.: Jungermanniarobusta Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 219, 1869 (Austin 1869).

** Denotarisia Grolle, Feddes Repert. 82 (1): 6, 1971 (Grolle 1971b).

*** Denotarisialinguifolia (De Not.) Grolle, Feddes Repert. 82 (1): 6, 1971 (Grolle 1971b). Bas.: Plagiochilalinguifolia De Not., Epat. Borneo: 13, 1874 (De Notaris 1874).

** Nothostrepta R.M.Schust., Phytologia 45 (5): 420, 1980 (Schuster 1980b).

*** Nothostreptabifida (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (Schuster 1980b). Bas.: Plagiochilabifida Steph., Annuario Reale Ist. Bot. Roma 2: 86, 1885 [1886] (Stephani 1885d).

*** Nothostreptalongissima (Steph.) R.M.Schust., Phytologia 45 (5): 420, 1980 (Schuster 1980b). Bas.: Anastrophyllumlongissimum Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 13, 1901 (Stephani 1901b).

** Pisanoa Hässel, Lindbergia 14 (3): 179, 1988 [1989] (Hässel 1988).

*** Pisanoachilensis Hässel, Lindbergia 14 (3): 179, 1988 [1989] (Hässel 1988).

** Protosyzygiella (Inoue) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (Schuster 1980c). Bas.: Syzygiellasubg.Protosyzygiella Inoue, J. Hattori Bot. Lab. 29: 180, 1966 (Inoue 1966c).

*** Protosyzygiellapseudoconnexa (Herzog) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 334, 1980 (Schuster 1980c). Bas.: Plagiochilapseudoconnexa Herzog, Rev. Bryol. Lichénol. 21 (3/4): 259, 1952 [1953] (Herzog 1952d).

*** Syzygiella Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). 24

*** subg.Anomalae (Inoue) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Syzygiellasect.Anomalae Inoue, J. Hattori Bot. Lab. 29: 183, 1966 (Inoue 1966c).

*** Syzygiellaanomala (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 471 (190), 1902 (Stephani 1902f). Bas.: Plagiochilaanomala Lindenb. et Gottsche, Syn. Hepat. 5: 646, 1847 (Gottsche et al. 1847).

*** Syzygiellabilobata Inoue, J. Hattori Bot. Lab. 29: 186, 1966 (Inoue 1966c).

** Syzygiellaciliata Gradst. et A.R.Benitez, Nova Hedwigia 99 (1/2): 115, 2014 (Gradstein and Benitez 2014).

*** Syzygiellaconcreta (Gottsche) Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). Bas.: Jungermanniaconcreta Gottsche, Mexik. Leverm.: 82, 1863 (Gottsche 1863).

*** Syzygiellamanca (Mont.) Steph., Hedwigia 31 (1): 14, 1892 (Jack and Stephani 1892). Bas.: Chiloscyphusmancus Mont., Syll. Gen. Sp. Crypt.: 63, 1856 (Montagne 1856b).

*** Syzygiellapectiniformis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 501, 1885 (Spruce 1885).

*** Syzygiellatonduzana Steph., Sp. Hepat. (Stephani) 6: 118, 1917 (Stephani 1917a).

*** Syzygiellatrigonifolia (Steph.) Herzog, Hedwigia 74 (2): 87, 1934 (Herzog 1934a). Bas.: Jamesoniellatrigonifolia Steph., Biblioth. Bot. 87 (2): 185, 1916 (Stephani 1916a).

*** subg.Cryptochila (R.M.Schust.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Cryptochila R.M.Schust., J. Hattori Bot. Lab. 26: 284, 1963 (Schuster 1963b).

*** Syzygiellaacinacifolia (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniaacinacifolia Hook.f. et Taylor, London J. Bot. 3: 367, 1844 (Hooker and Taylor 1844a).

*** Syzygiellanigrescens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniellanigrescens Steph., Hedwigia 34 (2): 48, 1895 (Stephani 1895c).

*** Syzygiellapaludosa (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniellapaludosa Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 11, 1901 (Stephani 1901b).

*** Syzygiellapseudocclusa (E.A.Hodgs.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniellapseudocclusa E.A.Hodgs., Trans. Roy. Soc. New Zealand 85 (4): 583, 1958 (Hodgson 1958).

*** Syzygiellasonderi (Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 143, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniasonderi Gottsche, Linnaea 28 (5): 550, 1856 [1857] (Gottsche 1856).

*** Syzygiellaspegazziniana (Spruce ex C.Massal.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniaspegazziniana Spruce ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 216, 1885 (Massalongo 1885).

*** subg.Pseudoplagiochila Inoue, J. Hattori Bot. Lab. 29: 182, 1966 (Inoue 1966c).

*** Syzygiellaovalifolia Inoue, J. Hattori Bot. Lab. 29: 191, 1966 (Inoue 1966c).

* Syzygiellasecurifolia (Nees) Inoue, J. Hattori Bot. Lab. 46: 232, 1979 (Inoue 1979a). Bas.: Plagiochilasecurifolia Nees, Sp. Hepat. (Lindenberg) 2-4: 58, 1840 (Lindenberg 1840). 25

*** Syzygiellasubintegerrima (Reinw., Blume et Nees) Spruce, J. Linn. Soc., Bot. 30 (210): 362, 1895 (Gepp 1895b). Bas.: Jungermanniasubintegerrima Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 238, 1824 [1825] (Reinwardt et al. 1824a).

*** Syzygiellatasmanica (Hook.f. et Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniatasmanica Hook.f. et Taylor, London J. Bot. 5: 274, 1846 (Taylor 1846a).

*** subg.Roivainenia (Perss.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Roivainenia Perss., Nova Hedwigia 3 (1): 43, 1961 (Persson and Grolle 1961).

*** Syzygiellajacquinotii (Mont.) Hentschel, K.Feldberg, Váňa et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniajacquinotii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 250, 1843 (Montagne 1843).

*** subg.Syzygiella

*** Syzygiellaautumnalis (DC.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniaautumnalis DC., Fl. Franç. (DC. & Lamarck), 5 (6): 202, 1815 (De Candolle and Lamarck 1815).

*** Syzygiellacampanulata Herzog, Rev. Bryol. Lichénol. 11 (1): 9, 1938 [1939] (Herzog 1938a).

*** Syzygiellacolorata (Lehm.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniacolorata Lehm., Linnaea 4: 366, 1829 (Lehmann 1829).

** Syzygiellacoloratavar.collenchymata J.J.Engel et Váňa, Mem. New York Bot. Gard. 105: 102, 2013 (Váňa and Engel 2013).

*** Syzygiellacontigua Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 470 (189), 1902 (Stephani 1902f). Nom. nov. pro Jungermannia contigua Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 118, 1864 (Gottsche 1864), nom. illeg.

*** Syzygiellaelongella (Taylor) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 144, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniaelongella Taylor, London J. Bot. 5: 274, 1846 (Taylor 1846a).

*** Syzygiellamacrocalyx (Mont.) Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). Bas.: Jungermanniamacrocalyx Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 248, 1843 (Montagne 1843).

*** Syzygiellanipponica (S.Hatt.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniellanipponica S.Hatt., J. Jap. Bot. 19 (11): 350, 1943 (Hattori 1943b).

** Syzygiellaoenops (Lindenb. et Gottsche) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniaoenops Lindenb. et Gottsche, Syn. Hepat. 5: 673, 1847 (Gottsche et al. 1847).

*** Syzygiellaperfoliata (Sw.) Spruce, J. Bot. 14: 234, 1876 (Spruce 1876a). Bas.: Jungermanniaperfoliata Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788).

*** Syzygiellapurpurascens (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jamesoniellapurpurascens Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 200, 1891 [1892] (Stephani 1891b).

*** Syzygiellarubricaulis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 468 (187), 1902 (Stephani 1902f). Bas.: Jungermanniarubricaulis Nees, Fl. Bras. (Martius) 1 (1): 344, 1833 (Nees 1833a).

*** Syzygiellasetulosa Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 469 (188), 1902 (Stephani 1902f).

*** Syzygiellateres (Carrington et Pearson) Váňa, Phytotaxa 76 (3): 35, 2013 (Váňa et al. 2013h). Bas.: Jungermanniateres Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 9, 1888 (Carrington and Pearson 1888b).

*** Syzygiellaundata (Mont.) K.Feldberg, Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 145, 2010 (Feldberg et al. 2010b). Bas.: Jungermanniaundata Mont., Ann. Sci. Nat. Bot. (sér. 4) 14: 183, 1860 (Montagne 1860).

Incertae sedis

*** Syzygiellaeatonii (Austin) Inoue, J. Jap. Bot. 37 (12): 359, 1962 (Inoue 1962a). Bas.: Plagiochilaeatonii Austin, Trans. Connecticut Acad. Arts 8 (15): 257, 1891 (Evans 1891).

*** Syzygiellauleana Steph., Hedwigia 44 (4): 224, 1905 (Stephani 1905a).

** Vanaea (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (Gradstein and Florschütz-de Waard 1989). Bas.: Anastrophyllumsubg.Vanaea Inoue et Gradst., Bull. Natl. Sci. Mus. Tokyo, B 14 (3): 88, 1988 (Inoue and Gradstein 1988).

*** Vanaeaplagiochiloides (Inoue et Gradst.) Inoue et Gradst., Trop. Bryol. 1: 33, 1989 (Gradstein and Florschütz-de Waard 1989). Bas.: Anastrophyllumplagiochiloides Inoue et Gradst., Bull. Natl. Sci. Mus. Tokyo, B 14 (3): 88, 1988 (Inoue and Gradstein 1988).

*** Anastrophyllaceae L.Söderstr., De Roo et Hedd

by J. Váňa and L. Söderström

Anastrophyllaceae was described by Söderström et. al (2010b) from elements usually included in Lophoziaceae. Further taxonomic and nomenclatural notes can be found in Váňa et al. (2013k, 2013a). The complex of Chandonanthus/Plicanthus/Tetralophozia should be checked using molecular methods before generic and specific status of most of the species can be confirmed. The placement of Hattoria and Zantenia in the family is still provisional. The placement of Isopaches is also unclear, de Roo et al. (2007) placed it in the family, but the study of Vilnet et al. (2010) showed that it can not be placed there.

*** Anastrepta (Lindb.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (Schiffner 1893b). Bas.: Jungermanniasect.Anastrepta Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 40, 1889 (Lindberg and Arnell 1889).

*** Anastreptaorcadensis (Hook.) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (Schiffner 1893b). Bas.: Jungermanniaorcadensis Hook., Brit. Jungermann.: tab. 71, 1815 (Hooker 1815).

*** Anastrophyllum (Spruce) Steph., Hedwigia 32 (3): 139, 1893 (Stephani 1893b). Bas.: Jungermanniasubg.Anastrophyllum Spruce, J. Bot. 14: 235, 1876 (Spruce 1876a).

*** Anastrophyllumalpinum Steph., Sp. Hepat. (Stephani) 6: 103, 1917 (Stephani 1917a).

*** Anastrophyllumassimile (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermanniaassimilis Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 93, 1860 [1861] (Mitten 1860c).

*** Anastrophyllumauritum (Lehm.) Steph., Bull. Herb. Boissier (sér. 2) 1 (11): 1137 (120), 1901 (Stephani 1901c). Bas.: Jungermanniaaurita Lehm., Linnaea 4: 368, 1829 (Lehmann 1829).

*** Anastrophyllumciliatum Steph., Hedwigia 32 (3): 139, 1893 (Stephani 1893b).

*** Anastrophyllumdonnianum (Hook.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermanniadonniana Hook., Brit. Jungermann.: tab. 39, 1813 (Hooker 1813).

*** Anastrophyllumellipticum Inoue, Bull. Natl. Sci. Mus. Tokyo, B 4 (1): 13, 1978 (Inoue 1978b). 26

** Anastrophyllumesenbeckii (Mont.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermanniaesenbeckii Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843 (Montagne 1843).

*** Anastrophyllumfissum Steph., Bull. Herb. Boissier 5 (10): 845, 1897 (Stephani 1897c).

*** Anastrophyllumjoergensenii Schiffn., Hedwigia 49 (4): 396, 1910 (Schiffner 1910b).

* Anastrophyllumlignicola D.B.Schill et D.G.Long, Ann. Bot. Fenn. 39 (2): 130, 2002 (Schill and Long 2002). 27

*** Anastrophyllummichauxii (F.Weber) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 289, 1932 [1933] (Buch 1932). Bas.: Jungermanniamichauxii F.Weber, Hist. Musc. Hepat. Prodr.: 76, 1815 (Weber 1815).

*** Anastrophyllumnigrescens (Mitt.) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermannianigrescens Mitt., Hooker’s J. Bot. Kew Gard. Misc. 3: 358, 1851 (Mitten 1851).

*** Anastrophyllumobtusum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 285, 1950 (Herzog 1950a).

*** Anastrophyllumpiligerum (Nees) Steph., Hedwigia 32 (3): 140, 1893 (Stephani 1893b). Bas.: Jungermanniapiligera Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 414, 1824 [1825] (Reinwardt et al. 1824b).

*** Anastrophyllumsquarrosum Herzog, Ann. Bryol. 5: 72, 1932 (Herzog 1932b).

*** Anastrophyllumstellatum R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a).

*** Anastrophyllumtubulosum (Nees) Grolle, J. Hattori Bot. Lab. 28: 101, 1965 (Grolle 1965e). Bas.: Jungermanniatubulosa Nees, Enum. Pl. Crypt. Javae: 32, 1830 (Nees 1830).

*** Barbilophozia Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908).

** subg.Barbilophozia

*** Barbilophoziabarbata (Schmidel ex Schreb.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908). Bas.: Jungermanniabarbata Schmidel ex Schreb., Spic. Fl. Lips.: 107, 1771 (Schreber 1771).

*** Barbilophoziahatcheri (A.Evans) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908). Bas.: Jungermanniahatcheri A.Evans, Bull. Torrey Bot. Club 25 (8): 417, 1898 (Evans 1898).

*** Barbilophozialycopodioides (Wallr.) Loeske, Verh. Bot. Vereins Prov. Brandenburg 49 (1): 37, 1908 (Loeske 1908). Bas.: Jungermannialycopodioides Wallr., Comp. fl. Germ. 2 (III): 76, 1831 (Bluff and Fingerhuth 1831).

*** Barbilophoziarubescens (R.M.Schust. et Damsh.) Kartt. et L.Söderstr., Ann. Bot. Fenn. 29 (2): 120, 1992 (Söderström et al. 1992). Bas.: Lophoziarubescens R.M.Schust. et Damsh., Phytologia 63 (5): 325, 1987 (Schuster and Damsholt 1987).

** subg.Sudeticae (Schljakov) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Lophoziasect.Sudeticae Schljakov, Pečen. Mchi Sev. SSSR 3: 113, 1980 (Shliakov 1980a).

*** Barbilophoziasudetica (Nees ex Huebener) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Jungermanniasudetica Nees ex Huebener, Hepaticol. germ.: 142, 1834 (Hübener 1834). 28

*** Biantheridion (Grolle) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jamesoniellasect.Biantheridion Grolle, Trans. Brit. Bryol. Soc. 4 (4): 662, 1964 (Grolle 1964i).

*** Biantheridionundulifolium (Nees) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermanniaschraderi β undulifolia Nees, Naturgesch. Eur. Leberm. 1: 306, 1833 (Nees 1833c).

*** Chandonanthus Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (Hooker 1867).

*** Chandonanthussquarrosus (Menzies) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (Hooker 1867). Bas.: Jungermanniasquarrosa Menzies, Musci Exot. 1: tab. 78, 1818 (Hooker 1818).

*** Crossocalyx Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (Meylan 1939).

*** Crossocalyxhellerianus (Nees ex Lindenb.) Meyl., Bull. Soc. Vaud. Sci. Nat. 60 (249): 266, 1939 (Meylan 1939). Bas.: Jungermanniahelleriana Nees ex Lindenb., Syn. hepat. eur: 64, 1829 (Lindenberg 1829).

*** Crossocalyxtenuis (Harry Williams) Schljakov, Novosti Sist. Nizš. Rast. 15: 246, 1978 (Shliakov 1978). Bas.: Anastrophyllumtenue Harry Williams, Bryologist 71 (1): 34, 1968 (Williams 1968).

*** Gymnocolea (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermanniasect.Gymnocolea Dumort., Syll. Jungerm. Europ.: 52, 1831 (Dumortier 1831).

*** Gymnocoleaborealis (Frisvoll et Moen) R.M.Schust., Lindbergia 12 (1): 7, 1986 (Schuster 1986). Bas.: Lophoziaborealis Frisvoll et Moen, Lindbergia 6 (2): 138, 1980 [1981] (Frisvoll and Moen 1980).

** Gymnocoleafascinifera Potemkin, Arctoa 2: 76, 1993 (Potemkin 1993).

*** Gymnocoleainflata (Huds.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermanniainflata Huds., Fl. Angl. (Hudson), ed. 2: 511, 1778 (Hudson 1778).

* Gymnocoleainflatasubsp.acutiloba (Schiffn.) R.M.Schust. et Damsh. ex L.Söderstr. et Váňa, Lindbergia 27 (1): 43, 2002 (Söderström et al. 2002). Bas.: Lophoziaacutiloba Schiffn., Hedwigia 48 (3): 187, 1909 (Schiffner 1909a). 29

*** Hamatostrepta Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (Váňa and Long 2008).

*** Hamatostreptaconcinna Váňa et D.G.Long, Fieldiana, Bot. (n.ser.) 47: 134, 2008 (Váňa and Long 2008).

** Hattoria R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 69, 1961 (Schuster 1961a).

*** Hattoriayakushimensis (Horik.) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 70, 1961 (Schuster 1961a). Bas.: Anastrophyllumyakushimense Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 149, 1934 (Horikawa 1934).

*** Isopaches H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 287, 1932 [1933] (Buch 1932).

*** Isopachesalboviridis (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (Shliakov 1979). Bas.: Lophoziaalboviridis R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 487, 1969 (Schuster 1969b).

*** Isopachesbicrenatus (Schmidel ex Hoffm.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (Buch 1932). Bas.: Jungermanniabicrenata Schmidel ex Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 11 (addenda), 1795 [1796] (Hoffmann 1795).

*** Isopachesdecolorans (Limpr.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 288, 1932 [1933] (Buch 1932). Bas.: Jungermanniadecolorans Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 316, 1879 [1880] (Limpricht 1879).

*** Isopachespumicicola (Berggr.) Bakalin, Arctoa 17: 162, 2008 [2009] (Bakalin 2008b). Bas.: Lophoziapumicicola Berggr., New Zealand Hepat.: 21, 1898 (Berggren 1898).

*** Neoorthocaulis L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (Söderström et al. 2010b).

*** Neoorthocaulisattenuatus (Mart.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (Söderström et al. 2010b). Bas.: Jungermanniaquinquedentata δ attenuata Mart., Fl. crypt. erlang.: 177, 1817 (Martius 1817).

*** Neoorthocaulisbinsteadii (Kaal.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 49, 2010 (Söderström et al. 2010b). Bas.: Jungermanniabinsteadii Kaal., Skr. Vidensk.-Selsk. Christiana, Math.-Naturvidensk. Kl. 1898 (9): 9, 1898 (Kaalaas 1898).

*** Neoorthocaulisfloerkei (F.Weber et D.Mohr) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Jungermanniafloerkei F.Weber et D.Mohr, Bot. Taschenb. (Weber): 410, 1807 (Weber and Mohr 1807).

** Neoorthocaulishyperboreus (R.M.Schust.) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b). Bas.: Lophoziafloerkeivar.hyperborea R.M.Schust., Bull. Natl. Mus. Canada 164: 21, 1959 (Schuster et al. 1959).

*** Orthocaulis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 293, 1932 [1933] (Buch 1932).

*** Orthocaulisatlanticus (Kaal.) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 294, 1932 [1933] (Buch 1932). Bas.: Jungermanniaatlantica Kaal., Skr. Vidensk.-Selsk. Christiana, Math.-Naturvidensk. Kl. 1898 (9): 11, 1898 (Kaalaas 1898).

* Orthocauliscavifolius H.Buch et S.W.Arnell, Memoranda Soc. Fauna Fl. Fennica 26: 71, 1951 (Buch 1951).

** Plicanthus R.M.Schust., Nova Hedwigia 74 (3/4): 484, 2002 (Schuster 2002a).

*** Plicanthusbirmensis (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 223, 2002 (Schuster 2002b). Bas.: Chandonanthusbirmensis Steph., Bull. Soc. Roy. Bot. Belgique 38 (1): 43, 1899 (Stephani 1899h). 30

** Plicanthusdifficilis (Steph.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 30, 2013 (Váňa et al. 2013a). Bas.: Chandonanthusdifficilis Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 101, 1914 (Stephani and Watts 1914).

** Plicanthusgiganteus (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 485, 2002 (Schuster 2002a). Bas.: Chandonanthusgiganteus Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 124, 1911 (Stephani 1911a). 31

*** Plicanthushirtellus (F.Weber) R.M.Schust., Nova Hedwigia 74 (3/4): 492, 2002 (Schuster 2002a). Bas.: Jungermanniahirtella F.Weber, Hist. Musc. Hepat. Prodr.: 50, 1815 (Weber 1815).

*** Schizophyllopsis Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Nom. nov. pro Anastrophyllumsubg.Schizophyllum R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 739, 1969 (Schuster 1969b). 32

** Schizophyllopsisaristata (Herzog ex N.Kitag.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllumbidensvar.aristatum Herzog ex N.Kitag., J. Hattori Bot. Lab. 33: 216, 1970 (Kitagawa 1970).

*** Schizophyllopsisbidens (Reinw., Blume et Nees) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Jungermanniabidens Reinw., Blume et Nees, Nova Acta Phys.-Med. Acad. Caes. Leop.-Carol. Nat. Cur. 12 (1): 208, 1824 [1825] (Reinwardt et al. 1824a).

** Schizophyllopsislanciloba (Steph.) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllumlancilobum Steph., Sp. Hepat. (Stephani) 6: 107, 1917 (Stephani 1917a).

*** Schizophyllopsispapillosa (J.J.Engel et Braggins) Váňa et L.Söderstr., Phytotaxa 152 (1): 48, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllumpapillosum J.J.Engel et Braggins, J. Bryol. 20 (2): 381, 1998 (Engel and Braggins 1998).

*** Schizophyllopsissphenoloboides (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 152 (1): 49, 2013 (Váňa et al. 2013f). Bas.: Anastrophyllumsphenoloboides R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 741, 1969 (Schuster 1969b).

*** Schljakovia Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009).

*** Schljakoviakunzeana (Huebener) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermanniakunzeana Huebener, Hepaticol. germ.: 115, 1834 (Hübener 1834).

*** Schljakovianthus Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009).

*** Schljakovianthusquadrilobus (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermanniaquadriloba Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 55, 1889 (Lindberg and Arnell 1889).

*** Sphenolobopsis R.M.Schust. et N.Kitag., Nova Hedwigia 22: 152, 1971 [1972] (Schuster 1971b).

*** Sphenolobopsispearsonii (Spruce) R.M.Schust., Nova Hedwigia 22: 153, 1971 [1972] (Schuster 1971b). Bas.: Jungermanniapearsonii Spruce, J. Bot. 19: 33, 1881 (Spruce 1881b).

*** Sphenolobus (Lindb.) Berggr., New Zealand Hepat.: 22, 1898 (Berggren 1898). Bas.: Jungermanniasect.Sphenolobus Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 369, 1874 (Lindberg 1874a).

*** Sphenolobusaustroamericanus (Váňa) Váňa, Phytotaxa 81 (1): 30, 2013 (Váňa et al. 2013a). Bas.: Anastrophyllumaustroamericanum Váňa, J. Hattori Bot. Lab. 48: 225, 1980 (Váňa 1980).

*** Sphenolobusminutus (Schreb. ex D.Crantz) Berggr., New Zealand Hepat.: 22, 1898 (Berggren 1898). Bas.: Jungermanniaminuta Schreb. ex D.Crantz, Forts. Hist. Grönland: 285, 1770 (Crantz 1770; non vidi).

*** Sphenolobussaxicola (Schrad.) Steph., Bull. Herb. Boissier (sér. 2) 2 (2): 168 (160), 1902 (Stephani 1902d). Bas.: Jungermanniasaxicola Schrad., Syst. Samml. Crypt. Gew. 2: 4, 1797 (Schrader 1797).

** Tetralophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 13: 227, 1976 (Shliakov 1976). Bas.: Chandonanthussubg.Tetralophozia R.M.Schust., J. Hattori Bot. Lab. 23: 206, 1960 [1961] (Schuster 1960a).

*** Tetralophoziacavallii (Gola) Váňa, Trop. Bryol. 8: 102, 1993 (Váňa 1993). Bas.: Blepharostomumcavallii Gola, Ann. Bot. (Rome) 6 (2): 274, 1907 (Gola 1907).

*** Tetralophoziafiliformis (Steph.) Urmi, J. Bryol. 12 (3): 394, 1983 (Urmi 1983). Bas.: Chandonanthusfiliformis Steph., Sp. Hepat. (Stephani) 3: 644, 1909 (Stephani 1909a).

** Tetralophoziapilifera (Steph.) R.M.Schust., Nova Hedwigia 74 (3/4): 482, 2002 (Schuster 2002a). Bas.: Chandonanthuspilifer Steph., Sp. Hepat. (Stephani) 3: 644, 1909 (Stephani 1909a).

*** Tetralophoziasetiformis (Ehrh.) Schljakov, Novosti Sist. Nizš. Rast. 13: 228, 1976 (Shliakov 1976). Bas.: Jungermanniasetiformis Ehrh., Hannover. Mag. 22 (8): 142, 1784 (Ehrhart 1784).

*** Zantenia (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllumsubg.Zantenia S.Hatt., Bot. Mag. (Tokyo) 79 (937): 342, 1966 (Hattori 1966a).

*** Zanteniaborneensis (Herzog) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 29, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllumborneense Herzog, Trans. Brit. Bryol. Soc. 1 (4): 282, 1950 (Herzog 1950a).

*** Zanteniadenticulata (Grolle) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllumdenticulatum Grolle, Nova Hedwigia 16: 148, 1968 (Grolle 1968d).

*** Zanteniakarstenii (Schiffn.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllumkarstenii Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 268, 1893 (Schiffner 1893a).

*** Zanteniaprionophylla (S.Hatt.) Váňa et J.J.Engel, Mem. New York Bot. Gard. 105: 30, 2013 (Váňa and Engel 2013). Bas.: Anastrophyllumprionophyllum S.Hatt., Bot. Mag. (Tokyo) 79 (937): 342, 1966 (Hattori 1966a).

*** Cephaloziaceae Mig

by J. Váňa with contributions by S.R. Gradstein (Odontoschisma)

Cephaloziaceae was recently studied by Vilnet et al. (2012) and Feldberg et al. (2013). Some taxonomic and nomenclatural notes were published by Váňa et al. (2013g, 2013i) and Gradstein et al. (2014b).

** Alobielloideae R.M.Schust

** Alobiella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (Schiffner 1893b). Bas.: Cephaloziasubg.Alobiella Spruce, Cephalozia: 28, 1882 (Spruce 1882).

*** Alobiellahusnotii (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (Schiffner 1893b). Bas.: Cephaloziahusnotii Spruce, Cephalozia: 30, 1882 (Spruce 1882).

** Alobiellopsis R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b).

*** Alobiellopsisacroscypha (Spruce) R.M.Schust., Nova Hedwigia 10 (1/2): 25, 1965 (Schuster 1965b). Bas.: Cephaloziaacroscypha Spruce, Cephalozia: 30, 1882 (Spruce 1882).

*** Alobiellopsisdominicensis (Spruce) Fulford, Mem. New York Bot. Gard. 11 (3): 350, 1968 (Fulford 1968). Bas.: Alobielladominicensis Spruce, J. Linn. Soc., Bot. 30 (210): 355, 1895 (Gepp 1895b).

*** Alobiellopsisheteromorpha (Lehm.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 682, 1969 (Schuster 1969a). Bas.: Jungermanniaheteromorpha Lehm., Linnaea 4: 362, 1829 (Lehmann 1829).

*** Alobiellopsisparvifolia (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 679, 1969 (Schuster 1969a). Bas.: Alobiellaparvifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 568 (352), 1908 (Stephani 1908e).

*** Alobiellopsispillansii (Sim) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 683, 1969 (Schuster 1969a). Bas.: Cephaloziapillansii Sim, Trans. Roy. Soc. South Africa 15 (1): 87, 1926 (Sim 1926).

** Cephalozioideae Müll.Frib

*** Cephalozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermanniasect.Cephalozia Dumort., Syll. Jungerm. Europ.: 60, 1831 (Dumortier 1831).

*** Cephaloziaacuminata (Herzog) Váňa, Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Hygrobiellaacuminata Herzog, Trans. Brit. Bryol. Soc. 1 (4): 292, 1950 (Herzog 1950a).

** Cephaloziaacutiloba (Inoue) Váňa, Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Metahygrobiellaacutiloba Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 10 (2): 155, 1967 (Inoue 1967c).

*** Cephaloziaalbula Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Nom. nov. pro Jungermannia albula Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 93, 1860 [1861] (Mitten 1860c), nom. illeg.

*** Cephaloziaambigua C.Massal., Malpighia 21 (7/8): 310, 1907 (Massalongo 1907).

*** Cephaloziaaustrigena R.M.Schust. ex J.J.Engel, Novon 17 (3): 312, 2007 (Engel 2007). Based on: Cephaloziabicuspidatasubsp.austrigena R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 712, 1974 (Schuster 1974), nom. inval.

*** Cephaloziabadia (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 483 (313), 1908 (Stephani 1908f). Bas.: Jungermanniabadia Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890 (Gottsche 1890).

*** Cephaloziabicuspidata (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermanniabicuspidata L., Sp. Pl. 1: 1132, 1753 (Linnaeus 1753).

* Cephaloziabicuspidatasubsp.lammersiana (Huebener) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 730, 1974 (Schuster 1974). Bas.: Jungermannialammersiana Huebener, Flora 15 (20): 306, 1832 (Hübener 1832).

*** Cephaloziachilensis (J.J.Engel et R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 29, 2002 (Schuster 2002b). Bas.: Metahygrobiellachilensis J.J.Engel et R.M.Schust., Brittonia 40 (2): 203, 1988 (Engel and Schuster 1988).

*** Cephaloziaconchata (Grolle et Váňa) Váňa, Syst. Bot. 40 (1): 38, 2015 (Shaw et al. 2015). Bas.: Jungermanniaconchata Grolle et Váňa, Fragm. Florist. Geobot. 37 (1): 3, 1992 (Grolle and Váňa 1992).

*** Cephaloziacrossii Spruce, Cephalozia: 46, 1882 (Spruce 1882).

*** Cephaloziadarjeelingensis Udar et D.Kumar, Geophytology 6 (1): 36, 1976 (Udar and Kumar 1976).

*** Cephaloziadrucei (R.M.Schust.) Váňa, Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Metahygrobielladrucei R.M.Schust., J. Hattori Bot. Lab. 26: 273, 1963 (Schuster 1963b).

*** Cephaloziafuegiensis Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Nom. nov. pro Hygrobiella dusenii Steph., Sp. Hepat. (Stephani) 6: 444, 1924 (Stephani 1924).

*** Cephaloziahamatiloba Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 427 (303), 1908 (Stephani 1908g).

*** Cephaloziahamatilobasubsp.siamensis (N.Kitag.) Váňa, Acta Bot. Fenn. 177: 16, 2004 (Koponen et al. 2004). Bas.: Cephaloziasiamensis N.Kitag., J. Hattori Bot. Lab. 32: 293, 1969 (Kitagawa 1969c).

*** Cephalozialacinulata (J.B.Jack ex Gottsche et Rabenh.) Spruce, Cephalozia: 45, 1882 (Spruce 1882). Bas.: Jungermannialacinulata J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 62-64: no. 624, 1877 (Gottsche and Rabenhorst 1877).

*** Cephalozialucens (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908 (Stephani 1908f). Bas.: Jungermannialucens A.Evans, Trans. Connecticut Acad. Arts 8 (15): 258, 1891 (Evans 1891).

*** Cephaloziamacgregorii (Steph.) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Hygrobiellamacgregorii Steph., Hedwigia 34 (2): 45, 1895 (Stephani 1895c).

*** Cephaloziamacounii (Austin) Austin, Hepat. bor.-amer.: 14, 1873 (Austin 1873). Bas.: Jungermanniamacounii Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 222, 1869 (Austin 1869).

*** Cephaloziamaxima Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (Stephani 1924).

*** Cephaloziamollusca (De Not.) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Jungermanniamollusca De Not., Epat. Borneo: 16, 1874 (De Notaris 1874).

* Cephalozianeesiana Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 429 (305), 1908 (Stephani 1908g). 33

*** Cephalozianishimurae (N.Kitag.) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Hygrobiellanishimurae N.Kitag., Misc. Bryol. Lichenol. 9 (4): 69, 1982 (Kitagawa 1982).

** Cephaloziapachygyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007).

** Cephaloziaphysocaula (Hook.f. et Taylor) Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 485 (315), 1908 (Stephani 1908f). Bas.: Jungermanniaphysocaula Hook.f. et Taylor, London J. Bot. 3: 455, 1844 (Hooker and Taylor 1844b).

** Cephaloziaschusteriana J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007).

*** Cephaloziastolonacea (Herzog) Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Hygrobiellastolonacea Herzog, Trans. Brit. Bryol. Soc. 1 (4): 293, 1950 (Herzog 1950a).

*** Cephaloziatubulata (Hook.f. et Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877). Bas.: Jungermanniatubulata Hook.f. et Taylor, London J. Bot. 3: 463, 1844 (Hooker and Taylor 1844b).

Excluded from the genus

* Cephaloziahians Steph., Sp. Hepat. (Stephani) 6: 441, 1924 (Stephani 1924). 34

* Cephaloziaindica Udar et D.Kumar, Geophytology 6 (1): 42, 1976 (Udar and Kumar 1976). 35

* Cephaloziakodaikanalensis G.Asthana et Saumya Srivast., Geophytology 43 (1): 63, 2013 (Asthana and Srivastava 2013). 36

* Cephaloziaparvifolia Arnell, Rev. Bryol. 25 (1): 1, 1898 (Arnell 1898). 37

* Cephaloziatricuspidata (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877). Bas.: Jungermanniatricuspidata Nees, Enum. Pl. Crypt. Javae: 31, 1830 (Nees 1830). 38

*** Fuscocephaloziopsis Fulford, Mem. New York Bot. Gard. 11 (3): 353, 1968 (Fulford 1968).

** Fuscocephaloziopsisaffinis (Lindb. ex Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephaloziaaffinis Lindb. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (4): 277 (291), 1908 (Stephani 1908j).

*** Fuscocephaloziopsisafricana (Váňa) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephaloziaafricana Váňa, Beih. Nova Hedwigia 90: 187, 1988 (Váňa 1988).

*** Fuscocephaloziopsisalbescens (Hook.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermanniaalbescens Hook., Brit. Jungermann.: tab. 72, 1815 (Hooker 1815).

* Fuscocephaloziopsisalbescensvar.islandica (Nees) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermanniaislandica Nees, Naturgesch. Eur. Leberm. 2: 29, 1836 (Nees 1836).

*** Fuscocephaloziopsisbaldwinii (C.M.Cooke) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephaloziabaldwinii C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 35, 1904 (Cooke 1904).

** Fuscocephaloziopsisbiloba (Herzog) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (Fulford 1968). Bas.: Alobiellabiloba Herzog, Feddes Repert. Spec. Nov. Regni Veg. 57 (1/2): 166, 1955 (Herzog 1955).

*** Fuscocephaloziopsiscatenulata (Huebener) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermanniacatenulata Huebener, Hepaticol. germ.: 169, 1834 (Hübener 1834).

*** Fuscocephaloziopsiscatenulatasubsp.nipponica (S.Hatt.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephalozianipponica S.Hatt., Bull. Tokyo Sci. Mus. 11: 74, 1944 (Hattori 1944d).

*** Fuscocephaloziopsisconnivens (Dicks.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermanniaconnivens Dicks., Fasc. Pl. Crypt. Brit. 4: 19, 1801 (Dickson 1801).

*** Fuscocephaloziopsisconnivenssubsp.fissa (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Cephaloziafissa Steph., Hedwigia 30 (5): 204, 1891 (Stephani 1891a).

*** Fuscocephaloziopsisconnivenssubsp.sandwicensis (Mont.) Váňa et L.Söderstr., Phytotaxa 112 (1): 9, 2013 (Váňa et al. 2013g). Bas.: Jungermanniasandwicensis Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 249, 1843 (Montagne 1843).

*** Fuscocephaloziopsiscrassifolia (Lindenb. et Gottsche) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Jungermanniacrassifolia Lindenb. et Gottsche, Syn. Hepat. 5: 685, 1847 (Gottsche et al. 1847).

*** Fuscocephaloziopsisgollanii (Steph.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephaloziagollanii Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 428 (304), 1908 (Stephani 1908g).

*** Fuscocephaloziopsisleucantha (Spruce) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozialeucantha Spruce, Cephalozia: 68, 1882 (Spruce 1882).

*** Fuscocephaloziopsisloitlesbergeri (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephalozialoitlesbergeri Schiffn., Österr. Bot. Z. 62 (1): 10, 1912 (Schiffner 1912b).

*** Fuscocephaloziopsislunulifolia (Dumort.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Jungermannialunulifolia Dumort., Syll. Jungerm. Europ.: 61, 1831 (Dumortier 1831). 39

*** Fuscocephaloziopsismacrostachya (Kaal.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephaloziamacrostachya Kaal., Rev. Bryol. 29 (1): 8, 1902 (Kaalaas 1902).

*** Fuscocephaloziopsismacrostachyasubsp.australis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephaloziamacrostachyasubsp.australis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 754, 1974 (Schuster 1974).

*** Fuscocephaloziopsismacrostachyasubsp.macrostachyavar.spiniflora (Schiffn.) Váňa et L.Söderstr., Phytotaxa 112 (1): 10, 2013 (Váňa et al. 2013g). Bas.: Cephaloziaspiniflora Schiffn., Hedwigia 54 (6): 323, 1914 (Schiffner 1914a).

*** Fuscocephaloziopsismonticola (J.D.Godfrey) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Schofieldiamonticola J.D.Godfrey, Bryologist 79 (3): 315, 1976 (Godfrey 1976).

*** Fuscocephaloziopsispachycaulis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Cephaloziapachycaulis R.M.Schust., Bryologist 96 (4): 623, 1993 (Schuster 1993b).

*** Fuscocephaloziopsispleniceps (Austin) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Jungermanniapleniceps Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 222, 1869 (Austin 1869).

** Fuscocephaloziopsisplenicepsvar.caroliniana (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Cephaloziaplenicepsvar.caroliniana R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 780, 1974 (Schuster 1974).

*** Fuscocephaloziopsispulvinata (Steph.) Fulford, Mem. New York Bot. Gard. 11 (3): 355, 1968 (Fulford 1968). Bas.: Alobiellapulvinata Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 572 (356), 1908 (Stephani 1908e).

* Fuscocephaloziopsisschusteri (Sushil K.Singh et D.K.Singh) Váňa, Phytotaxa 183 (4): 291, 2014 (Váňa et al. 2014a). Bas.: Cephaloziaschusteri Sushil K.Singh et D.K.Singh, Lindbergia 32 (1): 1, 2007 (Singh and Singh 2007a).

*** Fuscocephaloziopsissubintegra Gradst. et Váňa, Cryptog. Bryol. 25 (3): 274, 2004 (Parolly et al. 2004).

*** Fuscocephaloziopsiszoopsioides (Horik.) Váňa et L.Söderstr., Phytotaxa 112 (1): 11, 2013 (Váňa et al. 2013g). Bas.: Cephaloziazoopsioides Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 178, 1934 (Horikawa 1934).

*** Nowellia Mitt., Nat. hist. Azores: 321, 1870 (Mitten 1870).

** sect.Acronowellia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (Schuster 1974).

*** Nowelliareedii H.Rob., Bryologist 73 (1): 150, 1970 (Robinson 1970).

*** Nowelliayunckeri Fulford, Mem. New York Bot. Gard. 11 (3): 329, 1968 (Fulford 1968).

** sect.Metanowellia (Grolle) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 816, 1974 (Schuster 1974). Bas.: Nowelliasubg.Metanowellia Grolle, J. Hattori Bot. Lab. 31: 33, 1968 (Grolle 1968b).

*** Nowelliaborneensis (De Not.) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (Schiffner 1893b). Bas.: Jungermanniacurvifoliavar.borneensis De Not., Epat. Borneo: 19, 1874 (De Notaris 1874).

*** Nowelliadominicensis Steph., Sp. Hepat. (Stephani) 6: 443, 1924 (Stephani 1924).

*** Nowelliaevansii Grolle, J. Hattori Bot. Lab. 31: 33, 1968 (Grolle 1968b).

*** Nowellialangii Pearson, J. Linn. Soc., Bot. 46 (305): 25, 1922 (Pearson 1922b).

*** Nowelliapusilla Grolle, J. Hattori Bot. Lab. 31: 37, 1968 (Grolle 1968b).

*** Nowelliawrightii (Gottsche ex Spruce) Steph. ex Duss, Enum. musc. Antilles franç., Hép.: 21, 1903 (Duss 1903). Bas.: Cephaloziawrightii Gottsche ex Spruce, J. Linn. Soc., Bot. 30 (210): 354, 1895 (Gepp 1895b).

** sect.Nowellia

*** Nowelliaaciliata (P.C.Chen et P.C.Wu) Mizut., Hikobia 11: 469, 1994 (Mizutani 1994). Bas.: Nowelliacurvifoliavar.aciliata P.C.Chen et P.C.Wu, Obs. fl. Hwangs.: 6, 1965 (Chen and Wu 1965).

*** Nowelliacurvifolia (Dicks.) Mitt., Nat. hist. Azores: 321, 1870 (Mitten 1870). Bas.: Jungermanniacurvifolia Dicks., Fasc. Pl. Crypt. Brit. 2: 15, 1790 (Dickson 1790).

** Odontoschismatoideae H.Buch ex Grolle

*** Odontoschisma (Dumort.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835). Bas.: Pleuroschismasect.Odontoschisma Dumort., Syll. Jungerm. Europ.: 68, 1831 (Dumortier 1831).

** sect.Cladopodiella (H.Buch) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (Aranda et al. 2014). Bas.: Cladopodiella H.Buch, Memoranda Soc. Fauna Fl. Fennica 1: 89, 1927 (Buch 1927).

*** Odontoschismafrancisci (Hook.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Jungermanniafrancisci Hook., Brit. Jungermann.: tab. 49, 1813 (Hooker 1813).

** sect.Denudata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 833, 1974 (Schuster 1974).

*** Odontoschismabrasiliense Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 585 (369), 1908 (Stephani 1908e).

** Odontoschismacleefii Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (Aranda et al. 2014).

*** Odontoschismadenudatum (Mart.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835). Bas.: Jungermanniascalarisvar.denudata Mart., Fl. crypt. erlang.: 183, 1817 (Martius 1817).

** Odontoschismadenudatumsubsp.naviculare (Steph.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (Aranda et al. 2014). Bas.: Jamesoniellanavicularis Steph., Sp. Hepat. (Stephani) 6: 101, 1917 (Stephani 1917a).

** Odontoschismadenudatumsubsp.sandvicense (Ångstr.) Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (Aranda et al. 2014). Bas.: Sphagnoecetissandvicensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 29 (4): 22, 1872 (Ångström 1872).

*** Odontoschismaelongatum (Lindb.) A.Evans, Rhodora 14 (157): 13, 1912 (Evans 1912b). Bas.: Odontoschismadenudatumf.elongatum Lindb., Helsingf. Dagbl. 1874 (45, 16 Feb): 2, 1874 (Lindberg 1874b).

*** Odontoschismaengelii Gradst. et Burghardt, Fieldiana, Bot. (n.ser.) 47: 194, 2008 (Gradstein and Burghardt 2008).

*** Odontoschismalongiflorum (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (Trevisan 1877). Bas.: Sphagnoecetislongiflora Taylor, London J. Bot. 5: 281, 1846 (Taylor 1846a).

*** Odontoschismamacounii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 92, 1884 (Underwood 1884). Bas.: Sphagnoecetismacounii Austin, Bull. Torrey Bot. Club 3 (3): 13, 1872 (Austin 1872).

*** Odontoschismaportoricense (Hampe et Gottsche) Steph., Hedwigia 27 (11/12): 296, 1888 (Stephani 1888c). Bas.: Sphagnoecetisportoricensis Hampe et Gottsche, Linnaea 25 (3): 343, 1852 [1853] (Hampe and Gottsche 1852).

** Odontoschismapseudogrosseverrucosum Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1019, 2014 (Aranda et al. 2014).

*** Odontoschismapurpuratum Herzog, Trans. Brit. Bryol. Soc. 1 (4): 297, 1950 (Herzog 1950a).

*** Odontoschismasoratamum Fulford, Mem. New York Bot. Gard. 11 (3): 338, 1968 (Fulford 1968).

*** Odontoschismavariabile (Lindenb. et Gottsche) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 419, 1877 (Trevisan 1877). Bas.: Sphagnoecetisvariabilis Lindenb. et Gottsche, Syn. Hepat. 5: 688, 1847 (Gottsche et al. 1847).

** Odontoschismazhui Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1020, 2014 (Aranda et al. 2014).

** sect.Iwatsukia (N.Kitag.) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (Gradstein et al. 2014b). Bas.: Iwatsukia N.Kitag., J. Hattori Bot. Lab. 27: 178, 1964 (Kitagawa 1964).

*** Odontoschismabifidum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (Gradstein et al. 2014b). Bas.: Cladomastigumbifidum Fulford, Acta Bot. Venez. 2 (5/8): 80, 1967 (Fulford 1967).

*** Odontoschismajishibae (Steph.) L.Söderstr. et Váňa, Phytotaxa 112 (1): 13, 2013 (Váňa et al. 2013g). Bas.: Cephaloziajishibae Steph., Sp. Hepat. (Stephani) 6: 437, 1924 (Stephani 1924).

*** Odontoschismaspinosum (Fulford) Gradst., S.C.Aranda et Vanderp., Phytotaxa 162 (4): 232, 2014 (Gradstein et al. 2014b). Bas.: Cladomastigumspinosum Fulford, Mem. New York Bot. Gard. 23: 840, 1972 (Fulford 1972).

** sect.Neesia Gradst., S.C.Aranda et Vanderp., Taxon 63 (5): 1017, 2014 (Aranda et al. 2014).

*** Odontoschismafluitans (Nees) L.Söderstr. et Váňa, Phytotaxa 112 (1): 12, 2013 (Váňa et al. 2013g). Bas.: Jungermanniafluitans Nees, Flora 6 (2): 30, 1823 (Link 1823).

** sect.Odontoschisma

*** Odontoschismagrosseverrucosum Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 593 (377), 1908 (Stephani 1908e).

*** Odontoschismasphagni (Dicks.) Dumort., Recueil Observ. Jungerm.: 19, 1835 (Dumortier 1835). Bas.: Jungermanniasphagni Dicks., Fasc. Pl. Crypt. Brit. 1: 6, 1785 (Dickson 1785).

** Odontoschismasteyermarkii Gradst. et Ilk.-Borg., Nova Hedwigia 100 (1/2): 38, 2015 [2014 online] (Gradstein and Ilkiu-Borges 2015).

Incertae sedis

* Odontoschismaobcordatum (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 586 (370), 1908 (Stephani 1908e). Bas.: Cephaloziaobcordata Spruce, Cephalozia: 61, 1882 (Spruce 1882). 40

** Schiffnerioideae R.M.Schust

** Schiffneria Steph., Österr. Bot. Z. 44 (1): 1, 1894 (Stephani 1894c).

*** Schiffneriahyalina Steph., Österr. Bot. Z. 44 (1): 1, 1894 (Stephani 1894c).

** Trabacelluloideae R.M.Schust

** Haesselia Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (Grolle and Gradstein 1988).

*** Haesseliaacuminata Gradst., Trop. Bryol. 1: 30, 1989 (Gradstein and Florschütz-de Waard 1989).

*** Haesseliaroraimensis Grolle et Gradst., J. Hattori Bot. Lab. 64: 327, 1988 (Grolle and Gradstein 1988).

** Trabacellula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (Fulford 1967).

*** Trabacellulatumidula Fulford, Acta Bot. Venez. 2 (5/8): 86, 1967 (Fulford 1967).

*** Cephaloziellaceae Douin

by J. Váňa

The treatment of Cephaloziellaceae follows what was outlined in Váňa et al. (2013e). Some nomenclatural and taxonomic notes can also be found in Söderström et al. (2013a) and Váňa et al. (2013b, 2013l). The placement of Phycolepidozia follows Gradstein et al. (2014a).

*** Allisoniella E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 80, 1965 (Hodgson 1965).

** sect.Allisoniella

*** Allisoniellanigra (Rodway) R.M.Schust., Nova Hedwigia 22: 137, 1971 [1972] (Schuster 1971b). Bas.: Sphenolobusniger Rodway, Tasm. Bryoph.: 33, 1917 (Rodway 1917b).

** Allisoniellanigravar.acutiloba J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007).

** Allisoniellanigrasubsp.novaezelandiae R.M.Schust., Nova Hedwigia 22: 143, 1971 [1972] (Schuster 1971b).

*** Allisoniellarecurva R.M.Schust., Nova Hedwigia 22: 146, 1971 [1972] (Schuster 1971b).

*** Allisoniellasubbipartita (C.Massal.) R.M.Schust. et J.J.Engel, Nova Hedwigia 22: 147, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziasubbipartita C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 235, 1885 (Massalongo 1885).

*** Allisoniellatasmanica R.M.Schust., Nova Hedwigia 22: 145, 1971 [1972] (Schuster 1971b).

** sect.Protomarsupella (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 150, 1971 (Schuster 1971b). Bas.: Protomarsupella R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 264, 1966 (Schuster 1966b).

*** Allisoniellascottii (R.M.Schust.) R.M.Schust., Nova Hedwigia 22: 151, 1971 [1972] (Schuster 1971b). Bas.: Protomarsupellascottii R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 267, 1966 (Schuster 1966b).

*** Amphicephalozia R.M.Schust., Nova Hedwigia 22: 131, 1971 [1972] (Schuster 1971b).

*** Amphicephaloziaafricana Váňa et M.Wigginton, J. Bryol. 30 (1): 55, 2008 (Váňa and Wigginton 2008).

*** Amphicephaloziaamplexicaulis R.M.Schust., Nova Hedwigia 22: 133, 1971 [1972] (Schuster 1971b).

*** Amphicephaloziageisslerae Pócs et Váňa, Polish Bot. J. 46 (2): 145, 2001 (Pócs and Váňa 2001).

*** Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (Váňa et al. 2013k). Bas.: Anastrophyllumsect.Anastrophyllopsis R.M.Schust., Beih. Nova Hedwigia 119: 310, 2002 (Schuster 2002b).

*** Anastrophyllopsisinvolutifolia (Mont. ex Gottsche, Lindenb. et Nees) Váňa et L.Söderstr., Phytotaxa 81 (1): 15, 2013 (Váňa et al. 2013k). Bas.: Jungermanniainvolutifolia Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 1: 81, 1844 (Gottsche et al. 1844).

*** Anastrophyllopsisrevoluta (Steph.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (Váňa et al. 2013k). Bas.: Anastrophyllumrevolutum Steph., Hedwigia 32 (3): 139, 1893 (Stephani 1893b).

*** Anastrophyllopsissubcomplicata (Lehm. et Lindenb.) Váňa et L.Söderstr., Phytotaxa 81 (1): 16, 2013 (Váňa et al. 2013k). Bas.: Jungermanniasubcomplicata Lehm. et Lindenb., Nov. Stirp. Pug. 7: 4, 1838 (Lehmann 1838).

** Cephalojonesia Grolle, Rev. Bryol. Lichénol. 37 (4): 763, 1970 [1971] (Grolle and Vanden Berghen 1970).

*** Cephalojonesiaincuba Grolle et Vanden Berghen, Rev. Bryol. Lichénol. 37 (4): 764, 1970 [1971] (Grolle and Vanden Berghen 1970).

*** Cephalojonesiaincubasubsp.mexicana Burghardt, Gradst. et Váňa, J. Hattori Bot. Lab. 100: 35, 2006 (Burghardt et al. 2006).

** Cephalomitrion R.M.Schust., Nova Hedwigia 61 (3/4): 550, 1995 (Schuster 1995b).

*** Cephalomitrionaterrimum (Steph.) R.M.Schust., Nova Hedwigia 61 (3/4): 554, 1995 (Schuster 1995b). Bas.: Cephaloziaaterrima Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 501 (331), 1908 (Stephani 1908f).

*** Cephaloziella (Spruce) Schiffn., Hepat. (Engl.-Prantl): 98, 1893 (Schiffner 1893b) nom. conserv. Bas.: Cephaloziasubg.Cephaloziella Spruce, Cephalozia: 62, 1882 (Spruce 1882). 41

** subg.Cephaloziella

*** Cephaloziellaaenigmatica R.M.Schust., Nova Hedwigia 63 (1/2): 20, 1996 (Schuster 1996d).

*** Cephaloziellaanthelioides S.W.Arnell, Bot. Not. 105: 322, 1952 (Arnell 1952a).

*** Cephaloziellaarctogena (R.M.Schust.) Konstant., Arctoa 3: 126, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Cephaloziellarubellavar.arctogena R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 125, 1980 (Schuster 1980c).

** Cephaloziellaarenaria (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (Schuster 1963b). Bas.: Cephaloziaarenaria Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 504 (334), 1908 (Stephani 1908f).

*** Cephaloziellaaspericaulis Jørg., Bergens Mus. Skr. (n.ser.) 16: 197, 1934 (Jørgensen 1934).

*** Cephaloziellabaumgartneri Schiffn., Verh. K.K. Zool.-Bot. Ges. Wien 56 (3): 273, 1906 (Schiffner 1906c).

** Cephaloziellabreviperianthia C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 208, 1981 (Gao and Chang 1981).

** Cephaloziellabrinkmannii Douin, Mém. Soc. Bot. France 29: 75, 1920 (Douin 1920).

*** Cephaloziellacapensis (Sim) S.W.Arnell, Bot. Not. 105: 326, 1952 (Arnell 1952a). Bas.: Cephaloziacapensis Sim, Trans. Roy. Soc. South Africa 15 (1): 87, 1926 (Sim 1926).

*** Cephaloziellacapillaris (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920). Bas.: Cephaloziacapillaris Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 496 (326), 1908 (Stephani 1908f).

*** Cephaloziellacrassigyna (R.M.Schust.) R.M.Schust., Nova Hedwigia 61 (3/4): 556, 1995 (Schuster 1995b). Bas.: Cephaloziellaaterrimavar.crassigyna R.M.Schust., Nova Hedwigia 22: 211, 1971 [1972] (Schuster 1971b).

*** Cephaloziellacrispata N.Kitag., J. Hattori Bot. Lab. 32: 301, 1969 (Kitagawa 1969c).

*** Cephalozielladensifolia R.M.Schust., Nova Hedwigia 22: 199, 1971 [1972] (Schuster 1971b).

** Cephalozielladensifoliavar.dubia R.M.Schust., Nova Hedwigia 22: 202, 1971 [1972] (Schuster 1971b).

*** Cephalozielladivaricata (Sm.) Schiffn., Hepat. (Engl.-Prantl): 99, 1893 (Schiffner 1893b). Bas.: Jungermanniadivaricata Sm., Engl. Bot. 10: tab. 719, 1800 (Smith and Sowerby 1800).

** Cephalozielladivaricatavar.scabra (M.Howe) Haynes, Bryologist 12 (4): 68, 1909 (Haynes 1909). Bas.: Cephaloziadivaricatavar.scabra M.Howe, Mem. Torrey Bot. Club 7: 129, 1899 (Howe 1899). 42

** Cephalozielladusenii Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 6): 49, 1900 (Stephani 1900b).

*** Cephaloziellaelachista (J.B.Jack ex Gottsche et Rabenh.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (Schiffner 1900d). Bas.: Jungermanniaelachista J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 58-59: no. 574, 1873 (Gottsche and Rabenhorst 1873a).

** Cephaloziellaelachistavar.spinophylla (C.Gao) C.Gao, Fl. Bryoph. Sin. 9: 181, 2003 (Gao 2003). Bas.: Cephaloziellaspinophylla C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 208, 1981 (Gao and Chang 1981).

** Cephaloziellaelegans (Heeg) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 336, 1900 (Schiffner 1900d). Bas.: Cephaloziaelegans Heeg, Rev. Bryol. 20 (5): 82, 1893 (Heeg 1893).

*** Cephaloziellaexigua R.M.Schust., Nova Hedwigia 63 (1/2): 49, 1996 (Schuster 1996d).

*** Cephaloziellaexiliflora (Taylor) Douin, Mém. Soc. Bot. France 29: 72, 1920 (Douin 1920). Bas.: Jungermanniaexiliflora Taylor, London J. Bot. 5: 279, 1846 (Taylor 1846a).

** Cephaloziellafragillima (Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 409, 1976 (Fulford 1976). Bas.: Cephaloziafragillima Spruce, Mem. Torrey Bot. Club 1 (3): 131, 1890 (Spruce 1890).

*** Cephaloziellagarsidei S.W.Arnell, Rev. Bryol. Lichénol. 23 (1/2): 173, 1954 (Arnell 1954a).

*** Cephaloziellagrandiretis (R.M.Schust.) R.M.Schust., Nova Hedwigia 63 (1/2): 35, 1996 (Schuster 1996d). Bas.: Cephaloziellabyssaceasubsp.grandiretis R.M.Schust., Nova Hedwigia 22: 195, 1971 [1972] (Schuster 1971b).

*** Cephaloziellagrimsulana (J.B.Jack ex Gottsche et Rabenh.) Lacout., Hépat. France: 52, 1905 (Lacouture 1905). Bas.: Jungermanniagrimsulana J.B.Jack ex Gottsche et Rabenh., Hepat. Eur., Leberm. 53–55: no. 526, 1872 (Gottsche and Rabenhorst 1872).

*** Cephaloziellahampeana (Nees) Schiffn. ex Loeske, Moosfl. Harz.: 92, 1903 (Loeske 1903). Bas.: Jungermanniahampeana Nees, Naturgesch. Eur. Leberm. 3: 560, 1838 (Nees 1838b).

** Cephaloziellahebridensis Steph., Hedwigia 32 (5): 316, 1893 (Stephani 1893d).

*** Cephaloziellaherzogiana (Pandé et K.P.Srivast.) Udar et D.Kumar, Geophytology 6 (1): 45, 1976 (Udar and Kumar 1976). Bas.: Cephaloziaherzogiana Pandé et K.P.Srivast., Feddes Repert. Spec. Nov. Regni Veg. 58: 75, 1955 (Pandé and Srivastava 1955).

** Cephaloziellaheteroica (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 76, 1920 (Douin 1920). Bas.: Cephaloziaheteroica C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 38, 1904 (Cooke 1904).

** Cephaloziellahyalina Douin, Mém. Soc. Bot. France 29: 77, 1920 (Douin 1920).

* Cephaloziellahyalinavar.rappii (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 56, 1980 (Schuster 1980c). Bas.: Cephaloziellarappii Douin, Mém. Soc. Bot. France 29: 77, 1920 (Douin 1920).

*** Cephaloziellainaequalis R.M.Schust., Nova Hedwigia 22: 186, 1971 [1972] (Schuster 1971b).

*** Cephaloziellainvisa R.M.Schust., Nova Hedwigia 63 (1/2): 30, 1996 (Schuster 1996d).

** Cephaloziellakilohanensis (C.M.Cooke) Douin, Mém. Soc. Bot. France 29: 85, 1920 (Douin 1920). Bas.: Cephaloziakilohanensis C.M.Cooke, Trans. Connecticut Acad. Arts 12 (1): 37, 1904 (Cooke 1904).

*** Cephaloziellalongii Váňa, Folia Geobot. Phytotax. 27 (2): 193, 1992 (Váňa 1992).

*** Cephaloziellalycopodioides (Sim) S.W.Arnell, Bot. Not. 105: 321, 1952 (Arnell 1952a). Bas.: Cephalozialycopodioides Sim, Trans. Roy. Soc. South Africa 15 (1): 85, 1926 (Sim 1926).

** Cephaloziellamammillifera R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987).

*** Cephaloziellamassalongi (Spruce) Müll.Frib., Lebermoose 2 (17): 191, 1913 (Müller 1913a). Bas.: Cephaloziamassalongi Spruce, Cephalozia: 71, 1882 (Spruce 1882).

*** Cephaloziellamuelleriana R.M.Schust., Nova Hedwigia 63 (1/2): 24, 1996 (Schuster 1996d).

*** Cephaloziellanatalensis (Sim) S.W.Arnell, Hepat. South Africa: 338, 1963 (Arnell 1963b). Bas.: Cephalozianatalensis Sim, Trans. Roy. Soc. South Africa 15 (1): 86, 1926 (Sim 1926).

** Cephaloziellanicholsonii Douin, Rev. Bryol. 40 (6): 81, 1913 (Douin 1913b).

*** Cephaloziellanothogena R.M.Schust., Nova Hedwigia 63 (1/2): 37, 1996 (Schuster 1996d).

** Cephaloziellaobtusilobula R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 108, 1980 (Schuster 1980c).

** Cephaloziellapatulifolia (Steph.) Douin, Mém. Soc. Bot. France 29: 70, 1920 (Douin 1920). Bas.: Cephaloziapatulifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 509 (339), 1908 (Stephani 1908f).

*** Cephaloziellaphyllacantha (C.Massal. et Carestia) Müll.Frib., Lebermoose 2 (17): 194, 1913 (Müller 1913a). Bas.: Antheliaphyllacantha C.Massal. et Carestia, Nuovo Giorn. Bot. Ital. 12 (4): 340, 1880 (Massalongo and Carestia 1880).

*** Cephaloziellapolystratosa (R.M.Schust. et Damsh.) Konstant., Bot. Zhurn. (Moscow & Leningrad) 85 (10): 127, 2000 (Konstantinova 2000). Bas.: Cephaloziellabyssaceavar.polystratosa R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987).

*** Cephaloziellapseudocrassigyna R.M.Schust. ex J.J.Engel, Novon 17 (3): 313, 2007 (Engel 2007).

*** Cephaloziellapulcherrima R.M.Schust., Nova Hedwigia 22: 203, 1971 [1972] (Schuster 1971b).

** Cephaloziellapulcherrimasubsp.sphagnicola R.M.Schust., Nova Hedwigia 22: 205, 1971 [1972] (Schuster 1971b).

** Cephaloziellapungens Steph. ex Fulford, Mem. New York Bot. Gard. 11 (4): 410, 1976 (Fulford 1976).

*** Cephaloziellarubella (Nees) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 231, 1902 (Warnstorf 1902). Bas.: Jungermanniarubella Nees, Naturgesch. Eur. Leberm. 2: 236, 1836 (Nees 1836).

*** Cephaloziellaschelpei S.W.Arnell, Bot. Not. 110 (1): 18, 1957 (Arnell 1957a).

*** Cephaloziellaspegazziniana (C.Massal.) Douin, Mém. Soc. Bot. France 29: 69, 1920 (Douin 1920). Bas.: Cephaloziaspegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 234, 1885 (Massalongo 1885).

*** Cephaloziellaspinicaulis Douin, Mém. Soc. Bot. France 29: 62, 1920 (Douin 1920).

*** Cephaloziellaspinigera (Lindb.) Jørg., Bergens Mus. Skr. (n.ser.) 16: 189, 1934 (Jørgensen 1934). Bas.: Cephaloziaspinigera Lindb., Musci Scand.: 4, 1879 (Lindberg 1879).

*** Cephaloziellastellulifera (Taylor ex Carrington et Pearson) Croz., Rev. Bryol. 30 (2): 31, 1903 (Crozals 1903a). Bas.: Jungermanniastellulifera Taylor ex Carrington et Pearson, Hepat. Brit. Exsicc. Fasc. I: no. 32, 1878 (Carrington and Pearson 1878).

*** Cephaloziellastephanii Schiffn. ex Douin, Mém. Soc. Bot. France 29: 85, 1920 (Douin 1920).

** Cephaloziellastolonifera R.M.Schust., Phytologia 39 (6): 426, 1978 (Schuster 1978b).

** Cephaloziellasubtilis (Lindenb. et Gottsche) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Jungermanniasubtilis Lindenb. et Gottsche, Syn. Hepat. 5: 685, 1847 (Gottsche et al. 1847).

*** Cephaloziellasumatrana Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920).

*** Cephaloziellatabularis S.W.Arnell, Bot. Not. 105: 318, 1952 (Arnell 1952a).

*** Cephaloziellatransvaalensis S.W.Arnell, Bot. Not. 110 (1): 19, 1957 (Arnell 1957a).

*** Cephaloziellatriplicata S.W.Arnell, Bot. Not. 115: 203, 1962 (Arnell 1962a).

*** Cephaloziellaumtaliensis S.W.Arnell, Bot. Not. 110 (1): 20, 1957 (Arnell 1957a).

*** Cephaloziellauncinata R.M.Schust., Meddel. Grønland 199 (1): 316, 1974 (Schuster and Damsholt 1974).

** Cephaloziellauncinatavar.brevigyna R.M.Schust. et Damsh., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987).

** Cephaloziellauncinatavar.sphagnicola R.M.Schust., Meddel. Grønland 199 (1): 323, 1974 (Schuster and Damsholt 1974).

*** Cephaloziellavaginans Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 119, 1911 (Stephani 1911a).

*** Cephaloziellavarians (Gottsche) Steph., Wiss. Ergebn. Schwed. Südpolar-Exped. [1901–1903] 4 (1): 4, 1905 (Stephani 1905e). Bas.: Jungermanniavarians Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 452, 1890 (Gottsche 1890).

*** Cephaloziellaverrucosa Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d).

* Cephaloziellavillaumei (Steph.) Váňa, Phytotaxa 112 (1): 3, 2013 (Váňa et al. 2013e). Bas.: Cephaloziavillaumei Steph., Sp. Hepat. (Stephani) 6: 437, 1924 (Stephani 1924). 43

** Cephaloziellaviolacea Schljakov, Novosti Sist. Nizš. Rast. 15: 242, 1978 (Shliakov 1978).

** Cephaloziellawelwitschii (Steph.) Douin, Mém. Soc. Bot. France 29: 58, 1920 (Douin 1920). Bas.: Cephaloziawelwitschii Steph., Bull. Herb. Boissier (sér. 2) 8 (6): 432 (308), 1908 (Stephani 1908g).

** subg.Dichiton (Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (Müller 1916). Bas.: Dichiton Mont., Syll. Gen. Sp. Crypt.: 52, 1856 (Montagne 1856b).

*** Cephaloziellacalyculata (Durieu et Mont.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (Müller 1916). Bas.: Jungermanniacalyculata Durieu et Mont., Ann. Sci. Nat. Bot. (sér. 3) 11: 34, 1849 (Montagne 1849).

*** Cephaloziellaintegerrima (Lindb.) Warnst., Krypt.-Fl. Brandenburg, Leber- & Torfmoose: 232, 1902 (Warnstorf 1902). Bas.: Cephaloziaintegerrima Lindb., Acta Soc. Sci. Fenn. 10: 502, 1875 (Lindberg 1875).

** subg.Evansia (Douin et Schiffn.) Müll.Frib., Lebermoose 2 (27): 787, 1916 (Müller 1916). Bas.: Evansia Douin et Schiffn., Rev. Bryol. 40 (5): 66, 1913 (Douin 1913a).

*** Cephaloziellaantillana (Besch. et Spruce) Fulford, Mem. New York Bot. Gard. 11 (4): 405, 1976 (Fulford 1976). Bas.: Cephaloziaantillana Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxiii, 1889 [1890] (Bescherelle and Spruce 1889).

*** Cephalozielladentata (Raddi) Steph., Bull. Herb. Boissier 5 (2): 78, 1897 (Stephani 1897b). Bas.: Jungermanniadentata Raddi, Jungermanniogr. Etrusca: 21, 1818 (Raddi 1818a).

*** Cephaloziellahirta (Steph.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (Schuster 1963b). Bas.: Cephaloziahirta Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 561 (345), 1908 (Stephani 1908e).

** Cephaloziellasquarrosula (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Cephaloziasquarrosula Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877).

*** Cephaloziellasubspinosa R.M.Schust., Nova Hedwigia 22: 210, 1971 [1972] (Schuster 1971b).

** subg.Prionolobus (Spruce) Müll.Frib., Lebermoose 2 (16): 110, 1912 (Müller 1912). Bas.: Cephaloziasubg.Prionolobus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 508, 1885 (Spruce 1885).

*** Cephaloziellaacanthophora (S.Hatt.) Horik., Hikobia 1 (2): 79, 1951 (Horikawa 1951c). Bas.: Prionolobusacanthophorus S.Hatt., Bull. Tokyo Sci. Mus. 11: 29, 1944 (Hattori 1944d).

*** Cephaloziellabiokoensis Váňa et Frank Müll., Trop. Bryol. 24: 1, 2003 (Váňa and Müller 2003).

*** Cephaloziellagranatensis (J.B.Jack ex Steph.) Fulford, Mem. New York Bot. Gard. 11 (4): 411, 1976 (Fulford 1976). Bas.: Cephaloziagranatensis J.B.Jack ex Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 500 (330), 1908 (Stephani 1908f).

*** Cephaloziellagrisea R.M.Schust., Phytologia 39 (6): 425, 1978 (Schuster 1978b).

* Cephaloziellameghalayensis Udar et Ad.Kumar, Lindbergia 8 (1): 34, 1982 (Udar and Kumar 1982d). 44

*** Cephaloziellamicrophylla (Steph.) Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920). Bas.: Cephaloziamicrophylla Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 513 (343), 1908 (Stephani 1908f).

*** Cephaloziellatenuissima (Lehm.) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Jungermanniatenuissima Lehm., Linnaea 4: 367, 1829 (Lehmann 1829).

*** Cephaloziellaturneri (Hook.) Müll.Frib., Lebermoose 2 (17): 202, 1913 (Müller 1913a). Bas.: Jungermanniaturneri Hook., Brit. Jungermann.: tab. 29, 1812 (Hooker 1812).

Incertae sedis

** Cephalozielladentifolia Udar et Ad.Kumar, Lindbergia 8 (1): 30, 1982 (Udar and Kumar 1982d).

** Cephaloziellafilum (Trevis.) Steph., Hedwigia 32 (5): 318, 1893 (Stephani 1893d). Bas.: Cephaloziafilum Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 417, 1877 (Trevisan 1877).

* Cephaloziellaflexuosa C.Gao et K.C.Chang, Bull. Bot. Res., Harbin 4 (3): 88, 1984 (Chang and Gao 1984). 45

** Cephaloziellaintricata Schiffn. ex Douin, Mém. Soc. Bot. France 29: 59, 1920 (Douin 1920).

** Cephaloziellalevieri Schiffn. ex Douin, Mém. Soc. Bot. France 29: 80, 1920 (Douin 1920).

*** Cephaloziellapellucida R.M.Schust., Nova Hedwigia 63 (1/2): 57, 1996 (Schuster 1996d). 46

* Cephaloziellapygmaea (Spruce) Váňa, Phytotaxa 183 (4): 290, 2014 (Váňa et al. 2014a). Bas.: Cephaloziapygmaea Spruce, Cephalozia: 69, 1882 (Spruce 1882). 47

* Cephaloziellasecundifolia Pearson, Ann. Bryol. 4: 106, 1931 (Pearson 1931b). 48

** Cephaloziellasinensis Douin, Rev. Bryol. 41 (1): 8, 1914 (Douin 1914).

** Cephaloziopsis (Spruce) Schiffn., Hepat. (Engl.-Prantl): 85, 1893 (Schiffner 1893b). Bas.: Jungermanniasect.Cephaloziopsis Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 511, 1885 (Spruce 1885).

*** Cephaloziopsisintertexta (Gottsche) R.M.Schust., Nova Hedwigia 22: 183, 1971 [1972] (Schuster 1971b). Bas.: Jungermanniaintertexta Gottsche, Syn. Hepat. 1: 107, 1844 (Gottsche et al. 1844).

** Chaetophyllopsis R.M.Schust., J. Hattori Bot. Lab. 23: 69, 1960 [1961] (Schuster 1960b).

*** Chaetophyllopsiswhiteleggei (Carrington et Pearson) R.M.Schust. ex Hamlin, Rec. Domin. Mus. 7: 255, 1972 (Hamlin 1972). Bas.: Jungermanniawhiteleggei Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1051, 1888 (Carrington and Pearson 1888a).

** Cylindrocolea R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a).

** subg.Cylindrocolea

*** Cylindrocoleakiaeri (Austin) Váňa, Phytotaxa 112 (1): 2, 2013 (Váňa et al. 2013e). Bas.: Jungermanniakiaeri Austin, Bull. Torrey Bot. Club 6 (3): 18, 1875 (Austin 1875b).

*** Cylindrocoleasanctae-helenae M.Wigginton, Polish Bot. J. 58 (1): 107, 2013 (Wigginton 2013).

** sect.Cylindrocolea

*** Cylindrocoleaandersonii R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 33, 1980 (Schuster 1980c).

*** Cylindrocoleabrasiliensis D.P.Costa, N.D.Santos et Váňa, Bryologist 111 (4): 667, 2008 (Costa et al. 2008).

*** Cylindrocoleachevalieri (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). Bas.: Alobiellachevalieri Steph., Bull. Herb. Boissier (sér. 2) 8 (8): 567 (351), 1908 (Stephani 1908e).

*** Cylindrocoleagittinsii (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziellagittinsii E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 437, 1958 (Jones 1958).

*** Cylindrocoleamadagascariensis (Steph.) R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 12 (3): 666, 1969 (Schuster 1969a). Bas.: Lophoziamadagascariensis Steph., Sp. Hepat. (Stephani) 6: 112, 1917 (Stephani 1917a).

*** Cylindrocoleanigerica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 172, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziellanigerica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 435, 1958 (Jones 1958).

*** Cylindrocoleanovae-caledoniae (Grolle) R.M.Schust., Nova Hedwigia 22: 161, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziellanovae-caledoniae Grolle, Rev. Bryol. Lichénol. 29 (3/4): 208, 1960 [1961] (Grolle 1960a).

*** Cylindrocoleaplanifolia (Steph.) R.M.Schust., Nova Hedwigia 22: 164, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziellaplanifolia Steph., Hedwigia 32 (5): 317, 1893 (Stephani 1893d).

*** Cylindrocoleasprucei R.M.Schust., Nova Hedwigia 22: 163, 1971 [1972] (Schuster 1971b).

*** Cylindrocoleaugandica (E.W.Jones) R.M.Schust., Nova Hedwigia 22: 171, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziellaugandica E.W.Jones, Trans. Brit. Bryol. Soc. 3 (3): 433, 1958 (Jones 1958).

** sect.Platycaulis R.M.Schust., Nova Hedwigia 22: 173, 1971 (Schuster 1971b).

*** Cylindrocolearecurvifolia (Steph.) Inoue, J. Jap. Bot. 47 (11): 348, 1972 (Inoue 1972a). Bas.: Cephaloziarecurvifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (7): 497 (327), 1908 (Stephani 1908f).

*** Cylindrocoleatagawae (N.Kitag.) R.M.Schust., Nova Hedwigia 22: 174, 1971 [1972] (Schuster 1971b). Bas.: Cephaloziellatagawae N.Kitag., J. Hattori Bot. Lab. 32: 303, 1969 (Kitagawa 1969c).

** subg.Cylindroscyphus (Douin) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 20, 1980 (Schuster 1980c). Bas.: Cephaloziellasubg.Cylindroscyphus Douin, Mém. Soc. Bot. France 29: 56, 1920 (Douin 1920).

*** Cylindrocoleaabyssinica (Gola) Váňa, Phytotaxa 112 (1): 2, 2013 (Váňa et al. 2013e). Bas.: Cephaloziellaabyssinica Gola, Ann. Bot. (Rome) 13 (1): 68, 1914 (Gola 1914a).

** Cylindrocoleaobtusifolia Fulford, Mem. New York Bot. Gard. 11 (4): 401, 1976 (Fulford 1976).

** Cylindrocoleareticulata Udar et Ad.Kumar, Lindbergia 8 (3): 181, 1982 [1983] (Udar and Kumar 1982c).

*** Cylindrocolearhizantha (Mont.) R.M.Schust., Nova Hedwigia 22: 175, 1971 [1972] (Schuster 1971b). Bas.: Jungermanniarhizantha Mont., Hist. Phys. Cuba, Bot., Pl. Cell.: 454, 1842 (Montagne 1842a).

*** Gottschelia Grolle, J. Hattori Bot. Lab. 31: 13, 1968 (Grolle 1968c).

*** Gottscheliamaxima (Steph.) Grolle, J. Bryol. 25 (1): 6, 2003 (Grolle et al. 2003). Bas.: Tylimanthusmaximus Steph., Sp. Hepat. (Stephani) 6: 249, 1922 (Stephani 1922).

*** Gottscheliaschizopleura (Spruce) Grolle, J. Hattori Bot. Lab. 31: 16, 1968 (Grolle 1968c). Bas.: Jungermanniaschizopleura Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 517, 1885 (Spruce 1885).

** Gymnocoleopsis (R.M.Schust.) R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a). Bas.: Gymnocoleasubg.Gymnocoleopsis R.M.Schust., Bryologist 70 (1): 111, 1967 (Schuster 1967b).

*** Gymnocoleopsiscapensis (S.W.Arnell) R.M.Schust., J. Hattori Bot. Lab. 78: 123, 1995 (Schuster 1995c). Bas.: Lophoziacapensis S.W.Arnell, Svensk Bot. Tidskr. 47 (1): 112, 1953 (Arnell 1953a).

*** Gymnocoleopsiscylindriformis (Mitt.) R.M.Schust., J. Hattori Bot. Lab. 78: 126, 1995 (Schuster 1995c). Bas.: Jungermanniacylindriformis Mitt., J. Linn. Soc., Bot. 15 (84): 196, 1876 (Mitten 1876b).

*** Herzogobryum Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (Grolle 1963d). Nom. nov. pro Chondrophyllum Herzog, Rev. Bryol. Lichénol. 21 (1/2): 46, 1952 (Herzog 1952f).

*** Herzogobryumatrocapillum (Hook.f. et Taylor) Grolle, Österr. Bot. Z. 113 (2): 228, 1966 (Grolle 1966j). Bas.: Gymnomitrionatrocapillum Hook.f. et Taylor, London J. Bot. 5: 258, 1846 (Taylor 1846a).

*** Herzogobryumfiliforme R.M.Schust., Phytologia 45 (5): 422, 1980 (Schuster 1980b).

*** Herzogobryummolle Grolle, Österr. Bot. Z. 113 (2): 226, 1966 (Grolle 1966j).

*** Herzogobryumvermiculare (Schiffn.) Grolle, J. Hattori Bot. Lab. 28: 103, 1965 (Grolle 1965e). Bas.: Gymnomitrionvermiculare Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 2, 1890 (Schiffner 1890).

** Kymatocalyx Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 56, 1950 (Herzog 1950c).

*** Kymatocalyxafricanus Váňa et M.Wigginton, Haussknechtia, Beih. 9: 158, 1999 (Gradstein and Váňa 1999).

*** Kymatocalyxdominicensis (Spruce) Váňa, Österr. Bot. Z. 118 (5): 575, 1970 (Váňa 1970b). Bas.: Jungermanniadominicensis Spruce, J. Linn. Soc., Bot. 30 (210): 363, 1895 (Gepp 1895b).

*** Kymatocalyxmadagascariensis (Steph.) Gradst. et Váňa, Haussknechtia, Beih. 9: 164, 1999 (Gradstein and Váňa 1999). Bas.: Acrobolbusmadagascariensis Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902 (Stephani 1902f).

*** Kymatocalyxrhizomaticus (Herzog) Gradst. et Váňa, Haussknechtia, Beih. 9: 166, 1999 (Gradstein and Váňa 1999). Bas.: Stenorrhipisrhizomatica Herzog, Trans. Brit. Bryol. Soc. 1 (4): 290, 1950 (Herzog 1950a).

*** Lophonardia R.M.Schust., Phytologia 39 (4): 244, 1978 (Schuster 1978a).

*** Lophonardiajamesonii (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 19, 2013 (Söderström et al. 2013a). Bas.: Jungermanniajamesonii Mont., Syll. Gen. Sp. Crypt.: 60, 1856 (Montagne 1856b).

*** Lophonardialaxifolia (Mont.) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (Söderström et al. 2013a). Bas.: Sarcocyphoslaxifolius Mont., Ann. Sci. Nat. Bot. (sér. 3) 4: 346, 1845 (Montagne 1845b).

*** Lophonardiatristaniana (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 81 (1): 20, 2013 (Söderström et al. 2013a). Bas.: Jungermanniatristaniana S.W.Arnell, Results Norweg. Sci. Exped. Tristan da Cunha 42: 12, 1958 (Arnell 1958b).

*** Nothogymnomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 43, 1996 (Schuster 1996a).

*** Nothogymnomitrionerosum (Carrington et Pearson) R.M.Schust., J. Hattori Bot. Lab. 80: 44, 1996 (Schuster 1996a). Bas.: Cesiuserosus Carrington et Pearson, Pap. & Proc. Roy. Soc. Tasmania 1887: 8, 1888 (Carrington and Pearson 1888b).

*** Obtusifolium S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 309, 1956 (Arnell 1956b).

*** Obtusifoliumobtusum (Lindb.) S.W.Arnell, Ill. Moss Fl. Fennosc. Hep.: 133, 1956 (Arnell 1956b). Bas.: Jungermanniaobtusa Lindb., Musci Scand.: 7, 1879 (Lindberg 1879).

*** Oleolophozia L.Söderstr., De Roo et Hedd., Phytotaxa 3: 50, 2010 (Söderström et al. 2010b).

*** Oleolophoziaperssonii (H.Buch et S.W.Arnell) L.Söderstr., De Roo et Hedd., Phytotaxa 3: 51, 2010 (Söderström et al. 2010b). Bas.: Lophoziaperssonii H.Buch et S.W.Arnell, Bot. Not. 97: 382, 1944 (Buch 1944).

** Phycolepidozia R.M.Schust., Bull. Torrey Bot. Club 93 (6): 438, 1966 [1967] (Schuster 1966f).

** subg.Metaphycolepidozia Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 506, 2014 (Gradstein et al. 2014a).

** Phycolepidoziaindica Gradst., J.-P.Frahm et U.Schwarz, Taxon 63 (3): 499, 2014 (Gradstein et al. 2014a).

** subg.Phycolepidozia

*** Phycolepidoziaexigua R.M.Schust., Bull. Torrey Bot. Club 93 (6): 440, 1966 [1967] (Schuster 1966f).

** Protolophozia (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (Shliakov 1979). Bas.: Lophoziasubg.Protolophozia R.M.Schust., Nova Hedwigia 15: 472, 1968 (Schuster 1968b).

** Protolophoziaandrogyna R.M.Schust. ex Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Based on: Lophoziaandrogyna R.M.Schust., Beih. Nova Hedwigia 119: 266, 2002 (Schuster 2002b), nom. inval.

** Protolophoziaautoica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophoziaautoica R.M.Schust., Nova Hedwigia 15: 479, 1968 (Schuster 1968b).

** Protolophoziacrispata (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophoziacrispata R.M.Schust., Nova Hedwigia 15: 474, 1968 (Schuster 1968b).

*** Protolophoziadruceae (Grolle et E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophoziadruceae Grolle et E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 112, 1972 (Hodgson 1972).

*** Protolophoziaelongata (Steph.) Schljakov, Novosti Sist. Nizš. Rast. 16: 204, 1979 (Shliakov 1979). Bas.: Lophoziaelongata Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 41 (141), 1902 (Stephani 1902c).

*** Protolophoziaherzogiana (E.A.Hodgs. et Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Lophoziaherzogiana E.A.Hodgs. et Grolle, Rev. Bryol. Lichénol. 31 (3/4): 152, 1962 [1963] (Grolle 1962c).

*** Protolophozialancistipa (Grolle) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Andrewsianthuslancistipus Grolle, Marion Prince Edw. Is: 230, 1971 (Grolle 1971d).

*** Protolophozialeucorhiza (Mitt.) Váňa et L.Söderstr., Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Jungermannialeucorhiza Mitt., J. Linn. Soc., Bot. 15 (82): 68, 1876 (Mitten 1876a).

*** Protolophozialongiflora (Herzog) L.Söderstr. et Váňa, Phytotaxa 76 (3): 51, 2013 (Váňa et al. 2013l). Bas.: Orthocaulislongiflorus Herzog, Rev. Bryol. Lichénol. 23 (1/2): 32, 1954 (Herzog 1954).

** Protolophoziamonoica (E.A.Hodgs.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Metahygrobiellamonoica E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 76, 1965 (Hodgson 1965).

*** Protolophoziamulticuspidata (Hook.f. et Taylor) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Jungermanniamulticuspidata Hook.f. et Taylor, London J. Bot. 3: 375, 1844 (Hooker and Taylor 1844a).

** Protolophozianivicola (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophozianivicola R.M.Schust., Nova Hedwigia 15: 477, 1968 (Schuster 1968b).

*** Protolophoziaperssoniana (H.A.Mill.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophoziaperssoniana H.A.Mill., Ark. Bot. (n.ser.) 5 (2): 508, 1963 (Miller 1963).

** Protolophoziasubalpina (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophoziaautoicavar.subalpina R.M.Schust., Nova Hedwigia 15: 482, 1968 (Schuster 1968b).

*** Protolophoziatasmanica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophoziatasmanica R.M.Schust., Nova Hedwigia 15: 484, 1968 (Schuster 1968b).

*** Protolophoziaverruculosa (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 52, 2013 (Váňa et al. 2013l). Bas.: Lophoziaverruculosa R.M.Schust., Phytologia 39 (4): 242, 1978 (Schuster 1978a).

*** Lophoziaceae Cavers

by L. Söderström and J. Váňa

Lophoziaceae was broadly defined by de Roo et al. (2007) excluding Anastrophyllaceae and Jamesonielloideae and also moving several traditionally included genera to other families. Vilnet et al. (2010) further refined the family. Nomenclatural and taxonomic notes can also be found in Vilnet et al. (2007b, 2008), Söderström et al. (2013c) and Váňa et al. (2013m). The placement of Gerhildiella and Pseudocephaloziella in the family is provisional.

*** Andrewsianthus R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 66, 1961 (Schuster 1961a).

** subg.Andrewsianthus

*** Andrewsianthusaberrans (Nees et Mont.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 440, 1963 (Grolle 1963a). Bas.: Jungermanniaaberrans Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 250, 1843 (Montagne 1843).

*** Andrewsianthusbidens (Mitt. ex Steph.) R.M.Schust., Nova Hedwigia 15: 492, 1968 (Schuster 1968b). Bas.: Lophoziabidens Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 41 (141), 1902 (Stephani 1902c).

*** Andrewsianthusbilobus (Mitt.) Grolle, Trans. Brit. Bryol. Soc. 4 (3): 437, 1963 (Grolle 1963a). Bas.: Gymnanthebiloba Mitt., J. Proc. Linn. Soc., Bot. 7 (27): 166, 1863 (Mitten 1863).

*** Andrewsianthuscarinatus Grolle, Marion Prince Edw. Is: 229, 1971 (Grolle 1971d).

*** Andrewsianthuscavifolius Grolle et Váňa, J. Hattori Bot. Lab. 38: 640, 1974 (Váňa 1974b).

*** Andrewsianthuschimbuensis R.M.Schust., Nova Hedwigia 15: 491, 1968 (Schuster 1968b).

*** Andrewsianthuskinabaluensis N.Kitag., J. Hattori Bot. Lab. 33: 205, 1970 (Kitagawa 1970).

*** Andrewsianthuskoponenii Váňa et Piippo, Ann. Bot. Fenn. 26 (3): 284, 1989 (Váňa and Piippo 1989).

*** Andrewsianthusmizutanii N.Kitag., J. Hattori Bot. Lab. 32: 307, 1969 (Kitagawa 1969a).

*** Andrewsianthuspapillosus N.Kitag., J. Hattori Bot. Lab. 33: 207, 1970 (Kitagawa 1970).

*** Andrewsianthusperigonialis (Hook.f. et Taylor) R.M.Schust., Beih. Nova Hedwigia 119: 336, 2002 (Schuster 2002b). Bas.: Jungermanniaperigonialis Hook.f. et Taylor, London J. Bot. 3: 368, 1844 (Hooker and Taylor 1844a).

*** Andrewsianthuspuniceus (Nees) R.M.Schust. ex Grolle, Trans. Brit. Bryol. Soc. 4 (3): 439, 1963 (Grolle 1963a). Bas.: Jungermanniapunicea Nees, Enum. Pl. Crypt. Javae: 32, 1830 (Nees 1830).

*** Andrewsianthusrecurvifolius (Nees) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (Schuster 1969b). Bas.: Jungermanniarecurvifolia Nees, Enum. Pl. Crypt. Javae: 32, 1830 (Nees 1830).

*** Andrewsianthussundaicus (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 710, 1969 (Schuster 1969b). Bas.: Anastrophyllumsundaicum Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 202, 1898 (Schiffner 1898a).

*** Andrewsianthuszantenii Váňa, J. Hattori Bot. Lab. 38: 645, 1974 (Váňa 1974b).

** subg.Cephalolobus (R.M.Schust.) R.M.Schust., Beih. Nova Hedwigia 119: 326, 2002 (Schuster 2002b). Bas.: Cephalolobus R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 244, 1966 (Schuster 1966b).

*** Andrewsianthushodgsoniae (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Bas.: Cephalolobushodgsoniae R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 254, 1966 (Schuster 1966b).

*** Andrewsianthusmarionensis (S.W.Arnell) Grolle, Marion Prince Edw. Is: 232, 1971 (Grolle 1971d). Bas.: Lophoziamarionensis S.W.Arnell, Svensk Bot. Tidskr. 47 (3): 421, 1953 (Arnell 1953c).

*** Andrewsianthusscabrellus (C.Massal.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Bas.: Cephaloziascabrella C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 233, 1885 (Massalongo 1885).

** Andrewsianthussphenoloboides (R.M.Schust.) R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Bas.: Cephalolobussphenoloboides R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 251, 1966 (Schuster 1966b).

** Gerhildiella Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (Grolle 1966a).

*** Gerhildiellarossneriana Grolle, Rev. Bryol. Lichénol. 34 (1/2): 187, 1966 (Grolle 1966a).

*** Heterogemma (Jørg.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziasect.Heterogemma Jørg., Bergens Mus. Skr. (n.ser.) 16: 146, 1934 (Jørgensen 1934).

*** Heterogemmacapitata (Hook.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermanniacapitata Hook., Brit. Jungermann.: tab. 80, 1816 (Hooker 1816a).

*** Heterogemmalaxa (Lindb.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannialaxa Lindb., Acta Soc. Sci. Fenn. 10: 529, 1875 (Lindberg 1875).

*** Heterogemmapatagonica (Herzog et Grolle) L.Söderstr. et Váňa, Phytotaxa 97 (2): 29, 2013 (Söderström et al. 2013c). Bas.: Lophoziapatagonica Herzog et Grolle, Rev. Bryol. Lichénol. 28 (3/4): 343, 1959 [1960] (Grolle 1959a).

*** Lophozia (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermanniasect.Lophozia Dumort., Syll. Jungerm. Europ.: 53, 1831 (Dumortier 1831). 49

*** Lophoziaascendens (Warnst.) R.M.Schust., Bryologist 55 (3): 180, 1952 (Schuster 1952). Bas.: Sphenolobusascendens Warnst., Hedwigia 57 (1/2): 63, 1916 (Warnstorf 1916).

* Lophoziaaustrosibirica Bakalin, Ann. Bot. Fenn. 40 (1): 49, 2003 (Bakalin 2003).

** Lophoziaciliata Damsh., L.Söderstr. et H.Weibull, Lindbergia 25 (1): 3, 2000 (Söderström et al. 2000).

*** Lophoziaguttulata (Lindb. et Arnell) A.Evans, Proc. Wash. Acad. Sci. 2: 302, 1900 (Evans 1900c). Bas.: Jungermanniaguttulata Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 51, 1889 (Lindberg and Arnell 1889).

* Lophoziajamaicensis (Nees) Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 128 (155), 1901 (Stephani 1901h). Bas.: Jungermanniajamaicensis Nees, Syn. Hepat. 1: 105, 1844 (Gottsche et al. 1844). 50

** Lophozialacerata N.Kitag., Hikobia 3 (3): 172, 1963 (Kitagawa 1963a).

** Lophozialantratoviae Bakalin, Ann. Bot. Fenn. 40 (1): 47, 2003 (Bakalin 2003).

** Lophoziamurmanica Kaal., Rep. Second Norweg. Arctic Exped. 11: 34, 1906 (Bryhn 1906).

** Lophoziapacifica Bakalin, Bryologist 114 (2): 302, 2011 (Bakalin 2011).

* Lophoziapallida (Steph.) Grolle, J. Jap. Bot. 39 (6): 174, 1964 (Grolle 1964a). Bas.: Anastrophyllumpallidum Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1131 (114), 1901 (Stephani 1901c). 51

** Lophoziasavicziae Schljakov, Novosti Sist. Nizš. Rast. 10: 299, 1973 (Shliakov 1973).

* Lophoziaschusterana Schljakov, Novosti Sist. Nizš. Rast. 12: 320, 1975 (Shliakov 1975).

*** Lophoziasilvicola H.Buch, Beret. 18 Skand. Naturforskarmøde: 228, 1929 (Buch 1929).

*** Lophoziasilvicoloides N.Kitag., J. Hattori Bot. Lab. 28: 276, 1965 (Kitagawa 1965).

* Lophoziasubapiculata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 104, 1974 (Schuster and Damsholt 1974). 52

** Lophoziaudarii S.Srivast., S.C.Srivast. et K.K.Rawat, Nelumbo 55: 130, 2013 (Srivastava et al. 2013). 53

*** Lophoziaventricosa (Dicks.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermanniaventricosa Dicks., Fasc. Pl. Crypt. Brit. 2: 14, 1790 (Dickson 1790). 54

* Lophoziawenzelii (Nees) Steph., Bull. Herb. Boissier (sér. 2) 2 (1): 35 (135), 1902 (Stephani 1902c). Bas.: Jungermanniawenzelii Nees, Naturgesch. Eur. Leberm. 2: 58, 1836 (Nees 1836). 55

Excluded from the genus

* Lophoziaserpens (Dumort.) Dumort., Recueil Observ. Jungerm.: 17, 1835 (Dumortier 1835). Bas.: Jungermanniaserpens Dumort., Syll. Jungerm. Europ.: 56, 1831 (Dumortier 1831). 56

** Lophoziopsis Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009).

*** Lophoziopsisexcisa (Dicks.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermanniaexcisa Dicks., Fasc. Pl. Crypt. Brit. 3: 11, 1793 (Dickson 1793).

** Lophoziopsisexcisavar.elegans (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziaexcisavar.elegans R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 522, 1969 (Schuster 1969b).

** Lophoziopsisexcisavar.infuscata (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziaexcisavar.infuscata R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 94, 1974 (Schuster and Damsholt 1974).

** Lophoziopsisexcisavar.succulenta (R.M.Schust. et Damsh.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziaexcisavar.succulenta R.M.Schust. et Damsh., Meddel. Grønland 199 (1): 96, 1974 (Schuster and Damsholt 1974).

*** Lophoziopsislongidens (Lindb.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermannialongidens Lindb., Helsingf. Dagbl. 1876 (323, 26 Nov.): 2, 1876 (Lindberg 1876a).

** Lophoziopsislongidenssubsp.arctica (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 97 (2): 28, 2013 (Söderström et al. 2013c). Bas.: Lophozialongidenssubsp.arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 539, 1969 (Schuster 1969b). 57

*** Lophoziopsispellucida (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziapellucida R.M.Schust., Canad. J. Bot. 39 (4): 978, 1961 (Schuster 1961b).

** Lophoziopsispellucidavar.minor (R.M.Schust.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 28, 2013 (Söderström et al. 2013c). Bas.: Lophoziapellucidavar.minor R.M.Schust., Canad. J. Bot. 39 (4): 984, 1961 (Schuster 1961b).

*** Lophoziopsispolaris (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziaalpestrissubsp.polaris R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 614, 1969 (Schuster 1969b).

** Lophoziopsispolarisvar.sphagnorum (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziaalpestrisf.sphagnorum R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 619, 1969 (Schuster 1969b).

* Lophoziopsispropagulifera (Gottsche) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Jungermanniapropagulifera Gottsche, Int. Polarforsch., Deutsch. Exped. 2: 451, 1890 (Gottsche 1890). 58

** Lophoziopsisrubrigemma (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 67, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Lophoziarubrigemma R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 621, 1969 (Schuster 1969b).

** Pseudocephaloziella R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a).

*** Pseudocephaloziellaepiphytica R.M.Schust., Phytologia 39 (4): 243, 1978 (Schuster 1978a).

*** Trilophozia (R.M.Schust.) Bakalin, Monogr. Lophozia: 34, 2005 (Bakalin 2005). Bas.: Tritomariasubg.Trilophozia R.M.Schust., Amer. Midl. Naturalist 49 (2): 382, 1953 (Schuster 1953).

*** Trilophoziaquinquedentata (Huds.) Bakalin, Monogr. Lophozia: 34, 2005 (Bakalin 2005). Bas.: Jungermanniaquinquedentata Huds., Fl. Angl. (Hudson): 433, 1762 (Hudson 1762).

** Trilophoziaquinquedentatavar.asymmetrica (Horik.) L.Söderstr. et Váňa, Phytotaxa 97 (2): 32, 2013 (Söderström et al. 2013c). Bas.: Lophoziaasymmetrica Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 153, 1934 (Horikawa 1934).

*** Tritomaria Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (Loeske 1909). Based on: Tritomaria Schiffn., Ber. Naturwiss.-Med. Vereins Innsbruck 31 [Beilage]: 12, 1908 (Schiffner 1908a).

*** Tritomariaexsecta (Schmidel) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (Loeske 1909). Bas.: Jungermanniaexsecta Schmidel, Syst. Samml. Crypt. Gew. 2: 5, 1797 (Schrader 1797), nom. conserv.

** Tritomariaexsectasubsp.novaezelandiae J.J.Engel, Bryologist 109 (1): 61, 2006 (Engel 2006b).

*** Tritomariaexsectiformis (Breidl.) Schiffn. ex Loeske, Hedwigia 49 (1/2): 13, 1909 (Loeske 1909). Bas.: Jungermanniaexsectiformis Breidl., Mitt. Naturwiss. Vereins Steiermark 30: 321, 1894 (Breidler 1894).

** Tritomariaexsectiformissubsp.arctica R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 661, 1969 (Schuster 1969b).

** Tritomariaexsectiformissubsp.camerunensis S.W.Arnell ex Váňa, Phytotaxa 167 (2): 216, 2014 (Váňa et al. 2014b). Based on: Tritomariacamerunensis S.W.Arnell, Svensk Bot. Tidskr. 52 (1): 64, 1958 (Arnell 1958a), nom. inval.

*** Tritomariaferruginea (Grolle) Váňa, Phytotaxa 81 (1): 24, 2013 (Váňa et al. 2013m). Bas.: Andrewsianthusferrugineus Grolle, Khumbu Himal 1 (4): 275, 1966 (Grolle 1966k).

* Tritomariakoreana Bakalin, S.S.Choi et B.Y.Sun, Arctoa 18: 163, 2009 [2010] (Bakalin et al. 2009a).

* Tritomariamexicana Bakalin, Arctoa 17: 162, 2008 [2009] (Bakalin 2008b).

*** Tritomariascitula (Taylor) Jørg., Bergens Mus. Aarbok 1919/20 (7): 3, 1922 (Jørgensen 1922). Bas.: Jungermanniascitula Taylor, London J. Bot. 5: 274, 1846 (Taylor 1846a).

*** Scapaniaceae Mig

by J. Váňa

Scapaniaceae, in its classical concept, was recognized as monophyletic and nested within Lophoziaceae s. lat. (cf. Davis 2004, Schill et al. 2004, Yatsentyuk et al. 2004, He-Nygrén et al. 2006). More recent molecular studies classified Scapaniaceae as a sister clade to Lophoziaceae, and the genus Schistochilopsis, a segregate of the genus Lophozia s. lat., more closely related to Scapaniaceae than to Lophoziaceae (cf. de Roo et al. 2007, Vilnet et al. 2007). The genus Scapania was recently studied by morphological methods by Potemkin (2002), by molecular methods by Vilnet et al. (2007) and especially Heinrichs et al. (2012a). Some taxonomic and nomenclatural notes can be found in Váňa et al. (2012a, 2013j, 2015) and Konstantinova et al. (2013a). The placement of Pseudotritomaria, Saccobasis and Schistochilopsis in the family follows Vilnet et al. (2010).

*** Diplophyllum (Dumort.) Dumort., Recueil Observ. Jungerm.: 15, 1835 (Dumortier 1835) nom. conserv. Bas.: Jungermanniasect.Diplophyllum Dumort., Syll. Jungerm. Europ.: 44, 1831 (Dumortier 1831).

*** subg.Austrodiplophyllum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 18, 1968 (Schuster 1968a).

** Diplophyllumrecurvifolium C.Massal., Atti Reale Ist. Veneto Sci. Lett. Arti 87 (2): 221, 1928 (Massalongo 1928).

*** Diplophyllumsquarrosum Steph., Sp. Hepat. (Stephani) 4: 116, 1910 (Stephani 1910b).

*** Diplophyllumverrucosum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 19, 1968 (Schuster 1968a).

*** subg.Diplophyllum

*** sect.Diplophyllum

*** Diplophyllumalbicans (L.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (Dumortier 1835). Bas.: Jungermanniaalbicans L., Sp. Pl. 1: 1133, 1753 (Linnaeus 1753).

*** sect.Protodiplophyllum (R.M.Schust.) Váňa et L.Söderstr., Phytotaxa 76 (3): 29, 2013 (Váňa et al. 2013j). Bas.: Diplophyllumsubg.Protodiplophyllum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 192, 1974 (Schuster 1974).

*** Diplophyllumafricanum S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 527, 1956 (Arnell 1956e).

*** Diplophyllumandicola R.M.Schust., Phytologia 39 (4): 248, 1978 (Schuster 1978a).

*** Diplophyllumandrewsii A.Evans, Bryologist 25 (2): 28, 1922 (Evans 1922).

* Diplophyllumandrogynum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 277, 1998 (Engel and Smith Merrill 1998).

* Diplophyllumangustifolium J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 262, 1998 (Engel and Smith Merrill 1998).

*** Diplophyllumapiculatum (A.Evans) Steph., Sp. Hepat. (Stephani) 4: 110, 1910 (Stephani 1910b). Bas.: Diplophylleiaapiculata A.Evans, Bot. Gaz. 34 (5): 372, 1902 (Evans 1902b).

** Diplophyllumdioicum R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 20, 1968 (Schuster 1968a).

** Diplophyllumexiguum Steph., Sp. Hepat. (Stephani) 6: 500, 1924 (Stephani 1924).

* Diplophyllumgemmiparum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 255, 1998 (Engel and Smith Merrill 1998).

* Diplophyllumincrassatum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 265, 1998 (Engel and Smith Merrill 1998).

*** Diplophyllumnanum Herzog, Memoranda Soc. Fauna Fl. Fennica 26: 48, 1950 [1951] (Herzog 1950b).

* Diplophyllumnovum J.J.Engel et G.L.Merr., J. Hattori Bot. Lab. 84: 274, 1998 (Engel and Smith Merrill 1998).

*** Diplophyllumobtusatum (R.M.Schust.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 215, 1974 (Schuster 1974). Bas.: Diplophyllumapiculatumvar.obtusatum R.M.Schust., Amer. Midl. Naturalist 49 (2): 432, 1953 (Schuster 1953).

*** Diplophyllumobtusifolium (Hook.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (Dumortier 1835). Bas.: Jungermanniaobtusifolia Hook., Brit. Jungermann.: tab. 26, 1812 (Hooker 1812). 59

** Diplophyllumobtusifoliumsubsp.domesticum (Gottsche) Váňa, Phytotaxa 76 (3): 29, 2013 (Váňa et al. 2013j). Bas.: Jungermanniadomestica Gottsche, Linnaea 28 (5): 548, 1856 [1857] (Gottsche 1856).

*** Diplophyllumserrulatum (Müll.Frib.) Steph., Sp. Hepat. (Stephani) 4: 112, 1910 (Stephani 1910b). Bas.: Diplophylleiaserrulata Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 34, 1903 (Müller 1903).

*** Diplophyllumtaxifolium (Wahlenb.) Dumort., Recueil Observ. Jungerm.: 16, 1835 (Dumortier 1835). Bas.: Jungermanniataxifolia Wahlenb., Fl. Lapp. (Wahlenberg): 389, 1812 (Wahlenberg 1812).

** Diplophyllumtaxifoliumvar.mucronatum R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 203, 1974 (Schuster 1974).

*** Diplophyllumtrollii Grolle, Khumbu Himal 1 (4): 273, 1966 (Grolle 1966k).

*** Douinia (C.E.O.Jensen) H.Buch, Scapan. N.-Eur. Sib.: 13, 1928 (Buch 1928). Bas.: Diplophylleiasubg.Douinia C.E.O.Jensen, Danmarks mosser: 145, 1915 (Jensen 1915).

*** Douiniaimbricata (M.Howe) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (Konstantinova et al. 2013a). Bas.: Scapaniaimbricata M.Howe, Bull. New York Bot. Gard. 2 (6): 104, 1901 (Howe 1901b).

*** Douiniaovata (Dicks.) H.Buch, Scapan. N.-Eur. Sib.: 14, 1928 (Buch 1928). Bas.: Jungermanniaovata Dicks., Fasc. Pl. Crypt. Brit. 3: 11, 1793 (Dickson 1793).

*** Douiniaplicata (Lindb.) Konstant. et Vilnet, Phytotaxa 76 (3): 31, 2013 (Konstantinova et al. 2013a). Bas.: Diplophyllumplicatum Lindb., Acta Soc. Sci. Fenn. 10: 235, 1872 [1873] (Lindberg 1872b).

** Pseudotritomaria Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009).

*** Pseudotritomariaheterophylla (R.M.Schust.) Konstant. et Vilnet, Arctoa 18: 66, 2009 [2010] (Konstantinova and Vilnet 2009). Bas.: Tritomariaheterophylla R.M.Schust., Canad. J. Bot. 36 (2): 272, 1958 (Schuster 1958b).

** Saccobasis H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 291, 1932 [1933] (Buch 1932).

*** Saccobasispolita (Nees) H.Buch, Memoranda Soc. Fauna Fl. Fennica 8: 292, 1932 [1933] (Buch 1932). Bas.: Jungermanniapolita Nees, Naturgesch. Eur. Leberm. 2: 145, 1836 (Nees 1836).

** Saccobasispolymorpha (R.M.Schust.) Schljakov, Novosti Sist. Nizš. Rast. 16: 205, 1979 (Shliakov 1979). Bas.: Tritomariapolitasubsp.polymorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 700, 1969 (Schuster 1969b).

*** Scapania (Dumort.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835) nom. conserv. Bas.: Radulasect.Scapania Dumort., Syll. Jungerm. Europ.: 38, 1831 (Dumortier 1831).

** subg.Ascapania Grolle, Khumbu Himal 1 (4): 268, 1966 (Grolle 1966k).

*** Scapaniacontorta Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (Mitten 1860c).

** subg.Gracilidae (H.Buch) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (Váňa et al. 2012a). Bas.: Scapaniasect.Gracilidae H.Buch, Scapan. N.-Eur. Sib.: 106, 1928 (Buch 1928).

*** Scapaniaampliata Steph., Bull. Herb. Boissier 5 (2): 106, 1897 (Stephani 1897b).

** Scapaniaampliatasubsp.queenslandica M.L.Hicks, J. Hattori Bot. Lab. 69: 130, 1991 (Hicks 1991).

*** Scapaniabolanderi Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 218, 1869 (Austin 1869).

*** Scapaniagracilis Lindb., Morgonbladet (Helsinki) 1873 (286, 9 Dec): 2, 1873 (Lindberg 1873a).

** Scapaniamacroparaphyllia T.Cao, C.Gao et J.Sun, Acta Phytotax. Sin. 42 (2): 180, 2004 (Cao et al. 2004).

*** Scapaniamaxima Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 223, 1934 (Horikawa 1934).

*** Scapanianipponica (Amakawa et S.Hatt.) Amakawa, J. Hattori Bot. Lab. 30: 319, 1967 (Amakawa 1967a). Bas.: Scapaniabolanderivar.nipponica Amakawa et S.Hatt., J. Hattori Bot. Lab. 14: 83, 1955 (Amakawa and Hattori 1955).

** Scapaniaparaphyllia T.Cao, C.Gao, J.Sun et B.R.Zuo, Acta Phytotax. Sin. 45 (3): 311, 2007 (Zuo et al. 2007b).

*** Scapaniasubnimbosa Steph., Sp. Hepat. (Stephani) 4: 150, 1910 (Stephani 1910b).

** subg.Macroscapania R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 248, 1974 (Schuster 1974).

* Scapaniageppii Steph., Hedwigia 44 (1): 14, 1904 (Stephani 1904i).

*** Scapaniaportoricensis Hampe et Gottsche, Linnaea 25 (3): 342, 1852 [1853] (Hampe and Gottsche 1852).

** Scapaniaportoricensisvar.boliviensis (Steph.) Herzog, Ann. Bryol. 1: 110, 1928 (Herzog 1928). Bas.: Scapaniaboliviensis Steph., Biblioth. Bot. 87 (2): 231, 1916 (Stephani 1916a).

** Scapaniaportoricensisvar.organensis (Herzog) Herzog, Ann. Bryol. 1: 110, 1928 (Herzog 1928). Bas.: Scapaniaorganensis Herzog, Repert. Spec. Nov. Regni Veg. 21 (1/7): 27, 1925 (Herzog 1925a).

** Scapaniaportoricensisvar.roraimensis Warnst., Hedwigia 63 (2): 109, 1921 (Warnstorf 1921).

** subg.Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903 (Müller 1903).

** sect.Planifoliae (Müll.Frib.) Potemkin, J. Hattori Bot. Lab. 85: 56, 1998 (Potemkin 1998). Bas.: Scapania Gruppe Planifoliae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 286, 1905 (Müller 1905).

*** Scapaniadavidii Potemkin, Ann. Bot. Fenn. 38 (2): 83, 2001 (Potemkin 2001).

** Scapaniaferruginaeoides T.Cao, C.Gao et J.Sun, Guihaia 24 (1): 23, 2004 (Sun et al. 2004).

** Scapaniagaochii X.Fu ex T.Cao, Phytotaxa 97 (2): 26, 2013 (Cao et al. 2013). Based on: Scapaniagaochii X.Fu ex T.Cao, Acta Bot. Yunnan. 25 (5): 541, 2003 (Cao et al. 2003), nom. inval.

*** Scapaniaharae Amakawa, J. Hattori Bot. Lab. 27: 5, 1964 (Amakawa 1964a).

*** Scapanianimbosa Taylor, Nov. Stirp. Pug. 8: 6, 1844 (Lehmann 1844).

*** Scapaniaornithopodioides (With.) Waddell, Moss Exch. Club Cat. Hepat.: 4, 1897 (Waddell 1897). Bas.: Jungermanniaornithopodioides With., Bot. arr. veg. Gr. Brit. 2: 695, 1776 (Withering 1776).

*** Scapaniarotundifolia W.E.Nicholson, Symb. Sin. 5: 31, 1930 (Nicholson et al. 1930).

*** Scapaniasecunda Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 226, 1894 (Stephani 1894b).

*** Scapaniazhukovae Potemkin, Arctoa 9: 121, 2000 (Potemkin 2000b).

** sect.Plicaticalyx (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 326, 2002 (Potemkin 2002). Bas.: Scapaniasubg.Plicaticalyx Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 36, 1903 (Müller 1903).

*** Scapaniaciliatospinosa Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 222, 1934 (Horikawa 1934).

*** Scapaniaferruginea (Lehm. et Lindenb.) Lehm. et Lindenb., Syn. Hepat. 1: 72, 1844 (Gottsche et al. 1844). Bas.: Jungermanniaferruginea Lehm. et Lindenb., Nov. Stirp. Pug. 4: 20, 1832 (Lehmann 1832).

*** Scapaniahians Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 223, 1905 (Müller 1905).

** Scapaniahianssubsp.salishensis J.D.Godfrey et G.Godfrey, Bryologist 81 (3): 362, 1978 (Godfrey and Godfrey 1978).

*** Scapaniaorientalis Steph. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 606, 1901 (Müller 1901a).

*** Scapaniapseudocontorta Potemkin, Arctoa 9: 115, 2000 (Potemkin 2000b).

*** Scapaniasinikkae Potemkin, Ann. Bot. Fenn. 38 (2): 85, 2001 (Potemkin 2001).

*** Scapaniaspiniloba Potemkin, Arctoa 9: 117, 2000 (Potemkin 2000b).

** subg.Pseudomacrodiplophyllum Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (Váňa et al. 2012a).

*** Scapaniamicrodonta (Mitt.) Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 262, 1905 (Müller 1905). Bas.: Martinelliusmicrodontus Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 196, 1891 (Mitten 1891).

*** subg.Scapania

** sect.Aequilobae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 110, 1928 (Buch 1928). Bas.: Scapania Gruppe Aequilobae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 219, 1905 (Müller 1905).

*** Scapaniaaequiloba (Schwägr.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermanniaaequiloba Schwägr., Hist. Musc. Hepat. Prodr.: 24, 1814 (Schwägrichen 1814).

*** Scapaniaaspera M.Bernet et Bernet, Cat. hép. Suisse: 42, 1888 (Bernet 1888).

** sect.Americanae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 15, 2012 (Váňa et al. 2012a).

*** Scapaniaamericana Müll.Frib., Bull. Herb. Boissier (sér. 2) 3 (1): 44, 1903 (Müller 1903).

** sect.Apiculatae H.Buch, Scapan. N.-Eur. Sib.: 53, 1928 (Buch 1928).

*** Scapaniaapiculata Spruce, Hep. Pyr. Exsic.: no. 15, 1847 (Spruce 1847).

*** Scapaniacarinthiaca J.B.Jack ex Lindb., Rev. Bryol. 7 (4): 77, 1880 (Lindberg 1880a).

*** Scapaniacarinthiacavar.massalongi Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 598, 1901 (Müller 1901a).

*** Scapaniaumbrosa (Schrad.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermanniaumbrosa Schrad., Syst. Samml. Crypt. Gew. 2: 5, 1797 (Schrader 1797).

** sect.Ciliatae Grolle, Khumbu Himal 1 (4): 272, 1966 (Grolle 1966k).

*** Scapaniabhutanensis Amakawa, Fl. E. Himalaya 2: 230, 1971 (Hattori 1971a).

*** Scapaniaciliata Sande Lac., Prolus. fl. jap.: 209, 1867 (Sande Lacoste 1867).

*** Scapaniaciliatasubsp.hawaiica (Müll.Frib.) Potemkin, Ann. Bot. Fenn. 39 (4): 321, 2002 (Potemkin 2002). Bas.: Scapaniahawaiica Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 160, 1905 (Müller 1905).

*** Scapaniahirosakiensis Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 120, 1905 (Müller 1905).

*** Scapaniahollandiae W.S.Hong, Bryologist 83 (1): 56, 1980 (Hong 1980).

*** Scapaniakoponenii Potemkin, Ann. Bot. Fenn. 37 (1): 41, 2000 (Potemkin 2000c).

*** Scapanialepida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 101, 1860 [1861] (Mitten 1860c).

*** Scapaniasandei Schiffn. ex Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 612, 1901 (Müller 1901a).

** sect.Compactae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 101, 1928 (Buch 1928). Bas.: Scapania Gruppe Compactae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 53, 1905 (Müller 1905).

*** Scapaniacompacta (Roth) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermanniacompacta Roth, Tent. Fl. Germ. 3: 375, 1800 (Roth 1800).

*** Scapaniakaurinii Ryan, Bot. Not. 42: 210, 1889 (Ryan 1889).

*** Scapaniaspitsbergensis (Lindb.) Müll.Frib., Bull. Herb. Boissier (sér. 2) 1 (6): 607, 1901 (Müller 1901a). Bas.: Martinelliusspitsbergensis Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 31, 1889 (Lindberg and Arnell 1889).

** sect.Curtae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 55, 1928 (Buch 1928). Bas.: Scapania Gruppe Curtae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 245, 1905 (Müller 1905).

*** Scapaniacurta (Mart.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermanniacurta Mart., Fl. crypt. erlang.: 148, 1817 (Martius 1817).

** Scapaniacurtavar.grandiretis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 393, 1974 (Schuster 1974).

** Scapaniacurtavar.isoloba R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 396, 1974 (Schuster 1974).

*** Scapaniadiplophylloides Amakawa et S.Hatt., J. Hattori Bot. Lab. 9: 59, 1953 (Amakawa and Hattori 1953).

*** Scapaniaesterhuyseniae S.W.Arnell, Bot. Not. 110 (1): 26, 1957 (Arnell 1957a).

*** Scapaniafulfordiae W.S.Hong, Bryologist 83 (1): 46, 1980 (Hong 1980).

*** Scapaniagamundiae R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 14, 1968 (Schuster 1968a).

*** Scapaniahelvetica Gottsche, Hepat. Eur., Leberm. 42-44: no. 426, 1868 (Gottsche and Rabenhorst 1868).

*** Scapaniairrigua (Nees) Nees, Syn. Hepat. 1: 67, 1844 (Gottsche et al. 1844). Bas.: Jungermanniairrigua Nees, Naturgesch. Eur. Leberm. 1: 193, 1833 (Nees 1833c).

** Scapaniairriguasubsp.rufescens (Loeske) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 471, 1974 (Schuster 1974). Bas.: Scapaniairriguaf.rufescens Loeske, Moosfl. Harz.: 71, 1903 (Loeske 1903).

*** Scapanialingulata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 92, 1916 (Buch 1916).

** Scapanialingulatavar.microphylla (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 415, 1974 (Schuster 1974). Bas.: Scapaniamicrophylla Warnst., Hedwigia 63 (2): 75, 1921 (Warnstorf 1921).

** Scapaniamagadanica S.S.Choi, Bakalin et B.Y.Sun, Bot. Pacifica 1: 46, 2012 (Choi et al. 2012).

*** Scapaniamucronata H.Buch, Meddel. Soc. Fauna Fl. Fenn. 42: 91, 1916 (Buch 1916).

*** Scapaniaobcordata (Berggr.) S.W.Arnell, Ark. Bot. (n.ser.) 4 (6): 117, 1959 (Arnell and Mårtensson 1959). Bas.: Sarcocyphosobcordatus Berggr., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 13 (7): 96, 1875 (Berggren 1875).

** Scapaniaparvifolia Warnst., Hedwigia 63 (2): 78, 1921 (Warnstorf 1921).

*** Scapaniapraetervisa Meyl., Jahresber. Naturf. Ges. Graubündens (n.f.) 64: 364, 1926 (Meylan 1926).

*** Scapaniascandica (Arnell et H.Buch) Macvicar, Stud. handb. Brit. hepat. (ed. 2): 394, 1926 (Macvicar 1926). Bas.: Martinelliusscandicus Arnell et H.Buch, Bot. Not. 74: 1, 1921 (Arnell and Buch 1921).

** Scapaniascandicavar.argutedentata H.Buch, Scapan. N.-Eur. Sib.: 75, 1928 (Buch 1928).

** Scapaniascandicavar.dimorpha R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 453, 1974 (Schuster 1974).

** Scapaniascandicavar.grandiretis (Schljakov) Schljakov, Pečen. Mchi Sev. SSSR 4: 152, 1981 (Shliakov 1981). Bas.: Scapaniaparvifoliavar.grandiretis Schljakov, Novosti Sist. Nizš. Rast. 8: 332, 1971 (Shliakov 1971).

*** Scapaniauliginosa (Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermanniaundulatavar.uliginosa Lindenb., Syn. hepat. eur: 58, 1829 (Lindenberg 1829).

*** Scapaniavaldonii Váňa, Bedn.-Ochyra et Cykowska, Nova Hedwigia 89 (1/2): 126, 2009 (Váňa et al. 2009).

*** Scapaniazemliae S.W.Arnell, Svensk Bot. Tidskr. 41: 215, 1947 (Arnell 1947).

** sect.Cuspiduligerae H.Buch, Scapan. N.-Eur. Sib.: 125, 1928 (Buch 1928).

*** Scapaniacuspiduligera (Nees) Müll.Frib., Lebermoose 2 (22): 472, 1915 (Müller 1915a). Bas.: Jungermanniacuspiduligera Nees, Naturgesch. Eur. Leberm. 1: 180, 1833 (Nees 1833c).

** Scapaniacuspiduligeravar.diplophyllopsis R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 361, 1974 (Schuster 1974).

** sect.Grolleoscapania Potemkin, Ann. Bot. Fenn. 39 (4): 329, 2002 (Potemkin 2002).

*** Scapaniakarl-muelleri Grolle, Khumbu Himal 1 (4): 270, 1966 (Grolle 1966k).

** sect.Hyperboreae Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (Váňa et al. 2012a).

*** Scapaniahyperborea Jørg., Forh. Vidensk.-Selsk. Kristiania 1894 (8): 56, 1894 (Jørgensen 1894).

*** Scapaniapaludicola Loeske et Müll.Frib., Lebermoose 2 (21): 425, 1915 (Müller 1915b).

** Scapaniapaludicolavar.viridigemma R.M.Schust., Bull. Natl. Mus. Canada 122: 20, 1950 [1951] (Schuster 1950).

*** Scapaniatundrae (Arnell) H.Buch, Scapan. N.-Eur. Sib.: 99, 1928 (Buch 1928). Bas.: Martinelliustundrae Arnell, Bot. Not. 74: 289, 1921 (Arnell 1921).

** sect.Kaalaasia (H.Buch) Jørg., Bergens Mus. Skr. (n.ser.) 16: 210, 1934 (Jørgensen 1934). Bas.: Scapaniasubg.Kaalaasia H.Buch, Scapan. N.-Eur. Sib.: 47, 1928 (Buch 1928).

*** Scapaniacalcicola (Arnell et J.Perss.) Ingham, Naturalist (Hull) 564: 11, 1904 (Ingham 1904). Bas.: Martinelliuscalcicola Arnell et J.Perss., Rev. Bryol. 30 (6): 97, 1903 (Arnell 1903).

*** Scapaniagymnostomophila Kaal., Bot. Not. 49: 21, 1896 (Kaalaas 1896).

*** Scapanialigulifolia R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 306, 1974 (Schuster 1974).

*** Scapaniapseudocalcicola R.M.Schust., Phytologia 63 (5): 327, 1987 (Schuster and Damsholt 1987).

* Scapaniasect.Muelleria Potemkin, Ann. Bot. Fenn. 39 (4): 320, 2002 (Potemkin 2002).

*** Scapaniahimalayica Müll.Frib. ex Herzog, Ann. Bryol. 12: 81, 1939 (Herzog 1939b).

*** Scapaniaschljakovii Potemkin, Ann. Bot. Fenn. 38 (2): 87, 2001 (Potemkin 2001).

** sect.Nemorosae (Müll.Frib.) H.Buch, Scapan. N.-Eur. Sib.: 152, 1928 (Buch 1928). Bas.: Scapania Gruppe Nemorosae Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 155, 1905 (Müller 1905).

*** Scapaniabrevicaulis Taylor, London J. Bot. 5: 272, 1846 (Taylor 1846a).

*** Scapaniacrassiretis Bryhn, Rev. Bryol. 19 (1): 7, 1892 (Bryhn 1892).

* Scapaniadegenii Schiffn. ex Müll.Frib., Lebermoose 2 (22): 497, 1915 (Müller 1915a). 60

* Scapaniaglaucoviridis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 221, 1934 (Horikawa 1934). 61

** Scapaniagrossidens Steph. ex Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 146, 1905 (Müller 1905).

*** Scapaniahedbergii S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 556, 1956 (Arnell 1956e).

*** Scapaniaintegerrima Steph., Sp. Hepat. (Stephani) 4: 148, 1910 (Stephani 1910b).

*** Scapaniamatveyevae Potemkin, Arctoa 9: 97, 2000 (Potemkin 2000a).

*** Scapanianemorea (L.) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 160, 1963 [1964] (Grolle 1963d). Bas.: Jungermannianemorea L., Syst. Nat., ed. 10., 2: 1337, 1759 (Linnaeus 1759).

** Scapaniaparvidens Steph., Hedwigia 44 (1): 15, 1904 (Stephani 1904i). 62

*** Scapaniaparvitexta Steph., Bull. Herb. Boissier 5 (2): 107, 1897 (Stephani 1897b).

*** Scapaniarigida Nees, Syn. Hepat. 1: 69, 1844 (Gottsche et al. 1844).

** sect.Scapania

*** Scapaniagigantea Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 1: 15, 1931 (Horikawa 1931b).

** Scapaniagrandiloba Steph., Sp. Hepat. (Stephani) 6: 502, 1924 (Stephani 1924).

*** Scapaniakomagadakensis Amakawa, J. Hattori Bot. Lab. 31: 96, 1968 (Amakawa 1968a).

*** Scapaniaobscura (Arnell et C.E.O.Jensen) Schiffn., Österr. Bot. Z. 58 (10): 377, 1908 (Schiffner 1908b). Bas.: Martinelliusobscurus Arnell et C.E.O.Jensen, Moose Sarekgeb.: 91, 1907 (Arnell and Jensen 1907).

** Scapaniapaludosa (Müll.Frib.) Müll.Frib., Mitt. Bad. Bot. Vereins 4 (182/183): 287, 1902 (Müller 1902). Bas.: Scapaniaundulatavar.paludosa Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 220, 1901 (Müller 1901b).

*** Scapaniarufidula Warnst., Hedwigia 63 (2): 94, 1921 (Warnstorf 1921).

*** Scapaniaserrulata R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 539, 1974 (Schuster 1974).

*** Scapaniasubalpina (Nees ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermanniasubalpina Nees ex Lindenb., Syn. hepat. eur: 55, 1829 (Lindenberg 1829).

** Scapaniasubalpinavar.haynesiae Frye et L.Clark, Univ. Wash. Publ. Biol. 6 (4): 638, 1946 (Frye and Clark 1946).

** Scapaniasubalpinavar.muddiae C.D.Bird et W.S.Hong, Bryologist 83 (1): 51, 1980 (Hong 1980).

*** Scapaniaundulata (L.) Dumort., Recueil Observ. Jungerm.: 14, 1835 (Dumortier 1835). Bas.: Jungermanniaundulata L., Sp. Pl. 1: 1132, 1753 (Linnaeus 1753).

** sect.Scapaniella (H.Buch) Potemkin, J. Hattori Bot. Lab. 85: 43, 1998 (Potemkin 1998). Bas.: Scapaniella H.Buch, Scapan. N.-Eur. Sib.: 33, 1928 (Buch 1928).

*** Scapaniaglaucocephala (Taylor) Austin, Bull. Torrey Bot. Club 6 (16): 85, 1876 (Austin 1876c). Bas.: Jungermanniaglaucocephala Taylor, London J. Bot. 5: 277, 1846 (Taylor 1846a).

*** Scapaniaglaucocephalavar.saxicola (R.M.Schust.) Potemkin, Bryologist 102 (1): 36, 1999 (Potemkin 1999). Bas.: Scapaniasaxicola R.M.Schust., Amer. Midl. Naturalist 49 (2): 448, 1953 (Schuster 1953).

* Scapaniascapanioides (C.Massal.) Grolle, Feddes Repert. 87 (3/4): 235, 1976 (Grolle 1976a). Bas.: Jungermanniascapanioides C.Massal., Hepaticol. ven.: 64, 1879 (Massalongo 1879). 63

** sect.Simmonsiae (R.M.Schust.) Váňa, Hentschel, Joch.Müll. et Heinrichs, PhytoKeys 10: 16, 2012 (Váňa et al. 2012a). Bas.: Scapaniasubsect.Simmonsiae R.M.Schust., Hepat. Anthocerotae N. Amer. 3: 612, 1974 (Schuster 1974).

*** Scapaniasimmonsii Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 51, 1906 (Bryhn 1906).

** sect.Sphaeriferae Konstant. et Potemkin, Ann. Bot. Fenn. 31 (2): 125, 1994 (Konstantinova and Potemkin 1994).

*** Scapaniasphaerifera H.Buch et Tuom., Memoranda Soc. Fauna Fl. Fennica 11: 227, 1936 (Buch and Tuomikoski 1936).

** sect.Stephaniae Potemkin, J. Hattori Bot. Lab. 85: 57, 1998 (Potemkin 1998). Based on: Scapaniasect.Stephaniae Amakawa et S.Hatt., J. Hattori Bot. Lab. 12: 94, 1954 (Amakawa and Hattori 1954). nom. inval.

*** Scapaniagriffithii Schiffn., Österr. Bot. Z. 49 (6): 204, 1899 (Schiffner 1899a).

*** Scapaniajavanica Gottsche, Natuurk. Tijdschr. Ned.-Indië 4: 575, 1853 (Gottsche 1853).

* Scapaniajavanicavar.scabra Schiffn., Arch. Hydrobiol., suppl. 21 (3/4): 397, 1955 (Schiffner 1955).

*** Scapanialigulata Steph., Hedwigia 44 (1): 14, 1904 (Stephani 1904i).

** Scapanialigulatasubsp.stephanii (Müll.Frib.) Potemkin, Piippo et T.J.Kop., Ann. Bot. Fenn. 41 (6): 423, 2004 (Potemkin et al. 2004). Bas.: Scapaniastephanii Müll.Frib., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 83: 273, 1905 (Müller 1905).

** sect.Verrucosae Potemkin, J. Hattori Bot. Lab. 85: 54, 1998 (Potemkin 1998).

*** Scapaniaudarii S.C.Srivast. et A.Srivast., J. Indian Bot. Soc. 72: 237, 1993 (Srivastava and Srivastava 1993).

*** Scapaniaverrucosa Heeg, Rev. Bryol. 20 (5): 81, 1893 (Heeg 1893).

*** Schistochilopsis (N.Kitag.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Lophoziasubg.Schistochilopsis N.Kitag., J. Hattori Bot. Lab. 28: 289, 1965 (Kitagawa 1965).

*** Schistochilopsiscornuta (Steph.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Schistochilacornuta Steph., Sp. Hepat. (Stephani) 4: 84, 1909 (Stephani 1909d).

*** Schistochilopsisgrandiretis (Lindb. ex Kaal.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Jungermanniagrandiretis Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 322, 1893 (Kaalaas 1893b).

** Schistochilopsishyperarctica Konstant. et L.Söderstr., Phytotaxa 162 (4): 240, 2014 (Konstantinova et al. 2014b). Based on: Lophoziahyperarctica R.M.Schust., Canad. J. Bot. 39 (4): 967, 1961 (Schuster 1961b), nom. inval.

*** Schistochilopsisincisa (Schrad.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Jungermanniaincisa Schrad., Syst. Samml. Crypt. Gew. 2: 5, 1797 (Schrader 1797).

* Schistochilopsisnakanishii (Inoue) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Lophozianakanishii Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (1): 37, 1966 (Inoue 1966a). 64

** Schistochilopsisopacifolia (Culm. ex Meyl.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Lophoziaopacifolia Culm. ex Meyl., Beitr. Kryptogamenfl. Schweiz 6 (4): 174, 1924 (Meylan 1924). 65

*** Schistochilopsissetosa (Mitt.) Konstant., Arctoa 3: 125, 1994 (Konstantinova and Vasil’ev 1994). Bas.: Jungermanniasetosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 92, 1860 [1861] (Mitten 1860c).

Jungermanniineae R.M.Schust. ex Stotler et Crand.-Stotl
*** Acrobolbaceae E.A.Hodgs

by L. Briscoe and J.J. Engel

** Enigmella G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (Beckmann and Scott 1992).

** Enigmellathallina G.A.M.Scott et K.G.Beckm., J. Bryol. 17 (2): 297, 1992 (Beckmann and Scott 1992).

** Acrobolboideae R.M.Schust. ex Briscoe

*** Acrobolbus Nees, Syn. Hepat. 1: 5, 1844 (Gottsche et al. 1844).

** Acrobolbusafricanus (Pearson) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusafricanus Pearson, Forh. Vidensk.-Selsk. Kristiania 1887 (9): 14, 1887 (Pearson 1887b).

*** Acrobolbusanisodontus (Hook.f. et Taylor) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Jungermanniaanisodonta Hook.f. et Taylor, London J. Bot. 4: 79, 1845 (Hooker and Taylor 1845).

** Acrobolbusantillanus R.M.Schust., J. Hattori Bot. Lab. 90: 143, 2001 (Schuster 2001a).

** Acrobolbusazoricus (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusazoricus Grolle et Perss., Svensk Bot. Tidskr. 60 (1): 169, 1966 (Grolle and Persson 1966).

*** Acrobolbuscaducifolius R.M.Schust., J. Hattori Bot. Lab. 90: 154, 2001 (Schuster 2001a).

*** Acrobolbusciliatus (Mitt.) Schiffn., Hepat. (Engl.-Prantl): 86, 1893 (Schiffner 1893b). Bas.: Gymnantheciliata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 100, 1860 [1861] (Mitten 1860c).

*** Acrobolbuscinerascens (Lehm. et Lindenb.) Bastow, Pap. & Proc. Roy. Soc. Tasmania 1887: 242, 1888 (Bastow 1888). Bas.: Jungermanniacinerascens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 46, 1832 (Lehmann 1832).

*** Acrobolbusconcinnus (Mitt.) Steph., Trans. & Proc. New Zealand Inst. 24: 399, 1892 (Colenso 1892). Bas.: Gymnantheconcinna Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 230, 1860 (Mitten 1860b).

** Acrobolbuscuneifolius (Steph.) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthuscuneifolius Steph., Bull. Herb. Boissier (sér. 2) 5 (12): 1138 (10), 1905 (Stephani 1905b).

*** Acrobolbusdiversifolius R.M.Schust., J. Hattori Bot. Lab. 90: 150, 2001 (Schuster 2001a).

*** Acrobolbusepiphytus (Colenso) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidiumepiphytum Colenso, Trans. & Proc. New Zealand Inst. 21: 64, 1889 (Colenso 1889).

*** Acrobolbusflavicans (J.J.Engel et Grolle) Briscoe et J.J.Engel, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidiumflavicans J.J.Engel et Grolle, J. Hattori Bot. Lab. 34: 438, 1971 (Engel and Grolle 1971).

*** Acrobolbusgradsteinii (Grolle) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidiumgradsteinii Grolle, J. Hattori Bot. Lab. 66: 337, 1989 (Grolle 1989a).

** Acrobolbusintegrifolius (A.Evans) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusintegrifolius A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (Evans 1891).

*** Acrobolbusknightii (Mitt.) Briscoe, Phytotaxa 202 (1): 59, 2015 (Briscoe et al. 2015). Bas.: Marsupidiumknightii Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (Hooker 1867).

** Acrobolbuskunkelii (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthuskunkelii Hässel et Solari, Darwiniana 17: 574, 1972 (Hässel and Solari 1972).

*** Acrobolbuslaxus (Lehm. et Lindenb.) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Plagiochilalaxa Lehm. et Lindenb., Sp. Hepat. (Lindenberg) 2-4: 68, 1840 (Lindenberg 1840).

*** Acrobolbuslimbatus (Steph.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthuslimbatus Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 25, 1911 (Stephani 1911b).

*** Acrobolbuslophocoleoides (Mitt.) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (Hooker 1867). Bas.: Gymnanthelophocoleoides Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 144, 1854 (Mitten 1854).

** Acrobolbusmadeirensis (Grolle et Perss.) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusmadeirensis Grolle et Perss., Svensk Bot. Tidskr. 60 (1): 166, 1966 (Grolle and Persson 1966).

** Acrobolbusmittenii Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 460 (179), 1902 (Stephani 1902f).

*** Acrobolbusochrophyllus (Hook.f. et Taylor) R.M.Schust., Rev. Bryol. Lichénol. 30 (1/2): 64, 1961 (Schuster 1961a). Bas.: Jungermanniaochrophylla Hook.f. et Taylor, London J. Bot. 3: 368, 1844 (Hooker and Taylor 1844a).

*** Acrobolbuspapillosus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Marsupidiumpapillosum J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 289, 2008 (Engel and Glenny 2008c).

*** Acrobolbusperpusillus (Colenso) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusperpusillus Colenso, Trans. & Proc. New Zealand Inst. 19: 286, 1887 (Colenso 1887).

*** Acrobolbusperpusillusvar.denticulatus (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Marsupidiumperpusillumvar.denticulatum J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 284, 2008 (Engel and Glenny 2008c).

*** Acrobolbusplagiochiloides (J.J.Engel et Glenny) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Marsupidiumplagiochiloides J.J.Engel et Glenny, Nova Hedwigia 87 (3/4): 284, 2008 (Engel and Glenny 2008c).

*** Acrobolbuspseudosaccatus (Grolle) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthuspseudosaccatus Grolle, Nova Hedwigia 6 (3/4): 391, 1963 (Grolle 1963c).

*** Acrobolbusrenifolius (Hässel et Solari) Briscoe et J.J.Engel, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusrenifolius Hässel et Solari, Darwiniana 17: 583, 1972 (Hässel and Solari 1972).

** Acrobolbusruwenzorensis (S.W.Arnell) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusruwenzorensis S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 560, 1956 (Arnell 1956e).

*** Acrobolbussaccatus (Hook.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Jungermanniasaccata Hook., Musci Exot. 1: tab. 16, 1818 (Hooker 1818).

*** Acrobolbussetulosus (Mitt.) Briscoe, Phytotaxa 202 (1): 60, 2015 (Briscoe et al. 2015). Bas.: Gymnanthesetulosa Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 144, 1854 (Mitten 1854).

*** Acrobolbusspinifolius R.M.Schust., J. Hattori Bot. Lab. 90: 137, 2001 (Schuster 2001a).

*** Acrobolbussumatranus (Schiffn.) Briscoe, Phytotaxa 202 (1): 61, 2015 (Briscoe et al. 2015). Bas.: Lophoziasumatrana Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 203, 1898 (Schiffner 1898a).

*** Acrobolbussurculosus (Nees) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Scapaniasurculosa Nees, Syn. Hepat. 1: 62, 1844 (Gottsche et al. 1844).

*** Acrobolbustenellus (Taylor) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Gymnanthetenella Taylor, Nov. Stirp. Pug. 8: 1, 1844 (Lehmann 1844).

*** Acrobolbustenellusvar.diversifolius (E.A.Hodgs.) Briscoe, Phytotaxa 202 (1): 61, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusdiversifolius E.A.Hodgs., Trans. Roy. Soc. New Zealand 85 (4): 575, 1958 (Hodgson 1958).

*** Acrobolbusurvilleanus (Mont.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 423, 1877 (Trevisan 1877). Bas.: Plagiochilaurvilleana Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843 (Montagne 1843).

*** Acrobolbusviridis (Mitt.) Briscoe et J.J.Engel, Phytotaxa 202 (1): 61, 2015 (Briscoe et al. 2015). Bas.: Tylimanthusviridis Mitt., J. Linn. Soc., Bot. 15 (84): 197, 1876 (Mitten 1876b).

*** Acrobolbuswilsonii Nees, Syn. Hepat. 1: 5, 1844 (Gottsche et al. 1844).

** Acrobolbuswilsoniivar.andinus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 522, 1885 (Spruce 1885).

Excluded from the genus

* Acrobolbusbispinosus (J.B.Jack et Steph.) Steph., Bull. Herb. Boissier (sér. 2) 2 (5): 459 (178), 1902 (Stephani 1902f). Bas.: Tylimanthusbispinosus J.B.Jack et Steph., Hedwigia 31 (1): 26, 1892 (Jack and Stephani 1892). 66

** Austrolophozioideae R.M.Schust. ex Crand.-Stotl., Váňa et Stotler

** Austrolophozia R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (Schuster 1963b).

** Austrolophoziaandina R.M.Schust., Nova Hedwigia 15: 495, 1968 (Schuster 1968b).

*** Austrolophoziacamensis (Steph.) Grolle ex Hässel et Solari, Revista Mus. Argent. Ci. Nat., Bernardino Rivadavia Inst. Nac. Invest. Ci. Nat. Bot. 3 (6): 240, 1970 (Hässel and Solari 1970). Bas.: Tylimanthuscamensis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 24, 1911 (Stephani 1911b).

*** Austrolophoziaparadoxa R.M.Schust., J. Hattori Bot. Lab. 26: 282, 1963 (Schuster 1963b).

*** Goebelobryum Grolle, J. Hattori Bot. Lab. 25: 135, 1962 (Grolle 1962b).

*** Goebelobryumgrossitextum (Steph.) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (Grolle 1962b). Bas.: Marsupidiumgrossitextum Steph., Sp. Hepat. (Stephani) 6: 446, 1924 (Stephani 1924).

*** Goebelobryumunguiculatum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 25: 137, 1962 (Grolle 1962b). Bas.: Jungermanniaunguiculata Hook.f. et Taylor, London J. Bot. 5: 279, 1846 (Taylor 1846a).

*** Goebelobryumvermiculare J.J.Engel et Glenny, Nova Hedwigia 95 (3/4): 320, 2012 (Engel and Glenny 2012).

** Lethocoleoideae Grolle

*** Lethocolea Mitt., Handb. N. Zeal. fl. 2: 751, 1867 (Hooker 1867) nom. conserv.

** Lethocoleacongesta (Lehm.) S.W.Arnell, Bot. Not. 108: 311, 1955 (Arnell 1955b). Bas.: Jungermanniacongesta Lehm., Linnaea 4: 365, 1829 (Lehmann 1829).

*** Lethocoleaglossophylla (Spruce) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (Grolle 1965c). Bas.: Symphyomitraglossophylla Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 503, 1885 (Spruce 1885).

** Lethocoleaindica G.Asthana et Maurya, Natl. Acad. Sci. Lett. 37 (6): 535, 2014 (Asthana and Maurya 2014).

*** Lethocoleajavanica (Schiffn.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (Grolle 1965c). Bas.: Symphyomitrajavanica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 193, 1898 (Schiffner 1898a).

** Lethocoleanaruto-toganensis Furuki, Bryologist 104 (2): 306, 2001 (Furuki 2001).

*** Lethocoleapansa (Taylor) G.A.M.Scott et K.G.Beckm., Symp. Biol. Hung. 35: 212, 1987 (Scott and Beckmann 1987). Bas.: Jungermanniapansa Taylor, London J. Bot. 5: 275, 1846 (Taylor 1846a).

*** Lethocolearadicosa (Lehm. et Lindenb.) Grolle, Bot. Mag. (Tokyo) 78 (921): 83, 1965 (Grolle 1965c). Bas.: Jungermanniaradicosa Lehm. et Lindenb., Nov. Stirp. Pug. 6: 35, 1834 (Lehmann 1834).

* Lethocolearepens S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 67, 1969 (Winkler 1969).

** Saccogynidioideae Crand.-Stotl., Váňa et Stotler

*** Saccogynidium Grolle, J. Hattori Bot. Lab. 23: 43, 1960 [1961] (Grolle 1960d).

** sect.Decurvum Grolle, J. Hattori Bot. Lab. 23: 59, 1960 (Grolle 1960d).

*** Saccogynidiumdecurvum (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 59, 1960 [1961] (Grolle 1960d). Bas.: Lophocoleadecurva Mitt., Bot. antarct. voy. III (Fl. Tasman. 2): 227, 1860 (Mitten 1860b).

** sect.Jugata Grolle, J. Hattori Bot. Lab. 23: 55, 1960 (Grolle 1960d).

** Saccogynidiumrigidulum (Nees) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (Grolle 1960d). Bas.: Jungermanniarigidula Nees, Enum. Pl. Crypt. Javae: 25, 1830 (Nees 1830).

** sect.Saccogynidium

*** Saccogynidiumaustrale (Mitt.) Grolle, J. Hattori Bot. Lab. 23: 49, 1960 [1961] (Grolle 1960d). Bas.: Saccogynaaustralis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 145, 1854 (Mitten 1854).

** Saccogynidiumcaldense (Ångstr.) Grolle, J. Hattori Bot. Lab. 23: 44, 1960 [1961] (Grolle 1960d). Bas.: Chiloscyphuscaldensis Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 33 (7): 80, 1876 [1877] (Ångström 1876).

** Saccogynidiumgoebelii (Herzog) Grolle, J. Hattori Bot. Lab. 23: 52, 1960 [1961] (Grolle 1960d). Bas.: Leioscyphusgoebelii Herzog, Ann. Bryol. 5: 89, 1932 (Herzog 1932a).

** Saccogynidiummuricellum (De Not.) Grolle, J. Hattori Bot. Lab. 36: 80, 1972 [1973] (Grolle and Schultze-Motel 1972). Bas.: Chiloscyphusmuricellus De Not., Epat. Borneo: 24, 1874 (De Notaris 1874).

*** Saccogynidiumvasculosum (Hook.f. et Taylor) Grolle, J. Hattori Bot. Lab. 23: 46, 1960 [1961] (Grolle 1960d). Bas.: Jungermanniavasculosa Hook.f. et Taylor, London J. Bot. 3: 461, 1844 (Hooker and Taylor 1844b).

Incertae sedis

** Saccogynidiumchiloscyphoides R.M.Schust., J. Hattori Bot. Lab. 26: 272, 1963 (Schuster 1963b).

** Saccogynidiumirregularospinum C.Gao, T.Cao et M.J.Lai, Bryologist 104 (1): 129, 2001 (Gao et al. 2001).

*** Antheliaceae R.M.Schust

*** Anthelia (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermanniasect.Anthelia Dumort., Syll. Jungerm. Europ.: 63, 1831 (Dumortier 1831).

*** Antheliajulacea (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermanniajulacea L., Sp. Pl., ed. 2: 1601, 1763 (Linnaeus 1763).

** Antheliajuratzkana (Limpr.) Trevis., Mem. Reale Ist. Lombardo Sci. (Ser. 3), C. Sci. Mat. 4 (13): 416, 1877 (Trevisan 1877). Bas.: Jungermanniajuratzkana Limpr., Hedwigia 15 (2): 18, 1876 (Limpricht 1876).

*** Arnelliaceae Nakai

Crandall-Stotler et al. (2009) placed Stephaniella and Stephaniellidium in Arnelliaceae following de Roo et al. (2007). However, Váňa et al. (2012e) argued based on further molecular evidence that Arnelliaceae should be retained as a monotypic family with a single species.

*** Arnellia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (Lindberg and Arnell 1889).

*** Arnelliafennica (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 35, 1889 (Lindberg and Arnell 1889). Bas.: Jungermanniafennica Gottsche, Hepat. Eur., Leberm. 42-44: no. 418, 1868 (Gottsche and Rabenhorst 1868).

*** Balantiopsidaceae H.Buch

by J.J. Engel with contribution by J. Váňa (Neesioscyphus)

The placement of Pseudoisotachis in Balantiopsidaceae is only preliminary (cf. Váňa 2013).

*** Acroscyphella N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Nom. nov. pro Acroscyphus N.Kitag., Acta Phytotax. Geobot. 35 (1/3): 1, 1984 (Kitagawa 1984).

*** Acroscyphellaiwatsukii (N.Kitag.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Bas.: Acroscyphusiwatsukii N.Kitag., Acta Phytotax. Geobot. 35 (1/3): 3, 1984 (Kitagawa 1984).

*** Acroscyphellaphoenicorhiza (Grolle) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Bas.: Neesioscyphusphoenicorhizus Grolle, Österr. Bot. Z. 111 (1): 27, 1964 (Grolle 1964e).

*** Acroscyphellatjiwideiensis (Sande Lac.) N.Kitag. et Grolle, Acta Phytotax. Geobot. 36 (1/3): 58, 1985 (Kitagawa and Grolle 1985). Bas.: Chiloscyphustjiwideiensis Sande Lac., Ned. Kruidk. Arch. 3: 418, 1854 [1855] (Sande Lacoste 1854).

** Pseudoisotachis Váňa, Polish Bot. J. 58 (1): 55, 2013 (Váňa 2013).

** Pseudoisotachispocsii Váňa, Polish Bot. J. 58 (1): 55, 2013 (Váňa 2013).

*** Balantiopsidoideae J.J.Engel et Váňa

*** Balantiopsis Mitt., Handb. N. Zeal. fl. 2: 751, 1867 (Hooker 1867).

*** Balantiopsisasymmetrica (Herzog) J.J.Engel, Nova Hedwigia 16: 93, 1968 (Engel 1968). Bas.: Balantiopsislatifoliavar.asymmetrica Herzog, Rev. Bryol. Lichénol. 23 (1/2): 53, 1954 (Herzog 1954).

*** Balantiopsisbisbifida (Steph.) Steph., Sp. Hepat. (Stephani) 4: 101, 1910 (Stephani 1910b). Bas.: Isotachisbisbifida Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 24, 1901 (Stephani 1901b).

*** Balantiopsisbrasiliensis Steph., Sp. Hepat. (Stephani) 4: 104, 1910 (Stephani 1910b).

*** Balantiopsiscancellata (Nees) Steph., Sp. Hepat. (Stephani) 4: 103, 1910 (Stephani 1910b). Bas.: Ptilidiumcancellatum Nees, Syn. Hepat. 2: 251, 1845 (Gottsche et al. 1845a).

*** Balantiopsisciliaris S.Hatt., J. Jap. Bot. 41 (5): 129, 1966 (Hattori 1966b).

** Balantiopsisciliarissubsp.novoguineensis S.Hatt., J. Jap. Bot. 41 (5): 131, 1966 (Hattori 1966b).

*** Balantiopsisconvexiuscula Berggr., New Zealand Hepat.: 44, 1898 (Berggren 1898).

*** Balantiopsiscrocea Herzog, Beih. Bot. Centralbl. 60B (1/2): 12, 1939 (Herzog 1939c).

*** Balantiopsisdiplophylla (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (Hooker 1867). Bas.: Jungermanniadiplophylla Hook.f. et Taylor, London J. Bot. 3: 377, 1844 (Hooker and Taylor 1844a).

** Balantiopsisdiplophyllavar.hockenii (Berggr.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 11, 1997 (Engel and Smith Merrill 1997). Bas.: Balantiopsishockenii Berggr., New Zealand Hepat.: 46, 1898 (Berggren 1898).

*** Balantiopsiserinacea (Hook.f. et Taylor) Mitt., Handb. N. Zeal. fl. 2: 753, 1867 (Hooker 1867). Bas.: Jungermanniaerinacea Hook.f. et Taylor, London J. Bot. 3: 462, 1844 (Hooker and Taylor 1844b).

*** Balantiopsislingulata R.M.Schust., Bull. Natl. Sci. Mus. Tokyo (n.ser.) 11 (1): 26, 1968 (Schuster 1968a).

*** Balantiopsismontana (Colenso) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 12, 1997 (Engel and Smith Merrill 1997). Bas.: Chiloscyphusmontanus Colenso, Trans. & Proc. New Zealand Inst. 21: 62, 1889 (Colenso 1889).

** Balantiopsisneocaledonica Pearson, J. Linn. Soc., Bot. 46 (305): 28, 1922 (Pearson 1922b).

** Balantiopsispaucidens Steph., Bih. Kongl. Svenska Vetensk.-Akad. Handl. 26 (III, 17): 29, 1901 (Stephani 1901b).

*** Balantiopsispurpurata Mitt., Rep. Challenger, Bot. 1 (3, 1): 86, 1884 (Mitten 1884b).

*** Balantiopsisrosea Berggr., New Zealand Hepat.: 43, 1898 (Berggren 1898).

*** Balantiopsissplendens (Steph.) J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 55, 1997 (Engel and Smith Merrill 1997). Bas.: Isotachissplendens Steph., Hedwigia 34 (2): 49, 1895 (Stephani 1895c).

*** Balantiopsistumida Berggr., New Zealand Hepat.: 45, 1898 (Berggren 1898).

*** Balantiopsisverrucosa J.J.Engel et G.L.Merr., Fieldiana, Bot. (n.ser.) 37: 16, 1997 (Engel and Smith Merrill 1997).

*** Isotachidoideae Grolle

*** Isotachis Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 148, 1854 (Mitten 1854).

*** Isotachisarmata (Nees) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (Gottsche 1864). Bas.: Jungermanniaarmata Nees, Syn. Hepat. 1: 129, 1844 (Gottsche et al. 1844).

*** Isotachisaubertii (Schwägr.) Mitt., J. Linn. Soc., Bot. 22 (146): 322, 1886 (Mitten 1886b). Bas.: Jungermanniaaubertii Schwägr., Hist. Musc. Hepat. Prodr.: 19, 1814 (Schwägrichen 1814).

* Isotachisboliviensis Gottsche, Sp. Hepat. (Stephani) 3: 670, 1909 (Stephani 1909a).

** Isotachischinensis C.Gao, T.Cao et J.Sun, Bryologist 105 (4): 694, 2002 [2003] (Gao et al. 2002).

** Isotachiserythrorhiza (Lehm. et Lindenb.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 338, 1885 (Spruce 1885). Bas.: Jungermanniaerythrorhiza Lehm. et Lindenb., Nov. Stirp. Pug. 4: 44, 1832 (Lehmann 1832).

*** Isotachisfragilis Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 67, 1911 (Stephani 1911b).

** Isotachisgrandis Carrington et Pearson, Proc. Linn. Soc. New South Wales (ser. 2) 2 (4): 1041, 1888 (Carrington and Pearson 1888a).

*** Isotachisgrossidens Steph., Kungl. Svenska Vetensk.-Akad. Handl. (n.ser.) 46 (9): 69, 1911 (Stephani 1911b).

** Isotachishastatistipula (Steph.) J.J.Engel, Phytotaxa 183 (4): 299, 2014 (Engel et al. 2014). Bas.: Balantiopsishastatistipula Steph., J. & Proc. Roy. Soc. New South Wales 48 (1/2): 98, 1914 (Stephani and Watts 1914).

* Isotachishians Steph., Sp. Hepat. (Stephani) 3: 665, 1909 (Stephani 1909a).

*** Isotachishumectata (Hook.f. et Taylor) Steph., Sp. Hepat. (Stephani) 3: 654, 1909 (Stephani 1909a). Bas.: Jungermanniahumectata Hook.f. et Taylor, London J. Bot. 3: 462, 1844 (Hooker and Taylor 1844b).

** Isotachisinflata Steph., Arch. Mus. Nac. Rio de Janeiro 13: 113, 1905 (Stephani 1905c).

*** Isotachisintortifolia (Hook.f. et Taylor) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (Gottsche 1864). Bas.: Jungermanniaintortifolia Hook.f. et Taylor, London J. Bot. 3: 374, 1844 (Hooker and Taylor 1844a).

** Isotachisjaponica Steph., Sp. Hepat. (Stephani) 3: 652, 1909 (Stephani 1909a).

* Isotachislacustris Herzog, Hedwigia 74 (2): 94, 1934 (Herzog 1934a).

*** Isotachislopezii (R.M.Schust.) Gradst., Mem. New York Bot. Gard. 84: 66, 1999 (Gradstein 1999). Bas.: Ruizanthuslopezii R.M.Schust., Phytologia 39 (4): 241, 1978 (Schuster 1978a).

*** Isotachislyallii Mitt., Bot. antarct. voy. II (Fl. Nov.-Zel. 2): 149, 1854 (Mitten 1854).

*** Isotachisminima Pearson, Univ. Calif. Publ. Bot. 10 (4): 322, 1923 (Pearson 1923).

*** Isotachismontana Colenso, Trans. & Proc. New Zealand Inst. 21: 68, 1889 (Colenso 1889).

*** Isotachismulticeps (Lindenb. et Gottsche) Gottsche, Mexik. Leverm.: 105, 1863 (Gottsche 1863). Bas.: Jungermanniamulticeps Lindenb. et Gottsche, Syn. Hepat. 5: 687, 1847 (Gottsche et al. 1847). 67

** Isotachismulticepsvar.fendleri Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 125, 1864 (Gottsche 1864).

** Isotachisnigella Herzog, Memoranda Soc. Fauna Fl. Fennica 27: 107, 1952 (Herzog 1952c).

*** Isotachisobtusa Steph., Sp. Hepat. (Stephani) 6: 354, 1922 (Stephani 1922).

** Isotachisolivacea R.M.Schust., J. Hattori Bot. Lab. 83: 207, 1997 (Schuster and Engel 1997).

*** Isotachisplicata J.J.Engel, J. Hattori Bot. Lab. 83: 210, 1997 (Schuster and Engel 1997).

** Isotachispusilla Steph., Sp. Hepat. (Stephani) 3: 655, 1909 (Stephani 1909a).

** Isotachisriparia Rodway, Tasm. Bryoph.: 63, 1917 (Rodway 1917b).

*** Isotachisserrulata (Sw.) Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 121, 1864 (Gottsche 1864). Bas.: Jungermanniaserrulata Sw., Prodr. (Swartz): 143, 1788 (Swartz 1788).

*** Isotachisspegazziniana C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 220, 1885 (Massalongo 1885).

* Isotachissprucei Beauverd, Sp. Hepat. (Stephani) 6: 572, 1924 (Stephani 1924). Nom. nov. pro Isotachis trifida Steph., Sp. Hepat. (Stephani) 6: 356, 1922 (Stephani 1922), nom. illeg.

*** Isotachisstephanii E.S.Salmon, Rev. Bryol. 28 (4): 75, 1901 (Salmon 1901).

* Isotachisvexans Steph., Sp. Hepat. (Stephani) 3: 662, 1909 (Stephani 1909a).

*** Isotachiswestlandica (E.A.Hodgs.) R.M.Schust., Nova Hedwigia 15: 455, 1968 (Schuster 1968b). Bas.: Rhizocauliawestlandica E.A.Hodgs., Trans. Roy. Soc. New Zealand, Bot. 3 (4): 78, 1965 (Hodgson 1965).

*** Neesioscyphus Grolle, Österr. Bot. Z. 111 (1): 19, 1964 (Grolle 1964e).

*** Neesioscyphusallionii (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 185, 1966 (Grolle 1966c). Bas.: Isotachisallionii Steph., Sp. Hepat. (Stephani) 6: 350, 1922 (Stephani 1922).

*** Neesioscyphusargillaceus (Nees) Grolle, Österr. Bot. Z. 111 (1): 24, 1964 (Grolle 1964e). Bas.: Jungermanniaargillacea Nees, Fl. Bras. (Martius) 1 (1): 338, 1833 (Nees 1833a).

*** Neesioscyphusbicuspidatus (Steph.) Grolle, Rev. Bryol. Lichénol. 34 (1/2): 182, 1966 (Grolle 1966c). Bas.: Isotachisbicuspidata Steph., Symb. Antill. 2: 471, 1901 (Stephani 1901f).

*** Neesioscyphuscarneus (Nees) Grolle, Österr. Bot. Z. 111 (1): 20, 1964 (Grolle 1964e). Bas.: Jungermanniacarnea Nees, Fl. Bras. (Martius) 1 (1): 337, 1833 (Nees 1833a).

*** Neesioscyphushomophyllus (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 188, 1964 (Grolle 1964f). Bas.: Jungermanniahomophylla Nees, Fl. Bras. (Martius) 1 (1): 336, 1833 (Nees 1833a).

** Ruizanthoideae R.M.Schust. ex J.J.Engel et G.L.Merr

*** Ruizanthus R.M.Schust., Phytologia 39 (4): 240, 1978 (Schuster 1978a).

*** Ruizanthusvenezuelanus R.M.Schust., Phytologia 39 (4): 240, 1978 (Schuster 1978a).

*** Blepharidophyllaceae R.M.Schust. ex J.J.Engel

by J.J. Engel

*** Blepharidophyllum Ångstr., Öfvers. Kongl. Vetensk.-Akad. Förh. 30 (5): 151, 1873 (Ångström 1873).

*** Blepharidophyllumdensifolium (Hook.) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (Massalongo 1885). Bas.: Jungermanniadensifolia Hook., Musci Exot. 1: tab. 36, 1818 (Hooker 1818).

*** Blepharidophyllumvertebrale (Gottsche) Ångstr. ex C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 208, 1885 (Massalongo 1885). Bas.: Scapaniavertebralis Gottsche, Syn. Hepat. 1: 72, 1844 (Gottsche et al. 1844).

*** Clandarium (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (Schuster 1983a). Bas.: Blepharidophyllumsubg.Clandarium Grolle, J. Hattori Bot. Lab. 28: 65, 1965 (Grolle 1965a).

*** Clandariumclandestinum (Mont.) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (Schuster 1983a). Bas.: Plagiochilaclandestina Mont., Ann. Sci. Nat. Bot. (sér. 2) 19: 247, 1843 (Montagne 1843).

*** Clandariumgottscheanum (Grolle) R.M.Schust., New Manual Bryol. 1: 541, 1983 [1984] (Schuster 1983a). Bas.: Blepharidophyllumgottscheanum Grolle, J. Hattori Bot. Lab. 28: 69, 1965 (Grolle 1965a).

*** Clandariumxiphophyllum (Grolle) R.M.Schust., Phytologia 56 (2): 68, 1984 (Schuster 1984). Bas.: Blepharidophyllumxiphophyllum Grolle, J. Hattori Bot. Lab. 28: 65, 1965 (Grolle 1965a).

*** Calypogeiaceae Arnell

by M.A.M. Renner

Calypogeiaceae is shown to be monophyletic (Masuzaki et al. 2010b) and the classification follows Schuster (2000a) and Masuzaki et al. (2010b).

*** Calypogeia Raddi, Jungermanniogr. Etrusca: 31, 1818 (Raddi 1818a) nom. conserv.

** subg.Asperifoliae (Warnst.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 115, 1969 (Schuster 1969b). Bas.: Calypogeia [unranked] Asperifoliae Warnst., Bryol. Z. 1 (7): 111, 1917 (Warnstorf 1917).

*** Calypogeiaarguta Nees et Mont., Naturgesch. Eur. Leberm. 3: 24, 1838 (Nees 1838b).

** Calypogeiasullivantii Austin, Hepat. bor.-amer.: 19, 1873 (Austin 1873).

** subg.Calypogeia

*** Calypogeiaandicola Bischl., Candollea 18: 79, 1962 (Bischler 1962a).

** Calypogeiaannabonensis Steph., Sp. Hepat. (Stephani) 6: 447, 1924 (Stephani 1924).

** Calypogeiaazorica Bischl., Rev. Bryol. Lichénol. 37 (1): 116, 1970 (Bischler 1970).

*** Calypogeiaazurea Stotler et Crotz, Taxon 32 (1): 74, 1983 (Stotler and Crotz 1983).

*** Calypogeiabidentula (F.Weber) Nees, Syn. Hepat. 2: 199, 1845 (Gottsche et al. 1845a). Bas.: Jungermanniabidentula F.Weber, Hist. Musc. Hepat. Prodr.: 38, 1815 (Weber 1815).

** Calypogeiafalcata Bischl., Candollea 18: 112, 1962 (Bischler 1962c).

*** Calypogeiafissa (L.) Raddi, Jungermanniogr. Etrusca: 33, 1818 (Raddi 1818a). Bas.: Mniumfissum L., Sp. Pl. 1: 1114, 1753 (Linnaeus 1753), nom. conserv. 68

** Calypogeiafissasubsp.neogaea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 169, 1969 (Schuster 1969b).

** Calypogeiagoebelii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (Stephani 1908d). Bas.: Kantiusgoebelii Schiffn., Nova Acta Acad. Caes. Leop.-Carol. German. Nat. Cur. 60 (2): 260, 1893 (Schiffner 1893a).

** Calypogeiagoebeliivar.siamensis N.Kitag., Beih. Nova Hedwigia 90: 165, 1988 (Kitagawa 1988).

*** Calypogeiagrandistipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (Stephani 1908d). Bas.: Kantiusgrandistipulus Steph., Hedwigia 34 (2): 52, 1895 (Stephani 1895c).

*** Calypogeiaintegristipula Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 662 (394), 1908 (Stephani 1908d).

*** Calypogeialaxa Gottsche et Lindenb., Syn. Hepat. 5: 713, 1847 (Gottsche et al. 1847).

*** Calypogeialechleri (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (Stephani 1908d). Bas.: Kantiuslechleri Steph., Hedwigia 34 (2): 53, 1895 (Stephani 1895c). 69

** Calypogeialechlerivar.densifolia (Steph.) Bischl., Candollea 18: 101, 1962 (Bischler 1962c). Bas.: Kantiusdensifolius Steph., Hedwigia 34 (2): 52, 1895 (Stephani 1895c).

** Calypogeialongifolia Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (Stephani 1924).

** Calypogeialophocoleoides Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 677 (409), 1908 (Stephani 1908d).

** Calypogeiamascarenensis Bischl., Rev. Bryol. Lichénol. 37 (1): 89, 1970 (Bischler 1970).

** Calypogeiamicrostipula (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 670 (402), 1908 (Stephani 1908d). Bas.: Kantiusmicrostipulus Steph., Hedwigia 34 (2): 53, 1895 (Stephani 1895c).

*** Calypogeiamiquelii Mont. ex Gottsche, Lindenb. et Nees, Syn. Hepat. 2: 200, 1845 (Gottsche et al. 1845a).

*** Calypogeiamuelleriana (Schiffn.) Müll.Frib., Beih. Bot. Centralbl. 10 (4/5): 217, 1901 (Müller 1901b). Bas.: Kantiusmuellerianus Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 48: 342, 1900 (Schiffner 1900d). 70

** Calypogeiamuellerianasubsp.blomquistii R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 187, 1969 (Schuster 1969b).

*** Calypogeianeesiana (C.Massal. et Carestia) Müll.Frib., Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (Loeske 1905). Bas.: Kantiustrichomanisvar.neesianus C.Massal. et Carestia, Nuovo Giorn. Bot. Ital. 12 (4): 351, 1880 (Massalongo and Carestia 1880).

** Calypogeianeesianasubsp.subalpina (Inoue) Inoue, Mem. Natl. Sci. Mus. (Tokyo) 4: 58, 1971 (Inoue 1971a). Bas.: Calypogeiasubalpina Inoue, J. Jap. Bot. 37 (4): 103, 1962 (Inoue 1962b).

*** Calypogeiaoblata Herzog, Svensk Bot. Tidskr. 51 (1): 189, 1957 (Herzog 1957a).

*** Calypogeiaperuviana Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 9: 47, 1838 (Montagne 1838).

*** Calypogeiarhombifolia (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 667 (399), 1908 (Stephani 1908d). Bas.: Kantiusrhombifolius Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 413, 1885 (Spruce 1885).

** Calypogeiarhombifoliavar.colombiana Bischl., Candollea 18: 104, 1962 (Bischler 1962c).

*** Calypogeiasphagnicola (Arnell et J.Perss.) Warnst. et Loeske, Verh. Bot. Vereins Prov. Brandenburg 47: 320, 1905 (Loeske 1905). Bas.: Kantiussphagnicola Arnell et J.Perss., Rev. Bryol. 29 (2): 26, 1902 (Arnell 1902). 71

*** Calypogeiasubintegra (Gottsche, Lindenb. et Nees) Bischl., Candollea 18: 75, 1962 (Bischler 1962a). Bas.: Calypogeiaperuviana β subintegra Gottsche, Lindenb. et Nees, Syn. Hepat. 5: 712, 1847 (Gottsche et al. 1847).

** Calypogeiasubintegravar.dussiana (Steph.) Bischl., Candollea 18: 77, 1962 (Bischler 1962a). Bas.: Calypogeiadussiana Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 672 (404), 1908 (Stephani 1908d).

*** Calypogeiasuecica (Arnell et J.Perss.) Müll.Frib., Beih. Bot. Centralbl. 17 (2): 224, 1904 (Müller 1904). Bas.: Kantiussuecicus Arnell et J.Perss., Rev. Bryol. 29 (2): 29, 1902 (Arnell 1902).

*** Calypogeiatenax (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 664 (396), 1908 (Stephani 1908d). Bas.: Kantiustenax Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 416, 1885 (Spruce 1885).

** Calypogeiauncinulatula Herzog, Hedwigia 67 (6): 250, 1927 (Herzog 1927).

Incertae sedis

*** Calypogeiaaeruginosa Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (Mitten 1860c).

* Calypogeiaamazonica (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 680 (412), 1908 (Stephani 1908d). Bas.: Kantiusamazonicus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 415, 1885 (Spruce 1885). 72

** Calypogeiaangusta Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908 (Stephani 1908d).

** Calypogeiaapiculata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 668 (400), 1908 (Stephani 1908d). Bas.: Kantiusapiculatus Steph., Hedwigia 34 (2): 51, 1895 (Stephani 1895c).

** Calypogeiaasakawana S.Hatt. ex Inoue, J. Jap. Bot. 39 (4): 107, 1964 (Inoue 1964a).

** Calypogeiaceylanica S.Hatt. et Mizut., Candollea 23: 288, 1968 (Hattori 1968).

** Calypogeiacontracta Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 139, 1975 (Inoue 1975b).

** Calypogeiacuspidata (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 669 (401), 1908 (Stephani 1908d). Bas.: Kantiuscuspidatus Steph., Bull. Herb. Boissier 5 (10): 846, 1897 (Stephani 1897c).

* Calypogeiadecurrens (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 675 (407), 1908 (Stephani 1908d). Bas.: Kantiusdecurrens Steph., Hedwigia 34 (2): 52, 1895 (Stephani 1895c).

** Calypogeiaformosana Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 186, 1934 (Horikawa 1934).

** Calypogeiafujisana Inoue, Bull. Natl. Sci. Mus. Tokyo, B 1 (4): 135, 1975 (Inoue 1975b).

** Calypogeiagranulata Inoue, J. Jap. Bot. 43 (10/11): 468, 1968 (Inoue 1968b).

** Calypogeiajaponica Steph., Sp. Hepat. (Stephani) 6: 448, 1924 (Stephani 1924).

** Calypogeiakhasiana Ajit P.Singh et V.Nath, Taiwania 52 (4): 320, 2007 (Singh and Nath 2007a).

** Calypogeialatissima Steph., Sp. Hepat. (Stephani) 6: 449, 1924 (Stephani 1924).

*** Calypogeialunata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 107, 1860 [1861] (Mitten 1860c).

** Calypogeiamarginella Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 106, 1860 [1861] (Mitten 1860c).

** Calypogeiaobovata R.M.Schust., Phytologia 39 (4): 242, 1978 (Schuster 1978a).

* Calypogeiasteyermarkii Fulford, Mem. New York Bot. Gard. 11 (3): 305, 1968 (Fulford 1968).

** Calypogeiatosana (Steph.) Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 678 (410), 1908 (Stephani 1908d). Bas.: Kantiustosanus Steph., Hedwigia 34 (2): 54, 1895 (Stephani 1895c).

** Calypogeiaudarii Sudipa Das et D.K.Singh, Nelumbo 53: 194, 2011 (Das and Singh 2011).

** Eocalypogeia (R.M.Schust.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (Schuster 1995a). Bas.: Metacalypogeiasubg.Eocalypogeia R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 107, 1969 (Schuster 1969b).

** Eocalypogeiaquelpaertensis (S.Hatt. et Inoue) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (Schuster 1995a). Bas.: Metacalypogeiaquelpaertensis S.Hatt. et Inoue, J. Hattori Bot. Lab. 25: 129, 1962 (Hattori et al. 1962).

** Eocalypogeiaschusterana (S.Hatt. et Mizut.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 861, 1995 (Schuster 1995a). Bas.: Metacalypogeiaschusterana S.Hatt. et Mizut., Misc. Bryol. Lichenol. 4 (8): 121, 1967 (Hattori and Mizutani 1967).

** Metacalypogeia (S.Hatt.) Inoue, J. Hattori Bot. Lab. 21: 231, 1959 (Inoue 1959b). Bas.: Calypogeiasubg.Metacalypogeia S.Hatt., J. Hattori Bot. Lab. 18: 83, 1957 (Hattori 1957b).

*** Metacalypogeiaalternifolia (Nees) Grolle, Österr. Bot. Z. 111 (2/3): 185, 1964 (Grolle 1964f). Bas.: Mastigobryumalternifolium Nees, Syn. Hepat. 2: 216, 1845 (Gottsche et al. 1845a).

** Metacalypogeiacordifolia (Steph.) Inoue, J. Hattori Bot. Lab. 21: 233, 1959 (Inoue 1959b). Bas.: Calypogeiacordifolia Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 661 (393), 1908 (Stephani 1908d).

** Mizutania Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (Furuki and Iwatsuki 1989).

*** Mizutaniariccardioides Furuki et Z.Iwats., J. Hattori Bot. Lab. 67: 291, 1989 (Furuki and Iwatsuki 1989).

*** Mnioloma Herzog, Ann. Bryol. 3: 115, 1930 (Herzog 1930a).

** subg.Caracoma (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 833, 1995 (Schuster 1995a). Bas.: Calypogeiasubg.Caracoma Bischl., Candollea 18: 26, 1962 (Bischler 1962b).

** Mniolomabolivianum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (Schuster 2000a). Bas.: Calypogeiaboliviana Fulford, Mem. New York Bot. Gard. 11 (3): 291, 1968 (Fulford 1968).

*** Mniolomacaespitosum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (Schuster 1995a). Bas.: Kantiuscaespitosus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 412, 1885 (Spruce 1885).

*** Mniolomacellulosum (Spreng.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 836, 1995 (Schuster 1995a). Bas.: Jungermanniacellulosa Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (1): 232, 1827 (Sprengel 1827a).

*** Mniolomacrenulatum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (Schuster 1995a). Bas.: Calypogeiacrenulata Bischl., Candollea 18: 35, 1962 (Bischler 1962b).

*** Mniolomacyclostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 843, 1995 (Schuster 1995a). Bas.: Kantiuscyclostipus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 411, 1885 (Spruce 1885).

** Mniolomaelliottii (Steph.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 841, 1995 (Schuster 1995a). Bas.: Calypogeiaelliottii Steph., Bull. Herb. Boissier (sér. 2) 8 (9): 663 (395), 1908 (Stephani 1908d).

** Mniolomafissistipulum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (Schuster 1995a). Bas.: Calypogeiafissistipula Bischl., Candollea 18: 47, 1962 (Bischler 1962b).

*** Mniolomafuscum (Lehm.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 848, 1995 (Schuster 1995a). Bas.: Jungermanniafusca Lehm., Linnaea 4: 360, 1829 (Lehmann 1829).

*** Mniolomanephrostipum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (Schuster 1995a). Bas.: Kantiusnephrostipus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 412, 1885 (Spruce 1885).

** Mniolomanovaezelandiae J.J.Engel, Cryptog. Bryol. 27 (1): 111, 2006 (Engel 2006a).

*** Mniolomaparallelogramum (Spruce) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 847, 1995 (Schuster 1995a). Bas.: Kantiusparallelogramus Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 413, 1885 (Spruce 1885).

** Mniolomaretusum (Bischl.) R.M.Schust., Fragm. Florist. Geobot. 40 (2): 839, 1995 (Schuster 1995a). Bas.: Calypogeiaretusa Bischl., Candollea 18: 33, 1962 (Bischler 1962b).

** Mniolomastamatotonum M.A.M.Renner et E.A.Br., Fieldiana, Bot. (n.ser.) 47: 173, 2008 (Renner and Brown 2008).

*** Mniolomavenezuelanum (Fulford) R.M.Schust., Beih. Nova Hedwigia 118: 509, 2000 (Schuster 2000a). Bas.: Calypogeiavenezuelana Fulford, Mem. New York Bot. Gard. 11 (3): 287, 1968 (Fulford 1968).

** subg.Mnioloma, Fragm. Florist. Geobot. 40 (2): 833, 1995 (Schuster 1995a).

*** Mniolomarhynchophyllum Herzog, Ann. Bryol. 3: 120, 1930 (Herzog 1930a).

*** Endogemmataceae Konstant., Vilnet et A.V.Troitsky

by N.A. Konstantinova

Vilnet et al. (2011) described the monotypic family Endogemmataceae based on molecular evidence after a re-evaluation of Solenostomataceae.

*** Endogemma Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (Vilnet et al. 2011).

*** Endogemmacaespiticia (Lindenb.) Konstant., Vilnet et A.V.Troitsky, Folia Cryptog. Estonica 48: 132, 2011 (Vilnet et al. 2011). Bas.: Jungermanniacaespiticia Lindenb., Syn. hepat. eur: 67, 1829 (Lindenberg 1829).

*** Geocalycaceae H.Klinggr

Placement of Geocalycaceae in Jungermanniinae follows Shaw et al. (2015).

*** Geocalyx Nees, Naturgesch. Eur. Leberm. 1: 97, 1833 (Nees 1833c).

*** Geocalyxcaledonicus Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 205 (265), 1908 (Stephani 1908h).

*** Geocalyxgraveolens (Schrad.) Nees, Naturgesch. Eur. Leberm. 2: 397, 1836 (Nees 1836). Bas.: Jungermanniagraveolens Schrad., Syst. Samml. Crypt. Gew. 2: 6, 1797 (Schrader 1797).

** Geocalyxlancistipulus (Steph.) S.Hatt., J. Jap. Bot. 28 (8): 234, 1953 (Hattori 1953a). Bas.: Lophocolealancistipula Steph., Sp. Hepat. (Stephani) 6: 281, 1922 (Stephani 1922).

** Geocalyxorientalis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxix, 1889 [1890] (Bescherelle and Spruce 1889).

*** Gymnomitriaceae H.Klinggr

by J. Váňa

The treatment of the family follows Váňa et al. (2010b) with some modifications. Nardioideae is included following Vilnet et al. (2010) and Váňa et al. (2014c). Some re-arrangements in Apomarsupella, Gymnomitrion and Marsupella were done by Vilnet et al (2010) and Shaw et al. (2015). Further notes on nomenclature and taxonomy can be found in Váňa et al. (2013d). Herzogobryum and Nothogymnomitrion were removed from the family by Váňa et al. (2013e). The placement of Acrolophozia, Nanomarsupella and Paramomitrion is provisional. Inclusion of Cryptocoleopsis follows Shaw et al (2015).

*** Acrolophozia R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 259, 1966 (Schuster 1966b).

*** Acrolophoziafuegiana R.M.Schust., Nova Hedwigia 15: 499, 1968 (Schuster 1968b).

*** Acrolophoziapectinata R.M.Schust., Rev. Bryol. Lichénol. 34 (1/2): 261, 1966 (Schuster 1966b).

*** Acrolophoziasulcata Hässel, J. Bryol. 11 (1): 108, 1980 (Hässel 1980).

** Nanomarsupella R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (Hagborg et al. 2013). Based on: Marsupellasubg.Nanomarsupella R.M.Schust., Phytologia 39 (4): 248, 1978 (Schuster 1978a).

*** Nanomarsupellaxenophylla (R.M.Schust.) R.M.Schust. ex A.Hagborg, L.Söderstr. et von Konrat, Phytotaxa 112 (1): 16, 2013 (Hagborg et al. 2013). Bas.: Marsupellaxenophylla R.M.Schust., Phytologia 39 (4): 248, 1978 (Schuster 1978a).

** Paramomitrion R.M.Schust., J. Hattori Bot. Lab. 80: 134, 1996 (Schuster 1996a).

*** Paramomitrionparadoxum R.M.Schust., J. Hattori Bot. Lab. 80: 135, 1996 (Schuster 1996a).

*** Gymnomitrioideae T.Jensen

** Cryptocoleopsis Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (Amakawa 1959a).

*** Cryptocoleopsisimbricata Amakawa, J. Hattori Bot. Lab. 21: 274, 1959 (Amakawa 1959a).

*** Gymnomitrion Corda, Gen. hepat.: 651, 1829 (Corda 1829) nom. conserv.

*** Gymnomitrionadustum Nees, Naturgesch. Eur. Leberm. 1: 120, 1833 (Nees 1833c).

*** Gymnomitrionafricanum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (Horikawa 1943). Bas.: Acoleaafricana Steph., Sp. Hepat. (Stephani) 6: 77, 1917 (Stephani 1917a).

*** Gymnomitrionalpinum (Gottsche ex Husn.) Schiffn., Österr. Bot. Z. 53 (7): 280, 1903 (Schiffner 1903a). Bas.: Sarcocyphosalpinus Gottsche ex Husn., Hepaticol. gall. 1: 13, 1875 (Husnot 1875).

*** Gymnomitrionasperulatum R.M.Schust., Acta Acad. Ped. Agr., Sect. Biol. 24: 114, 2003 (Váňa 2003).

*** Gymnomitrionatrofilum Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (Váňa 1976b).

*** Gymnomitrionbolivianum (Steph.) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Anastrophyllumbolivianum Steph., Biblioth. Bot. 87 (2): 186, 1916 (Stephani 1916a).

*** Gymnomitrionbrevissimum (Dumort.) Warnst., Hedwigia 53 (3): 196, 1913 (Warnstorf 1913). Bas.: Acoleabrevissima Dumort., Syll. Jungerm. Europ.: 76, 1831 (Dumortier 1831).

*** Gymnomitrioncommutatum (Limpr.) Schiffn., Magyar Bot. Lapok 13: 304, 1914 [1915] (Schiffner 1914b). Bas.: Sarcocyphoscommutatus Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 314, 1879 [1880] (Limpricht 1879).

*** Gymnomitrionconcinnatum (Lightf.) Corda, Gen. hepat.: 651, 1829 (Corda 1829). Bas.: Jungermanniaconcinnata Lightf., Fl. Scot. 2: 786, 1777 (Lightfoot 1777), nom. conserv.

*** Gymnomitrioncorallioides Nees, Naturgesch. Eur. Leberm. 1: 118, 1833 (Nees 1833c).

*** Gymnomitrioncrenatilobum Grolle, Khumbu Himal 1 (4): 278, 1966 (Grolle 1966k).

*** Gymnomitrioncrenulatum Gottsche ex Carrington, Trans. Bot. Soc. Edinburgh 7 (3): 444, 1863 (Carrington 1863).

*** Gymnomitrioncrystallocaulon (Grolle) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Marsupellacrystallocaulon Grolle, Khumbu Himal 1 (4): 281, 1966 (Grolle 1966k).

** Gymnomitrionincompletum (Gottsche) R.M.Schust. ex Váňa, J. Hattori Bot. Lab. 40: 186, 1976 (Váňa 1976a). Bas.: Jungermanniaincompleta Gottsche, Linnaea 28 (5): 551, 1856 [1857] (Gottsche 1856).

*** Gymnomitrionlaceratum (Steph.) Horik., Acta Phytotax. Geobot. 13: 212, 1943 (Horikawa 1943). Bas.: Acolealacerata Steph., Sp. Hepat. (Stephani) 6: 78, 1917 (Stephani 1917a).

*** Gymnomitrionminiatum Lindenb. et Gottsche, Syn. Hepat. 4: 617, 1846 (Gottsche et al. 1846).

*** Gymnomitrionminutulum (Hässel) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Marsupellaminutula Hässel, J. Bryol. 11 (1): 123, 1980 (Hässel 1980).

** Gymnomitrionmoralesae Váňa, J. Hattori Bot. Lab. 48: 230, 1980 (Váňa 1980).

*** Gymnomitrionmucronulatum (N.Kitag.) N.Kitag., Acta Phytotax. Geobot. 19 (2/3): 53, 1962 (Kitagawa 1962a). Bas.: Gymnomitrionconcinnatumvar.mucronulatum N.Kitag., Acta Phytotax. Geobot. 18 (2/3): 38, 1959 (Kitagawa 1959).

** Gymnomitrionmucrophorum R.M.Schust., Bryologist 98 (2): 243, 1995 (Schuster 1995d).

*** Gymnomitrionnigrum (Grolle et Váňa) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Marsupellanigra Grolle et Váňa, J. Hattori Bot. Lab. 40: 186, 1976 (Váňa 1976a).

*** Gymnomitrionnoguchianum S.Hatt., J. Jap. Bot. 27 (2): 55, 1952 (Hattori 1952b).

*** Gymnomitrionobtusilobum N.Kitag., Bull. Univ. Mus. Univ. Tokyo 8: 229, 1975 (Hattori 1975e).

*** Gymnomitrionobtusum Lindb., Morgonbladet (Helsinki) 1877 (30, 6 Feb): 2, 1877 (Lindberg 1877a).

*** Gymnomitrionpacificum Grolle, Trans. Brit. Bryol. Soc. 5 (1): 92, 1966 (Grolle 1966f).

*** Gymnomitrionrevolutum (Nees) H.Philib., Rev. Bryol. 17 (3): 34, 1890 (Philibert 1890). Bas.: Sarcocyphosrevolutus Nees, Naturgesch. Eur. Leberm. 2: 419, 1836 (Nees 1836).

** Gymnomitrionrevolutumsubsp.novoguineanensis (R.M.Schust.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Apomarsupellarevolutasubsp.novoguineanensis R.M.Schust., J. Hattori Bot. Lab. 80: 90, 1996 (Schuster 1996a).

*** Gymnomitrionrubidum (Mitt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Jungermanniarubida Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 90, 1860 [1861] (Mitten 1860c).

*** Gymnomitrionsetaceum Grolle et Váňa, J. Hattori Bot. Lab. 41: 411, 1976 (Váňa 1976b).

*** Gymnomitrionsinense Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 176, 1951 (Müller 1951b).

** Gymnomitrionstrictum (Berggr.) R.M.Schust., J. Hattori Bot. Lab. 26: 280, 1963 (Schuster 1963b). Bas.: Cesiusstrictus Berggr., New Zealand Hepat.: 2, 1898 (Berggren 1898).

** Gymnomitrionstrictumvar.inaequale R.M.Schust., J. Hattori Bot. Lab. 80: 118, 1996 (Schuster 1996a).

*** Gymnomitrionsubintegrum (S.W.Arnell) Váňa, Novon 20 (2): 225, 2010 (Váňa et al. 2010c). Bas.: Marsupellasubintegra S.W.Arnell, Ark. Bot. (n.ser.) 3 (16): 545, 1956 (Arnell 1956e).

*** Gymnomitriontruncatoapiculatum Herzog, Hedwigia 74 (2): 81, 1934 (Herzog 1934a).

*** Gymnomitrionverrucosum W.E.Nicholson, Symb. Sin. 5: 10, 1930 (Nicholson et al. 1930).

*** Marsupella Dumort., Commentat. Bot. (Dumortier): 114, 1822 (Dumortier 1822).

*** Marsupellaalata S.Hatt. et N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 79, 1960 (Kitagawa 1960b).

*** Marsupellaandreaeoides (Lindb.) Müll.Frib., Feddes Repert. Spec. Nov. Regni Veg. 54 (2/3): 214, 1951 (Müller 1951a). Bas.: Cesiusandreaeoides Lindb., Meddel. Soc. Fauna Fl. Fenn. 14: 68, 1887 (Lindberg 1887a).

*** Marsupellaapiculata Schiffn., Österr. Bot. Z. 53 (6): 249, 1903 (Schiffner 1903b).

*** Marsupellaaquatica (Lindenb.) Schiffn., Sitzungsber. deutsch. naturwiss.-med. Vereins Böhmen “Lotos” Prag 44 (8): 267, 1896 [1897] (Schiffner 1896a). Bas.: Jungermanniaemarginatavar.aquatica Lindenb., Syn. hepat. eur: 75, 1829 (Lindenberg 1829).

*** Marsupellaarctica (Berggr.) Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 26, 1906 (Bryhn 1906). Bas.: Sarcocyphosemarginatusvar.arcticus Berggr., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 13 (7): 96, 1875 (Berggren 1875).

*** Marsupellaboeckii (Austin) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 409, 1893 (Kaalaas 1893b). Bas.: Sarcocyphosboeckii Austin, Bull. Torrey Bot. Club 3 (3): 9, 1872 (Austin 1872).

*** Marsupellabolanderi (Austin) Underw., Zoe 1 (12): 365, 1891 (Underwood 1891). Bas.: Sarcocyphosbolanderi Austin, Bull. Torrey Bot. Club 3 (3): 9, 1872 (Austin 1872).

*** Marsupellacondensata (Ångstr. ex C.Hartm.) Lindb. ex Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 420, 1893 (Kaalaas 1893b). Bas.: Gymnomitrioncondensatum Ångstr. ex C.Hartm., Handb. Skand. fl. (ed. 10): 128, 1871 (Hartman 1871).

*** Marsupelladisticha Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 164 (25), 1901 (Stephani 1901h).

*** Marsupellaemarginata (Ehrh.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (Dumortier 1835). Bas.: Jungermanniaemarginata Ehrh., Hannover. Mag. 22 (8): 141, 1784 (Ehrhart 1784).

** Marsupellaemarginatasubsp.tubulosa (Steph.) N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 76, 1960 (Kitagawa 1960b). Bas.: Marsupellatubulosa Steph., Bull. Herb. Boissier 5 (2): 99, 1897 (Stephani 1897b).

** Marsupellaemarginatasubsp.tubulosavar.apertifolia (Steph.) N.Kitag., J. Hattori Bot. Lab. 26: 89, 1963 (Kitagawa 1963b). Bas.: Marsupellaapertifolia Steph., Bull. Herb. Boissier (sér. 2) 1 (2): 162 (23), 1901 (Stephani 1901h).

** Marsupellaemarginatasubsp.tubulosavar.patens N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 77, 1960 (Kitagawa 1960b).

** Marsupellaemarginatasubsp.tubulosavar.tubulosa (Steph.) N.Kitag. ex Váňa et L.Söderstr., Phytotaxa 183 (4): 288, 2014 (Váňa et al. 2014c). Bas.: Marsupellatubulosa Steph., Bull. Herb. Boissier 5 (2): 99, 1897 (Stephani 1897b).

*** Marsupellafunckii (F.Weber et D.Mohr) Dumort., Recueil Observ. Jungerm.: 24, 1835 (Dumortier 1835). Bas.: Jungermanniafunckii F.Weber et D.Mohr, Bot. Taschenb. (Weber): 422, 1807 (Weber and Mohr 1807).

*** Marsupellamicrophylla R.M.Schust., Phytologia 39 (4): 249, 1978 (Schuster 1978a).

** Marsupellaminutissima N.Kitag., Mem. Coll. Sci. Kyoto Imp. Univ., Ser. B, Biol. 27: 81, 1960 (Kitagawa 1960b).

*** Marsupellaneesii Sande Lac. ex Schiffn., Consp. Hepat. Arch. Ind.: 70, 1898 (Schiffner 1898b). Based on: Sarcocyphosneesii Nees ex Sande Lac., Ann. Mus. Bot. Lugduno-Batavi 1: 288, 1864 (Sande Lacoste 1864), nom. inval.

*** Marsupellaparoica R.M.Schust., Bryologist 60 (2): 145, 1957 (Schuster 1957b).

*** Marsupellaprofunda Lindb., Rev. Bryol. 14 (2): 19, 1887 (Lindberg 1887b).

*** Marsupellapseudofunckii S.Hatt., J. Hattori Bot. Lab. 4: 63, 1950 (Hattori 1950).

*** Marsupellasparsifolia (Lindb.) Dumort., Bull. Soc. Roy. Bot. Belgique 13: 128, 1874 (Dumortier 1874). Bas.: Sarcocyphossparsifolius Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 9: 280, 1868 (Lindberg 1868b).

** Marsupellasparsifoliasubsp.childii R.M.Schust., Phytotaxa 183 (4): 288, 2014 (Váňa et al. 2014c). Based on: Marsupellasparsifoliasubsp.childii R.M.Schust., J. Hattori Bot. Lab. 80: 61, 1996 (Schuster 1996a), nom. inval.

*** Marsupellasphacelata (Giesecke ex Lindenb.) Dumort., Recueil Observ. Jungerm.: 24, 1835 (Dumortier 1835). Bas.: Jungermanniasphacelata Giesecke ex Lindenb., Syn. hepat. eur: 76, 1829 (Lindenberg 1829).

*** Marsupellaspiniloba R.M.Schust. et Damsh., Phytologia 63 (5): 326, 1987 (Schuster and Damsholt 1987).

*** Marsupellasprucei (Limpr.) Bernet, Cat. hép. Suisse: 33, 1888 (Bernet 1888). Bas.: Sarcocyphossprucei Limpr., Flora 64 (5): 72, 1881 (Limpricht 1881).

** Marsupellastableri Spruce, Rev. Bryol. 8 (6): 96, 1881 (Spruce 1881a).

*** Marsupellastoloniformis N.Kitag., J. Hattori Bot. Lab. 30: 201, 1967 (Kitagawa 1967b).

** Marsupellastoloniformissubsp.vermiformis R.M.Schust., J. Hattori Bot. Lab. 80: 72, 1996 (Schuster 1996a).

*** Marsupellayakushimensis (Horik.) S.Hatt., Bull. Tokyo Sci. Mus. 11: 80, 1944 (Hattori 1944d). Bas.: Sphenolobusyakushimensis Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 156, 1934 (Horikawa 1934).

*** Poeltia Grolle, Khumbu Himal 1 (4): 280, 1966 (Grolle 1966k).

*** Poeltiacampylata Grolle, Khumbu Himal 1 (4): 280, 1966 (Grolle 1966k).

*** Prasanthus Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889 (Lindberg and Arnell 1889).

** Prasanthusjamalicus Potemkin, Ann. Bot. Fenn. 29 (4): 319, 1992 (Potemkin 1992).

*** Prasanthussuecicus (Gottsche) Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 62, 1889 (Lindberg and Arnell 1889). Bas.: Gymnomitrionsuecicum Gottsche, Fl. Danica 16 (48): 20, 1871 (Lange 1871).

*** Nardioideae Váňa

*** Nardia Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (Gray 1821) nom. conserv.

*** Nardiaarnelliana Grolle, Bot. Mag. (Tokyo) 77 (914): 297, 1964 (Grolle 1964b).

*** Nardiaassamica (Mitt.) Amakawa, J. Hattori Bot. Lab. 26: 23, 1963 (Amakawa 1963). Bas.: Jungermanniaassamica Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c).

*** Nardiabreidleri (Limpr.) Lindb., Helsingf. Dagbl. 1880 (311, 15 Nov.): 2, 1880 (Lindberg 1880b). Bas.: Aliculariabreidleri Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 57: 311, 1879 [1880] (Limpricht 1879).

*** Nardiacompressa (Hook.) Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (Gray 1821). Bas.: Jungermanniacompressa Hook., Brit. Jungermann.: tab. 58, 1813 (Hooker 1813).

*** Nardiaflagelliformis Inoue, J. Jap. Bot. 46 (1): 1, 1971 (Inoue 1971b).

*** Nardiageoscyphus (De Not.) Lindb., Not. Sällsk. Fauna Fl. Fenn. Förh. 13: 371, 1874 (Lindberg 1874a). Bas.: Aliculariageoscyphus De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 18: 486, 1859 (De Notaris 1859).

* Nardiageoscyphusvar.dioica Bakalin, Arctoa 18: 87, 2009 [2010] (Bakalin et al. 2009b).

* Nardiageoscyphusvar.suberecta (Lindb. ex Kaal.) Váňa, Phytotaxa 76 (3): 37, 2013 (Váňa et al. 2013d). Bas.: Nardiahaematostictavar.suberecta Kaal., Nyt Mag. Naturvidensk. 33 (4/5): 395, 1893 (Kaalaas 1893b).

*** Nardiagrollei Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 491, 2009 (Váňa and Long 2009).

*** Nardiainsecta Lindb., Helsingf. Dagbl. 1878 (315, 18 Nov.): 2, 1878 (Lindberg 1878).

*** Nardiajaponica Steph., Bull. Herb. Boissier 5 (2): 101, 1897 (Stephani 1897b).

* Nardiakamtschatica Arnell et C.E.O.Jensen, Hedwigia 67 (1/2): 111, 1927 (Arnell 1927). 73

* Nardialeptocaulis C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (Gao and Chang 1981).

*** Nardialescurii (Austin) Underw., Bull. Illinois State Lab. Nat. Hist. 2 (1): 115, 1884 (Underwood 1884). Bas.: Alicularialescurii Austin, Hepat. bor.-amer.: 4, 1873 (Austin 1873).

** Nardiaminutifolia Furuki, Bryol. Res. 9 (3): 73, 2006 (Furuki 2006b).

** Nardianuda (Lindenb. et Gottsche) Váňa, Folia Geobot. Phytotax. 8 (2): 193, 1973 (Váňa 1973b). Bas.: Jungermannianuda Lindenb. et Gottsche, Syn. Hepat. 5: 668, 1847 (Gottsche et al. 1847).

*** Nardiapoeltii Váňa, J. Hattori Bot. Lab. 36: 73, 1972 [1973] (Váňa 1972b).

*** Nardiascalaris Gray, Nat. Arr. Brit. Pl. 1: 694, 1821 (Gray 1821).

* Nardiascalarisvar.botryoidea (R.M.Schust.) Váňa, Phytotaxa 76 (3): 38, 2013 (Váňa et al. 2013d). Bas.: Nardiascalarissubsp.botryoidea R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 862, 1969 (Schuster 1969b).

* Nardiascalarisvar.harae (Amakawa) Váňa, Phytotaxa 76 (3): 38, 2013 (Váňa et al. 2013d). Bas.: Nardiaharae Amakawa, J. Jap. Bot. 32 (2): 38, 1957 (Amakawa 1957c).

*** Nardiasubclavata (Steph.) Amakawa, J. Jap. Bot. 32 (2): 40, 1957 (Amakawa 1957c). Bas.: Jungermanniasubclavata Steph., Sp. Hepat. (Stephani) 6: 93, 1917 (Stephani 1917a).

*** Nardiasucculenta (A.Rich.) Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 519, 1885 (Spruce 1885). Bas.: Jungermanniasucculenta A.Rich., Nov. Stirp. Pug. 4: 43, 1832 (Lehmann 1832).

*** Nardiaunispiralis Amakawa, J. Jap. Bot. 32 (6): 167, 1957 (Amakawa 1957a).

** Gyrothyraceae R.M.Schust

Placement of Gyrothyraceae in Jungermanniinae follows Shaw et al. (2015).

** Gyrothyra M.Howe, Bull. Torrey Bot. Club 24 (4): 201, 1897 (Howe 1897a).

*** Gyrothyraunderwoodiana M.Howe, Bull. Torrey Bot. Club 24 (4): 202, 1897 (Howe 1897a).

*** Harpanthaceae Arnell

Davis (2004) and Hentschel et al. (2006) resolved Harpanthus as an independent lineage which is recognized here as a monogeneric family as originally construed by Arnell (1928).

*** Harpanthus Nees, Naturgesch. Eur. Leberm. 2: 351, 1836 (Nees 1836).

*** Harpanthusdrummondii (Taylor) Grolle, Österr. Bot. Z. 112 (3): 274, 1965 (Grolle 1965g). Bas.: Chiloscyphusdrummondii Taylor, London J. Bot. 5: 283, 1846 (Taylor 1846a).

*** Harpanthusflotovianus (Nees) Nees, Naturgesch. Eur. Leberm. 2: 353, 1836 (Nees 1836). Bas.: Jungermanniaflotoviana Nees, Flora 16 (26): 408, 1833 (Nees 1833b).

*** Harpanthusscutatus (F.Weber et D.Mohr) Spruce, Trans. Bot. Soc. Edinburgh 3 (1/4): 209, 1850 (Spruce 1850). Bas.: Jungermanniascutata F.Weber et D.Mohr, Bot. Taschenb. (Weber): 408, 1807 (Weber and Mohr 1807).

** Hygrobiellaceae Konstant. et Vilnet

by N. Konstantinova

The family Hygrobiellaceae was validated by Konstantinova et al. (2014a).

*** Hygrobiella Spruce, Cephalozia: 73, 1882 (Spruce 1882).

** Hygrobiellaintermedia Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (Bakalin and Vilnet 2014).

*** Hygrobiellalaxifolia (Hook.) Spruce, Cephalozia: 74, 1882 (Spruce 1882). Bas.: Jungermannialaxifolia Hook., Brit. Jungermann.: tab. 59, 1813 (Hooker 1813).

** Hygrobiellasquamosa Bakalin et Vilnet, Pl. Syst. Evol. 300 (10): 2286, 2014 (Bakalin and Vilnet 2014).

*** Jackiellaceae R.M.Schust

by J. Váňa

The monogeneric status of Jackiella is supported by a molecular study based on three loci by Hendry et al. (2007).

*** Jackiella Schiffn., Hep. Fl. Buitenzorg: 211, 1900 (Schiffner 1900a).

** Jackiellaangustifolia Herzog, Trans. Brit. Bryol. Soc. 1 (4): 296, 1950 (Herzog 1950a).

** Jackiellaceylanica Schiffn. ex Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 212 (272), 1908 (Stephani 1908h).

*** Jackiellacurvata E.A.Hodgs. et Allison, Trans. Roy. Soc. New Zealand 85 (4): 571, 1958 (Hodgson 1958).

*** Jackiellajavanica Schiffn., Hep. Fl. Buitenzorg: 212, 1900 (Schiffner 1900a).

** Jackiellajavanicavar.cavifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900 (Schiffner 1900a).

** Jackiellajavanicavar.cordifolia Schiffn., Hep. Fl. Buitenzorg: 213, 1900 (Schiffner 1900a).

** Jackiellarenifolia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (Schiffner 1900c).

** Jackiellasinensis (W.E.Nicholson) Grolle, Österr. Bot. Z. 111 (2/3): 186, 1964 (Grolle 1964f). Bas.: Aploziasinensis W.E.Nicholson, Symb. Sin. 5: 12, 1930 (Nicholson et al. 1930).

** Jackiellasingapurensis Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 70: 218, 1900 [1901] (Schiffner 1900c).

** Jackiellasingapurensisvar.philippinensis N.Kitag., Misc. Bryol. Lichenol. 9 (1): 9, 1981 (Kitagawa 1981b).

Excluded from the genus

* Jackiellaunica Steph., Sp. Hepat. (Stephani) 6: 318, 1922 (Stephani 1922). 74

*** Jungermanniaceae Rchb

by J. Váňa

The treatment of Jungermanniaceae follows Shaw et al. (2015). Many old names in Jungermannia are still neither synonymized nor transferred and we do not know their value. Some of them may prove to be older names of currently accepted taxa. We list those doubtful names in a separate section below.

** Delavayelloideae Grolle

** Delavayella Steph., Hedwigia 33 (1): 4, 1894 (Stephani 1894a).

** Delavayellaserrata Steph., Mém. Soc. Nat. Sci. Nat. Math. Cherbourg 29: 211, 1894 (Stephani 1894b).

** Delavayellaserratavar.purpurea P.C.Chen, Feddes Repert. Spec. Nov. Regni Veg. 58: 38, 1955 (Chen 1955).

** Liochlaena Nees, Syn. Hepat. 2: 150, 1845 (Gottsche et al. 1845a).

*** Liochlaenalanceolata Nees, Syn. Hepat. 2: 150, 1845 (Gottsche et al. 1845a).

*** Liochlaenasubulata (A.Evans) Schljakov, Pečen. Mchi Sev. SSSR 4: 71, 1981 (Shliakov 1981). Bas.: Jungermanniasubulata A.Evans, Trans. Connecticut Acad. Arts 8 (15): 258, 1891 (Evans 1891).

** Jungermannioideae Dumort

** Eremonotus Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 200, 1900 (Pearson 1900).

*** Eremonotusmyriocarpus (Carrington) Lindb. et Kaal. ex Pearson, Hepat. Br. Isl. 1 (6-15): 201, 1900 (Pearson 1900). Bas.: Jungermanniamyriocarpa Carrington, Hepat. Brit. Exsicc. Fasc. II: no. 96, 1879 (Carrington and Pearson 1879).

*** Jungermannia L., Sp. Pl. 1: 1131, 1753 (Linnaeus 1753). 75

*** Jungermanniaatrovirens Dumort., Syll. Jungerm. Europ.: 51, 1831 (Dumortier 1831).

*** Jungermanniaborealis Damsh. et Váňa, Lindbergia 4 (1/2): 5, 1977 (Damsholt and Váňa 1977).

* Jungermanniaerectii Ajit P.Singh et V.Nath, Hepat. Khasi Jaintia Hills: E. Himal.: 117, 2007 (Singh and Nath 2007b). 76

*** Jungermanniaexsertifolia Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (Stephani 1917a).

** Jungermanniaexsertifoliasubsp.cordifolia (Dumort.) Váňa, Folia Geobot. Phytotax. 8 (3): 268, 1973 (Váňa 1973c). Bas.: Aploziacordifolia Dumort., Bull. Soc. Roy. Bot. Belgique 13: 59, 1874 (Dumortier 1874).

*** Jungermanniagollanii Steph., Sp. Hepat. (Stephani) 6: 86, 1917 (Stephani 1917a).

** Jungermanniakonstantinovae Bakalin et Vilnet, Arctoa 18: 161, 2009 [2010] (Bakalin and Vilnet 2009).

*** Jungermanniaovatotrigona (Steph.) Grolle, Feddes Repert. 82 (1): 90, 1971 (Grolle 1971b). Bas.: Jamesoniellaovatotrigona Steph., Biblioth. Bot. 87 (2): 184, 1916 (Stephani 1916a).

*** Jungermanniapolaris Lindb., Öfvers. Kongl. Vetensk.-Akad. Förh. 23 (10): 560, 1866 [1867] (Lindberg 1866).

*** Jungermanniapumila With., Arr. Brit. Pl., ed. 3, 3: 883, 1796 (Withering 1796).

Taxa doubtfully belonging to the genus 77

* Jungermanniaamentacea Bertol., Mem. Reale Accad. Sci. Ist. Bologna (ser. 2) 1: 19, 1862 (Bertoloni 1862).

* Jungermanniabrasiliensis Raddi, Critt. Brasil.: 15, 1822 (Raddi 1822). 78

* Jungermanniachinensis Osbeck, Dagb. Ostind. Resa: 221, 1757 (Osbeck 1757). 79

* Jungermanniacordata Vill., Hist. Pl. Dauphiné (Villars) 3: 923, 1789 (Villars 1789).

* Jungermanniacrenulata Schmidel, Jungerm. Char.: 20, 1760 (Schmidel 1760).

* Jungermanniacreutzeri Kremer, Monogr. hépat. Moselle: 26, 1837 (Krémer 1837).

* Jungermanniadigitata C.F.W.Meissn. ex Spreng. Syst. Veg. (ed. 16) [Sprengel] 4 (2): 326, 1827 (Sprengel 1827b).

* Jungermanniadubioides H.A.Mill., Phytologia 47 (4): 322, 1981 (Miller 1981). Nom. nov. pro Jungermannia dubia Nees, Prodr. Fl. Norfolk.: 5, 1833 (Endlicher 1833), nom. illeg.

* Jungermanniafernandeziana Mitt., Rep. Challenger, Bot. 1 (3, 1): 85, 1884 (Mitten 1884b).

* Jungermanniahexagona Schwägr., Hist. Musc. Hepat. Prodr.: 18, 1814 (Schwägrichen 1814).

* Jungermanniaholandriana Kremer, Monogr. hépat. Moselle: 25, 1837 (Krémer 1837).

* Jungermanniaincerta Gottsche, Abh. Naturwiss. Vereins Bremen 7: 344, 1882 (Gottsche 1882). 80

* Jungermannialateriflora Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (Gottsche 1863).

* Jungermannialescuriana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 67, 1866 (Peck 1866). 81

* Jungermannialongiretis Besch. et Spruce, Bull. Soc. Bot. France (Congr. Bot.) 36: clxxxv, 1889 [1890] (Bescherelle and Spruce 1889). 82

* Jungermanniamastigophora Spreng. Neue Entdeck. Pflanzenk. 2: 99, 1821 (Sprengel 1821). 83

* Jungermanniamichelii Mérat, Nouv. fl. env. Paris (ed. 2) 1: 219, 1821 (Mérat 1821). 84

* Jungermanniaminima Scop., Fl. Carniol. (ed. 2) 2: 350, 1772 (Scopoli 1772).

* Jungermanniaodorata With., Bot. arr. veg. Gr. Brit. 2: 693, 1776 (Withering 1776).

* Jungermanniapeltata Schmidel, Jungerm. Char.: 14, 1760 (Schmidel 1760).

* Jungermanniaquadridigitata Griff., Not. pl. asiat. 2: 314, 1849 (Griffith 1849). 85

* Jungermanniasauteri De Not. ex Rabenh., Hedwigia 1 (20): 121, 1857 (Rabenhorst 1857).

* Jungermanniasecunda Hampe ex Gottsche, Mexik. Leverm.: 82, 1863 (Gottsche 1863).

* Jungermanniastereocaulis Bory, Voy. Uranie, Bot. 4: 130, 1827 (Gaudichaud 1827). 86

* Jungermanniasubmersa Kremer, Monogr. hépat. Moselle: 36, 1837 (Krémer 1837).

* Jungermanniasullivantiana Austin, Rep. (Annual) Regents Univ. State New York State Cab. Nat. Hist. 19: 66, 1866 (Peck 1866). 87

* Jungermanniasupina Hoffm., Deutschl. Fl., Theil 2 (Hoffm.): 86, 1795 [1796] (Hoffmann 1795).

* Jungermanniatenuis Ehrh., Beitr. Naturk. (Ehrhart) 4: 45, 1789 (Ehrhart 1789). 88

* Jungermanniauncifolia Steph., Hedwigia 34 (2): 51, 1895 (Stephani 1895c).

* Jungermanniavernicosa Cass. ex Mérat, Nouv. fl. env. Paris (ed. 2) 1: 221, 1821 (Mérat 1821).

** Mesoptychioideae R.M.Schust

*** Mesoptychia (Lindb.) A.Evans, Ottawa Naturalist 17: 15, 1903 (Evans 1903a). Bas.: Jungermanniasect.Mesoptychia Lindb., Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 39, 1889 (Lindberg and Arnell 1889).

*** Mesoptychiabadensis (Gottsche ex Rabenh.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (Váňa et al. 2012b). Bas.: Jungermanniabadensis Gottsche ex Rabenh., Hepat. Eur., Leberm. 9-10: no. 95, 1859 (Rabenhorst 1859).

*** Mesoptychiabantriensis (Hook.) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (Váňa et al. 2012b). Bas.: Jungermanniabantriensis Hook., Brit. Jungermann.: tab. 41, 1813 (Hooker 1813).

** Mesoptychiabantriensissubsp.wallfischii (Ştefănuţ) L.Söderstr. et Váňa, Phytotaxa 65: 52, 2012 (Váňa et al. 2012b). Bas.: Leiocoleabantriensissubsp.wallfischii Ştefănuţ, Hornwort Liverwort Romania: 21, 2008 (Ştefănuţ 2008).

* Mesoptychiachichibuensis (Inoue) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Lophoziachichibuensis Inoue, J. Jap. Bot. 36 (2): 41, 1961 (Inoue 1961b).

* Mesoptychiacollaris (Nees) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Jungermanniacollaris Nees, Fl. crypt. erlang.: xv, 1817 (Martius 1817).

* Mesoptychiafitzgeraldiae (Paton et A.R.Perry) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Leiocoleafitzgeraldiae Paton et A.R.Perry, J. Bryol. 18 (3): 470, 1995 (Paton and Perry 1995).

*** Mesoptychiagillmanii (Austin) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Jungermanniagillmanii Austin, Bull. Torrey Bot. Club 3 (3): 12, 1872 (Austin 1872).

*** Mesoptychiaheterocolpos (Thed. ex Hartm.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Jungermanniaheterocolpos Thed. ex Hartm., Handb. Skand. fl. (ed. 3): 328, 1838 (Hartman 1838).

** Mesoptychiaheterocolposvar.arctica (S.W.Arnell) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Leiocoleaarctica S.W.Arnell, Svensk Bot. Tidskr. 44 (2): 374, 1950 (Arnell 1950).

** Mesoptychiaheterocolposvar.harpanthoides (Bryhn et Kaal.) L.Söderstr. et Váňa, Phytotaxa 65: 53, 2012 (Váňa et al. 2012b). Bas.: Lophoziaharpanthoides Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 31, 1906 (Bryhn 1906).

*** Mesoptychiaigiana (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophoziaigiana S.Hatt., J. Jap. Bot. 31 (7): 201, 1956 (Hattori 1956a).

* Mesoptychiamamatkulovii (Duda) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophoziamamatkulovii Duda, Trans. Brit. Bryol. Soc. 6 (1): 82, 1970 (Duda 1970).

*** Mesoptychiamayebarae (S.Hatt.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Cephaloziamayebarae S.Hatt., J. Hattori Bot. Lab. 3: 37, 1948 [1950] (Hattori 1948a).

*** Mesoptychiamorrisoncola (Horik.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophoziamorrisoncola Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 150, 1934 (Horikawa 1934).

* Mesoptychiapolymorpha Stotler, Crand.-Stotl. et Bakalin, Polish Bot. J. 58 (1): 82, 2013 (Crandall-Stotler et al. 2013).

*** Mesoptychiarutheana (Limpr.) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Jungermanniarutheana Limpr., Jahresber. Schles. Ges. Vaterl. Cult. 61: 207, 1884 (Limpricht 1884).

** Mesoptychiarutheanavar.laxa (Schiffn. ex Burrell) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophoziaschultziivar.laxa Schiffn. ex Burrell, J. Bot. 49: 217, 1911 (Burrell 1911).

*** Mesoptychiasahlbergii (Lindb. et Arnell) A.Evans, Ottawa Naturalist 17: 15, 1903 (Evans 1903a). Bas.: Jungermanniasahlbergii Lindb. et Arnell, Kongl. Svenska Vetensk.-Akad. Handl. (n.ser.) 23 (5): 40, 1889 (Lindberg and Arnell 1889).

*** Mesoptychiasubcrispa (Herzog) L.Söderstr. et Váňa, Phytotaxa 65: 54, 2012 (Váňa et al. 2012b). Bas.: Lophoziasubcrispa Herzog, Ann. Naturhist. Mus. Wien 53 (1): 362, 1942 [1943] (Herzog 1942b).

*** Mesoptychiaturbinata (Raddi) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (Váňa et al. 2012b). Bas.: Jungermanniaturbinata Raddi, Jungermanniogr. Etrusca: 18, 1818 (Raddi 1818a).

* Mesoptychiaussuriensis (Bakalin) L.Söderstr. et Váňa, Phytotaxa 65: 55, 2012 (Váňa et al. 2012b). Bas.: Leiocoleaussuriensis Bakalin, Arctoa 17: 103, 2008 [2009] (Bakalin 2008c).

** Rivulariella D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (Wagner 2013).

** Rivulariellagemmipara (A.Evans) D.H.Wagner, Phytoneuron 2013 (10): 2, 2013 (Wagner 2013). Bas.: Chiloscyphusgemmiparus A.Evans, Bryologist 41 (3): 50, 1938 (Evans 1938b).

** Notoscyphaceae Crand.-Stotl., Váňa et Stotler

Shaw et al. (2015) circumscribed Notoscyphaceae as monogeneric based on molecular and morphological evidence.

*** Notoscyphus Mitt., Fl. vit.: 407, 1871 [1873] (Mitten 1871).

*** Notoscyphuslutescens (Lehm. et Lindenb.) Mitt., Fl. vit.: 407, 1871 [1873] (Mitten 1871). Bas.: Jungermannialutescens Lehm. et Lindenb., Nov. Stirp. Pug. 4: 16, 1832 (Lehmann 1832).

** Saccogynaceae Heeg

Shaw et al. (2015) recognized Saccogynaceae as being monogeneric.

*** Saccogyna Dumort., Commentat. Bot. (Dumortier): 113, 1822 (Dumortier 1822) nom. conserv.

* Saccogynaligulata Steph., Bull. Herb. Boissier (sér. 2) 8 (3): 207 (267), 1908 (Stephani 1908h).

** Saccogynasubacuta Steph., Sp. Hepat. (Stephani) 6: 316, 1922 (Stephani 1922).

*** Saccogynaviticulosa (L.) Dumort., Syll. Jungerm. Europ.: 74, 1831 (Dumortier 1831). Bas.: Jungermanniaviticulosa L., Sp. Pl. 1: 1131, 1753 (Linnaeus 1753).

Excluded from the genus

* Saccogynatridens Steph., Sp. Hepat. (Stephani) 6: 317, 1922 (Stephani 1922). 89

*** Solenostomataceae Stotler et Crand.-Stotl

by J. Váňa

Solenostomataceae was first defined by Crandall-Stotler et al. (2009) and refined to the current circumscription in Shaw et al (2015). The placement of Aponardia is provisional following Váňa et al. (2012c) and the placement of Arctoscyphus, Cryptocolea and Diplocolea in the family is also provisional. Taxonomic and nomenclatural notes can also be found in Váňa and Long (2009) and Váňa et al. (2012d, 2013c).

** Aponardia (R.M.Schust.) Váňa, Phytotaxa 65: 46, 2012 (Váňa et al. 2012c). Bas.: Nardiasubg.Aponardia R.M.Schust., Beih. Nova Hedwigia 119: 360, 2002 (Schuster 2002b).

*** Aponardiahuerlimannii (Váňa et Grolle) Váňa, Phytotaxa 65: 46, 2012 (Váňa et al. 2012c). Bas.: Nardiahuerlimannii Váňa et Grolle, Österr. Bot. Z. 118 (3): 233, 1970 (Váňa 1970c).

** Arctoscyphus Hässel, Lindbergia 16 (4): 133, 1990 [1992] (Hässel 1990a).

** Arctoscyphusfuegiensis (C.Massal.) Hässel, Cryptog. Bryol. Lichénol. 17 (3): 164, 1996 (Hässel 1996). Bas.: Leioscyphusrepensvar.fuegiensis C.Massal., Nuovo Giorn. Bot. Ital. 17 (3): 212, 1885 (Massalongo 1885).

** Arctoscyphusronsmithii Hässel, Lindbergia 16 (4): 133, 1990 [1992] (Hässel 1990a).

** Cryptocolea R.M.Schust., Amer. Midl. Naturalist 49 (2): 414, 1953 (Schuster 1953).

*** Cryptocoleaimbricata R.M.Schust., Amer. Midl. Naturalist 49 (2): 417, 1953 (Schuster 1953).

** Diplocolea Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (Amakawa 1962).

*** Diplocoleasikkimensis Amakawa, J. Jap. Bot. 37 (9): 274, 1962 (Amakawa 1962).

*** Solenostoma Mitt., J. Proc. Linn. Soc., Bot. 8 (29): 51, 1864 [1865] (Mitten 1864c) nom. conserv. 90

** subg.Eucalyx (Lindb.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (Shaw et al. 2015). Bas.: Nardiasect.Eucalyx Lindb., Acta Soc. Sci. Fenn. 10: 525, 1875 (Lindberg 1875).

*** Solenostomabilobum (S.Hatt. ex Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 286, 2009 (Potemkin and Sofronova 2009). Bas.: Plectocoleabiloba S.Hatt. ex Amakawa, J. Jap. Bot. 32 (7): 216, 1957 (Amakawa 1957d).

*** Solenostomaemarginatum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Plectocoleaemarginata Amakawa, J. Jap. Bot. 33 (11): 340, 1958 (Amakawa 1958b).

*** Solenostomaflagellatum (S.Hatt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Plectocoleaflagellata S.Hatt., J. Hattori Bot. Lab. 3: 13, 1948 [1950] (Hattori 1948b).

*** Solenostomahokkaidense (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermanniahokkaidensis Váňa, J. Hattori Bot. Lab. 35: 314, 1972 (Váňa 1972a).

*** Solenostomaobovatum (Nees) C.Massal., Epat. erb. critt. ital.: 17, 1903 (Massalongo 1903). Bas.: Jungermanniaobovata Nees, Naturgesch. Eur. Leberm. 1: 332, 1833 (Nees 1833c).

*** Solenostomaobscurum (A.Evans) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1013, 1969 (Schuster 1969b). Bas.: Nardiaobscura A.Evans, Rhodora 21 (249): 159, 1919 (Evans 1919a).

*** Solenostomaschusteranum (J.D.Godfrey et G.Godfrey) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermanniaschusterana J.D.Godfrey et G.Godfrey, J. Hattori Bot. Lab. 46: 109, 1979 (Godfrey and Godfrey 1979).

*** Solenostomasubtilissimum (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (Schuster 1969b). Bas.: Nardiasubtilissima Schiffn., Ann. K. K. Naturhist. Hofmus. 23: 136, 1909 (Schiffner 1909b).

** subg.Metasolenostoma Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 38, 2015 (Shaw et al. 2015).

*** Solenostomafusiforme (Steph.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 944, 1969 (Schuster 1969b). Bas.: Nardiafusiformis Steph., Bull. Herb. Boissier 5 (2): 99, 1897 (Stephani 1897b).

*** Solenostomagracillimum (Sm.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 972, 1969 (Schuster 1969b). Bas.: Jungermanniagracillima Sm., Engl. Bot. 32: tab. 2238, 1811 (Smith and Sowerby 1811).

*** Solenostomahandelii (Schiffn.) Müll.Frib., Beitr. Kryptogamenfl. Schweiz 10 (2): 38, 1947 (Müller 1947). Bas.: Nardiahandelii Schiffn., Ann. K. K. Naturhist. Hofmus. 23: 135, 1909 (Schiffner 1909b).

** Solenostomalignicola (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Nardialignicola Schiffn., Ann. K. K. Naturhist. Hofmus. 23: 137, 1909 (Schiffner 1909b).

*** Solenostomalimbatifolium (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (Váňa and Long 2009). Bas.: Jungermannialimbatifolia Amakawa, J. Hattori Bot. Lab. 31: 112, 1968 (Amakawa 1968b).

** Solenostomaphilippinense Váňa, Phytotaxa 152 (1): 43, 2013 (Váňa et al. 2013c). Based on: Solenostomagracillimumsubsp.camiguinense Bakalin, Polish Bot. J. 58 (1): 134, 2013 (Bakalin 2013).

*** Solenostomarubrum (Gottsche) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 975, 1969 (Schuster 1969b). Bas.: Jungermanniarubra Gottsche, Bot. Gaz. 13 (5): 113, 1888 (Underwood 1888).

*** Solenostomasuborbiculatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermanniasuborbiculata Amakawa, J. Hattori Bot. Lab. 31: 112, 1968 (Amakawa 1968b).

** subg.Plectocolea Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 (Mitten 1864a).

*** Solenostomabalfourii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermanniabalfourii Váňa, Folia Geobot. Phytotax. 9 (3): 279, 1974 (Váňa 1974a).

*** Solenostomaborneense (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermanniaborneensis Amakawa, J. Hattori Bot. Lab. 33: 160, 1970 (Amakawa 1970).

*** Solenostomacallithrix (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 486 (48), 1901 (Stephani 1901a). Bas.: Jungermanniacallithrix Lindenb. et Gottsche, Syn. Hepat. 5: 673, 1847 (Gottsche et al. 1847).

*** Solenostomachampawatense (S.N.Srivast. et Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermanniachampawatensis S.N.Srivast. et Amakawa, Proc. Natl. Acad. Sci. India, B 61 (2): 205, 1991 (Srivastava and Amakawa 1991).

*** Solenostomacomatum (Nees) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 73, 1981 (Gao and Chang 1981). Bas.: Jungermanniacomata Nees, Enum. Pl. Crypt. Javae: 78, 1830 (Nees 1830).

*** Solenostomacomatumvar.novae-guineae (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermanniacomatavar.novae-guineae Váňa, J. Hattori Bot. Lab. 37: 187, 1973 (Váňa 1973a).

*** Solenostomacrenuliforme (Austin) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (Stephani 1901a). Bas.: Jungermanniacrenuliformis Austin, Bull. Torrey Bot. Club 3 (3): 10, 1872 (Austin 1872).

*** Solenostomadecolor (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermanniadecolor Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 10, 1890 (Schiffner 1890).

*** Solenostomadusenii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Nardiadusenii Steph., Hedwigia 30 (5): 209, 1891 (Stephani 1891a).

*** Solenostomaerectum (Amakawa) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 66, 1981 (Gao and Chang 1981). Bas.: Plectocoleaerecta Amakawa, J. Jap. Bot. 32 (10): 307, 1957 (Amakawa 1957b).

* Solenostomaflagellalioides C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 205, 1981 (Gao and Chang 1981).

*** Solenostomaflavialbicans (Amakawa et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermanniaflavialbicans Amakawa et Grolle, J. Hattori Bot. Lab. 31: 108, 1968 (Amakawa 1968b).

*** Solenostomafossombronioides (Austin) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 1027, 1969 (Schuster 1969b). Bas.: Jungermanniafossombronioides Austin, Proc. Acad. Nat. Sci. Philadelphia 21: 220, 1869 (Austin 1869).

*** Solenostomaglaucum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (Váňa and Long 2009). Bas.: Jungermanniaglauca Amakawa, Fl. E. Himalaya: 511, 1966 (Hattori 1966c).

* Solenostomagongshanense (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (Váňa and Long 2009). Bas.: Jungermanniagongshanensis C.Gao et J.Sun, Bull. Bot. Res., Harbin 27 (2): 140, 2007 (Sun and Duan 2007).

*** Solenostomahaskarlianum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 502, 2009 (Váňa and Long 2009). Bas.: Aliculariahaskarliana Nees, Syn. Hepat. 1: 12, 1844 (Gottsche et al. 1844).

*** Solenostomahattorianum (Amakawa) Potemkin et Nyushko, Liverworts and Hornworts of Russia 1: 287, 2009 (Potemkin and Sofronova 2009). Bas.: Plectocoleahattoriana Amakawa, J. Jap. Bot. 33 (11): 341, 1958 (Amakawa 1958b).

*** Solenostomahirticalyx (Steph.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermanniahirticalyx Steph., Sp. Hepat. (Stephani) 6: 87, 1917 (Stephani 1917a).

*** Solenostomahorikawanum (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Plectocoleahorikawana Amakawa, J. Jap. Bot. 32 (7): 219, 1957 (Amakawa 1957d).

*** Solenostomahyalinum (Lyell) Mitt., Nat. hist. Azores: 319, 1870 (Mitten 1870). Bas.: Jungermanniahyalina Lyell, Brit. Jungermann.: tab. 63, 1814 (Hooker 1814).

*** Solenostomainfuscum (Mitt.) Hentschel, Pl. Syst. Evol. 268 (1/4): 152, 2007 (Hentschel et al. 2007a). Bas.: Plectocoleainfusca Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 196, 1891 (Mitten 1891).

*** Solenostomainfuscumvar.ovicalyx (Steph.) Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 288, 2009 (Potemkin and Sofronova 2009). Bas.: Solenostomaovicalyx Steph., Sp. Hepat. (Stephani) 6: 82, 1917 (Stephani 1917a).

** Solenostomakurilense (Bakalin) Váňa, Phytotaxa 152 (1): 40, 2013 (Váňa et al. 2013c). Bas.: Plectocoleaflagellatavar.kurilensis Bakalin, Arctoa 18: 90, 2009 [2010] (Bakalin et al. 2009b).

* Solenostomalixingjiangii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (Váňa and Long 2009). Bas.: Jungermannialixingjiangii C.Gao et X.L.Bai, Philipp. Scientist 38: 128, 2001 (Gao and Bai 2001).

*** Solenostomamarginatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (Schuster 1969b). Bas.: Plectocoleamarginata S.Hatt., J. Hattori Bot. Lab. 3: 40, 1948 [1950] (Hattori 1948a).

*** Solenostomamicranthum (Mitt.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Plectocoleamicrantha Mitt., Fl. vit.: 405, 1871 [1873] (Mitten 1871).

** Solenostomamontanum (Steph.) Váňa, Phytotaxa 65: 44, 2012 (Váňa et al. 2012d). Bas.: Nardiamontana Steph., Hedwigia 28 (3): 164, 1889 (Stephani 1889d).

* Solenostomamulticarpum (C.Gao et J.Sun) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermanniamulticarpa C.Gao et J.Sun, Bull. Bot. Res., Harbin 27 (2): 139, 2007 (Sun and Duan 2007).

** Solenostomanilgiriense (A.Alam, Ad.Kumar et S.C.Srivast.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermannianilgiriensis A.Alam, Ad.Kumar et S.C.Srivast., Bull. Bot. Surv. India 49 (1/4): 220, 2007 (Alam et al. 2007).

*** Solenostomaobliquifolium (Schiffn.) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Nardiaobliquifolia Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 191, 1898 (Schiffner 1898a).

*** Solenostomaonraedtii (Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermanniaonraedtii Váňa, Folia Geobot. Phytotax. 9 (3): 282, 1974 (Váňa 1974a).

* Solenostomaorbicularifolium (Piippo ex C.Gao et Bai) Váňa, Phytotaxa 222 (2): 199, 2015 (Söderström et al. 2015d). Bas.: Jungermanniaorbicularifolia Piippo ex C.Gao et Bai, Philipp. Scientist 38: 129, 2001 (Gao and Bai 2001).

*** Solenostomaotianum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 984, 1969 (Schuster 1969b). Bas.: Plectocoleaotiana S.Hatt., J. Jap. Bot. 28 (6): 183, 1953 (Hattori 1953b).

*** Solenostomaovalifolium (Amakawa) Váňa, Phytotaxa 152 (1): 40, 2013 (Váňa et al. 2013c). Bas.: Plectocoleainfuscavar.ovalifolia Amakawa, J. Jap. Bot. 34 (4): 115, 1959 (Amakawa 1959b).

*** Solenostomaparoicum (Schiffn.) R.M.Schust., Amer. Midl. Naturalist 49 (2): 402, 1953 (Schuster 1953). Bas.: Nardiaparoica Schiffn., Lotos 58: 320, 1910 (Schiffner 1910d).

*** Solenostomaplagiochilaceum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermanniaplagiochilacea Grolle, J. Hattori Bot. Lab. 58: 197, 1985 (Grolle 1985b).

*** Solenostomapolyrhizoides (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermanniapolyrhizoides Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 262, 1966 (Amakawa 1966).

*** Solenostomaradicellosum Mitt., J. Linn. Soc., Bot. 8 (31): 156, 1864 [1865] (Mitten 1864a).

*** Solenostomarenauldii (Steph.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermanniarenauldii Steph., Bull. Soc. Roy. Bot. Belgique, Compt. Rend. 30 (2): 201, 1891 [1892] (Stephani 1891b).

*** Solenostomarigidulum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 983, 1969 (Schuster 1969b). Bas.: Plectocolearigidula S.Hatt., J. Jap. Bot. 27 (2): 53, 1952 (Hattori 1952b).

*** Solenostomarosulans (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Bas.: Nardiarosulans Steph., Bull. Herb. Boissier 5 (2): 101, 1897 (Stephani 1897b).

*** Solenostomarotundatum Amakawa, J. Jap. Bot. 31 (2): 50, 1956 (Amakawa 1956).

*** Solenostomarubripunctatum (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 898, 1969 (Schuster 1969b). Bas.: Plectocolearubripunctata S.Hatt., J. Hattori Bot. Lab. 3: 41, 1948 [1950] (Hattori 1948a).

* Solenostomarupicola (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Bas.: Jungermanniarupicola Amakawa, J. Hattori Bot. Lab. 22: 23, 1960 (Amakawa 1960b). 91

*** Solenostomasikkimense (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermanniasikkimensis Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 92, 1917 (Stephani 1917a).

*** Solenostomatetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (Váňa and Long 2009). Bas.: Jungermanniatetragona Lindenb., Bot. Zeitung (Berlin) 6 (25): 462, 1848 (Meissner 1848).

* Solenostomatetragonumvar.kodaikanalense A.Alam, D.Sharma et So.Yadav, Phytotaxonomy 12: 70, 2012 (Alam et al. 2012).

*** Solenostomatorticalyx (Steph.) C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 69, 1981 (Gao and Chang 1981). Bas.: Jungermanniatorticalyx Steph., Sp. Hepat. (Stephani) 6: 94, 1917 (Stephani 1917a).

*** Solenostomatruncatum (Nees) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 509, 2009 (Váňa and Long 2009). Bas.: Jungermanniatruncata Nees, Enum. Pl. Crypt. Javae: 29, 1830 (Nees 1830).

** Solenostomatruncatumvar.setulosum (Herzog) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Plectocoleasetulosa Herzog, J. Hattori Bot. Lab. 14: 33, 1955 (Herzog and Noguchi 1955).

*** Solenostomatuberculiferum (Herzog) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Aploziatuberculifera Herzog, Ann. Bryol. 5: 84, 1932 (Herzog 1932a).

*** Solenostomaunispire (Amakawa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Plectocoleaunispiris Amakawa, J. Jap. Bot. 29 (6): 178, 1954 (Amakawa 1954).

*** Solenostomavirgatum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Plectocoleavirgata Mitt., Trans. Linn. Soc. London, Bot. 3 (3): 197, 1891 (Mitten 1891).

*** Solenostomavulcanicola (Schiffn.) Nyushko ex Potemkin et Sofronova, Liverworts and Hornworts of Russia 1: 289, 2009 (Potemkin and Sofronova 2009). Bas.: Nardiavulcanicola Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 191, 1898 (Schiffner 1898a).

* Solenostomazangmuii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermanniazangmuii C.Gao et X.L.Bai, Philipp. Scientist 38: 135, 2001 (Gao and Bai 2001).

** subg.Solenostoma

*** Solenostomaamoenum (Lindenb. et Gottsche) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermanniaamoena Lindenb. et Gottsche, Syn. Hepat. 5: 674, 1847 (Gottsche et al. 1847).

*** Solenostomaamplexifolium (Hampe) Váňa et Schäf.-Verw., Acta Bot. Hung. 51 (3/4): 408, 2009 (Schäfer-Verwimp and Pócs 2009). Bas.: Plagiochilaamplexifolia Hampe, Nov. Stirp. Pug. 7: 6, 1838 (Lehmann 1838).

** Solenostomaappalachianum R.M.Schust. ex Bakalin, Arctoa 23: 127, 2014 (Bakalin 2014). Based on: Solenostomaappalachianum R.M.Schust., Rhodora 60 (717): 246, 1958 (Schuster 1958a), nom. inval. 92

*** Solenostomaappressifolium (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 494, 2009 (Váňa and Long 2009). Bas.: Jungermanniaappressifolia Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c). 93

*** Solenostomaappressifoliumvar.minor (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermanniaclavellatavar.minor Amakawa, J. Hattori Bot. Lab. 26: 25, 1963 (Amakawa 1963).

*** Solenostomaappressifoliumvar.nigricans (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermanniadecolyanavar.nigricans Amakawa, J. Hattori Bot. Lab. 30: 185, 1967 (Amakawa 1967b).

*** Solenostomaariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermanniaariadne Taylor, Nov. Stirp. Pug. 8: 9, 1844 (Lehmann 1844).

*** Solenostomaatrobrunneum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 495, 2009 (Váňa and Long 2009). Bas.: Jungermanniaatrobrunnea Amakawa, J. Hattori Bot. Lab. 30: 192, 1967 (Amakawa 1967b).

*** Solenostomaatrorevolutum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermanniaatrorevoluta Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 255, 1966 (Amakawa 1966).

*** Solenostomaatrovirens Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 493 (55), 1901 (Stephani 1901a).

*** Solenostomabaueri (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 495 (57), 1901 (Stephani 1901a). Bas.: Aploziabaueri Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 195, 1898 (Schiffner 1898a).

*** Solenostomabengalense (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermanniabengalensis Amakawa, J. Hattori Bot. Lab. 31: 112, 1968 (Amakawa 1968b).

** Solenostomabreviflorum Kashyap et R.S.Chopra, Liverworts W. Himal. 2: 85, 1932 (Kashyap and Chopra 1932).

*** Solenostomacaeleste (Inoue et Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermanniacaelestis Inoue et Váňa, Stud. Cryptog. Papua N. Guinea: 16, 1979 (Inoue 1979b).

* Solenostomacaoi (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermanniacaoi C.Gao et X.L.Bai, Philipp. Scientist 38: 152, 2001 (Gao and Bai 2001).

*** Solenostomacaucasicum (Váňa) Konstant., Arctoa 1: 123, 1992 (Konstantinova et al. 1992). Bas.: Jungermanniacaucasica Váňa, Preslia 42: 96, 1970 (Váňa 1970a).

*** Solenostomachenianum (C.Gao, Y.H.Wu et Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 496, 2009 (Váňa and Long 2009). Bas.: Jungermanniacheniana C.Gao, Y.H.Wu et Grolle, Nova Hedwigia 77 (1/2): 190, 2003 (Gao et al. 2003).

*** Solenostomaclavellatum Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 491 (53), 1901 (Stephani 1901a).

*** Solenostomaconfertissimum (Nees) Schljakov, Novosti Sist. Nizš. Rast. 17: 239, 1980 (Shliakov 1980b). Bas.: Jungermanniaconfertissima Nees, Naturgesch. Eur. Leberm. 1: 291, 1833 (Nees 1833c).

*** Solenostomaconiflorum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (Stephani 1901a). Bas.: Jungermanniaconiflora Schiffn., Leberm., Forschungsr. Gazelle 4 (4): 10, 1890 (Schiffner 1890).

*** Solenostomacrassulum (Nees et Mont.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 497 (59), 1901 (Stephani 1901a). Bas.: Jungermanniacrassula Nees et Mont., Ann. Sci. Nat. Bot. (sér. 2) 5: 54, 1836 (Nees and Montagne 1836).

** Solenostomacryptogynum R.M.Schust. ex J.J.Engel, Novon 17 (3): 311, 2007 (Engel 2007). Based on: Solenostomacryptogynum R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (Schuster 2002b), nom. inval.

*** Solenostomacyclops (S.Hatt.) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (Schuster 1969b). Bas.: Jungermanniacyclops S.Hatt., J. Hattori Bot. Lab. 3: 5, 1948 [1950] (Hattori 1948b).

*** Solenostomadiversiclavellatum (Amakawa et Grolle) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermanniadiversiclavellata Amakawa et Grolle, J. Hattori Bot. Lab. 31: 107, 1968 (Amakawa 1968b).

*** Solenostomadulongense Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 497, 2009 (Váňa and Long 2009).

*** Solenostomaexsertum (A.Evans) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (Stephani 1901a). Bas.: Nardiaexserta A.Evans, Trans. Connecticut Acad. Arts 8 (15): 259, 1891 (Evans 1891).

*** Solenostomafaurieanum (Beauverd) R.M.Schust., Hepat. Anthocerotae N. Amer. 2: 945, 1969 (Schuster 1969b). Bas.: Jungermanniafaurieana Beauverd, Sp. Hepat. (Stephani) 6: 571, 1924 (Stephani 1924).

*** Solenostomaflagellare (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermanniaflagellaris Amakawa, J. Hattori Bot. Lab. 29: 258, 1966 (Amakawa 1966).

*** Solenostomaflavorevolutum (Váňa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 501, 2009 (Váňa and Long 2009). Bas.: Jungermanniaflavorevoluta Váňa, J. Hattori Bot. Lab. 36: 63, 1972 [1973] (Váňa 1972b).

*** Solenostomagrollei (D.G.Long et Váňa) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (Feldberg et al. 2009). Bas.: Gottscheliagrollei D.G.Long et Váňa, J. Bryol. 29 (3): 167, 2007 (Long and Váňa 2007).

*** Solenostomagrosseverrucosum (Amakawa et S.Hatt.) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Horikawaellagrosseverrucosa Amakawa et S.Hatt., Bull. Univ. Mus. Univ. Tokyo 8: 216, 1975 (Hattori 1975e).

*** Solenostomaheterolimbatum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermanniaheterolimbata Amakawa, J. Hattori Bot. Lab. 30: 183, 1967 (Amakawa 1967b).

*** Solenostomahewsoniae (Amakawa et Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 136, 2010 (Váňa et al. 2010a). Bas.: Jungermanniahewsoniae Amakawa et Grolle, J. Hattori Bot. Lab. 31: 108, 1968 (Amakawa 1968b).

*** Solenostomahiugaense Amakawa, J. Jap. Bot. 31 (2): 47, 1956 (Amakawa 1956).

*** Solenostomaindrodayanum (Sushil K.Singh et D.K.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermanniaindrodayana Sushil K.Singh et D.K.Singh, Cryptog. Bryol. 28 (2): 103, 2007 (Singh and Singh 2007b).

*** Solenostomainundatum (Hook.f. et Taylor) Mitt. ex Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 490 (52), 1901 (Stephani 1901a). Bas.: Jungermanniainundata Hook.f. et Taylor, London J. Bot. 3: 559, 1844 (Hooker and Taylor 1844d).

*** Solenostomajavanicum (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 494 (56), 1901 (Stephani 1901a). Bas.: Aploziajavanica Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 193, 1898 (Schiffner 1898a).

** Solenostomakanaii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermanniakanaii Amakawa, J. Hattori Bot. Lab. 30: 194, 1967 (Amakawa 1967b).

*** Solenostomakashyapii (S.C.Srivast., S.Srivast. et D.Sharma) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermanniakashyapii S.C.Srivast., S.Srivast. et D.Sharma, Lindbergia 28 (3): 131, 2003 (Srivastava et al. 2003).

*** Solenostomalanigerum (Mitt.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 503, 2009 (Váňa and Long 2009). Bas.: Jungermannialanigera Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c).

*** Solenostomamacrocarpum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 504, 2009 (Váňa and Long 2009). Bas.: Jungermanniamacrocarpa Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 87, 1917 (Stephani 1917a).

*** Solenostomamamatkulovii (Váňa et Zerov) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermanniamamatkulovii Váňa et Zerov, Preslia 49: 181, 1977 (Váňa 1977).

* Solenostomamicrophyllum C.Gao, Fl. Hepat. Chin. Boreali-Orient.: 206, 1981 (Gao and Chang 1981).

* Solenostomamicrorevolutum (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermanniamicrorevoluta C.Gao et X.L.Bai, Philipp. Scientist 38: 146, 2001 (Gao and Bai 2001).

*** Solenostomamildbraedii (Steph.) R.M.Schust., Beih. Nova Hedwigia 119: 387, 2002 (Schuster 2002b). Bas.: Jungermanniamildbraedii Steph., Wiss. Ergebn. Deut. Zentr.-Afr. Exped. (1907-08), Bot. 2: 113, 1911 (Stephani 1911a).

*** Solenostomaniveum (Grolle) R.M.Schust. ex Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 137, 2010 (Váňa et al. 2010a). Bas.: Jungermannianivea Grolle, Misc. Bryol. Lichenol. 6 (1): 1, 1971 (Grolle 1971c).

* Solenostomanovazelandiae R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (Schuster 2002b). 94

*** Solenostomaohbae (Amakawa) C.Gao, Bryofl. Xizang: 495, 1985 (Li 1985). Bas.: Jungermanniaohbae Amakawa, Bull. Univ. Mus. Univ. Tokyo 8: 218, 1975 (Hattori 1975e).

*** Solenostomaorbiculatum (Colenso) R.M.Schust., Beih. Nova Hedwigia 119: 380, 2002 (Schuster 2002b). Bas.: Gymnomitrionorbiculatum Colenso, Trans. & Proc. New Zealand Inst. 18: 236, 1886 (Colenso 1886b).

*** Solenostomaparvitextum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermanniaparvitexta Amakawa, J. Hattori Bot. Lab. 30: 187, 1967 (Amakawa 1967b).

*** Solenostomapatoniae (Grolle, D.B.Schill et D.G.Long) K.Feldberg, Hentschel, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (Feldberg et al. 2009). Bas.: Gottscheliapatoniae Grolle, D.B.Schill et D.G.Long, J. Bryol. 25 (1): 3, 2003 (Grolle et al. 2003).

*** Solenostomapocsii (Váňa) Bakalin, Arctoa 18: 160, 2009 [2010] (Bakalin and Vilnet 2009). Bas.: Jungermanniapocsii Váňa, Folia Geobot. Phytotax. 10 (4): 365, 1975 (Váňa 1975b).

*** Solenostomapoeltii (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 505, 2009 (Váňa and Long 2009). Bas.: Jungermanniapoeltii Amakawa, J. Hattori Bot. Lab. 29: 258, 1966 (Amakawa 1966).

*** Solenostomapseudocyclops (Inoue) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (Váňa and Long 2009). Bas.: Jungermanniapseudocyclops Inoue, Bull. Natl. Sci. Mus. Tokyo (n.ser.) 9 (1): 37, 1966 (Inoue 1966a).

** Solenostomapseudopyriflorum Bakalin et Vilnet, Arctoa 18: 159, 2009 [2010] (Bakalin and Vilnet 2009).

*** Solenostomapurpuratum (Mitt.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (Stephani 1901a). Bas.: Jungermanniapurpurata Mitt., J. Proc. Linn. Soc., Bot. 5 (18): 91, 1860 [1861] (Mitten 1860c).

*** Solenostomapyriflorum Steph., Sp. Hepat. (Stephani) 6: 83, 1917 (Stephani 1917a). 95

*** Solenostomapyriflorumvar.gracillimum (Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 506, 2009 (Váňa and Long 2009). Bas.: Jungermanniapyrifloravar.gracillima Amakawa, Fl. E. Himalaya 2: 228, 1971 (Hattori 1971a).

** Solenostomapyriflorumvar.major (S.Hatt.) Bakalin, Arctoa 17: 230, 2008 [2009] (Bakalin 2008a). Bas.: Jungermanniamonticolaf.major S.Hatt., J. Hattori Bot. Lab. 3: 8, 1948 [1950] (Hattori 1948b).

*** Solenostomapyriflorumvar.minutissimum (Amakawa) Bakalin, Arctoa 16: 208, 2007 [2008] (Bakalin 2007). Bas.: Jungermanniapyrifloravar.minutissima Amakawa, J. Hattori Bot. Lab. 22: 61, 1960 (Amakawa 1960b). 96

*** Solenostomaraujeanum (Grolle ex Amakawa) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Bas.: Jungermanniaraujeana Grolle ex Amakawa, J. Hattori Bot. Lab. 29: 262, 1966 (Amakawa 1966).

*** Solenostomariclefii Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 507, 2009 (Váňa and Long 2009). Nom. nov. pro Jungermannia grollei Amakawa, J. Hattori Bot. Lab. 29: 260, 1966 (Amakawa 1966).

*** Solenostomasanguinolentum (Griff.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 489 (51), 1901 (Stephani 1901a). Bas.: Jungermanniasanguinolenta Griff., Not. pl. asiat. 2: 302, 1849 (Griffith 1849).

*** Solenostomaschaulianum (Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermanniaschauliana Steph., Sp. Hepat. (Stephani) 6: 90, 1917 (Stephani 1917a).

*** Solenostomashimizuanum (S.Hatt. ex Váňa) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Jungermanniashimizuana S.Hatt. ex Váňa, J. Hattori Bot. Lab. 35: 315, 1972 (Váňa 1972a).

*** Solenostomaspeciosum (Horik.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 244, 2009 (Feldberg et al. 2009). Bas.: Anastrophyllumspeciosum Horik., J. Sci. Hiroshima Univ., Ser. B, Div. 2, Bot. 2: 147, 1934 (Horikawa 1934).

** Solenostomaspeciosumsubsp.villosum (R.M.Schust.) Hentschel, K.Feldberg, Bombosch, D.G.Long, Váňa et Heinrichs, Pl. Syst. Evol. 280 (3/4): 246, 2009 (Feldberg et al. 2009). Bas.: Scaphophyllumspeciosumsubsp.villosum R.M.Schust., Bryologist 101 (3): 434, 1998 (Schuster 1998b).

*** Solenostomasphaerocarpum (Hook.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 499 (61), 1901 (Stephani 1901a). Bas.: Jungermanniasphaerocarpa Hook., Brit. Jungermann.: tab. 74, 1815 (Hooker 1815).

*** Solenostomastephanii (Schiffn.) Steph., Bull. Herb. Boissier (sér. 2) 1 (5): 496 (58), 1901 (Stephani 1901a). Bas.: Aploziastephanii Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 195, 1898 (Schiffner 1898a).

*** Solenostomastoloniferum (Steph.) S.W.Arnell, Hepat. South Africa: 316, 1963 (Arnell 1963b). Bas.: Nardiastolonifera Steph., Hedwigia 31 (3): 128, 1892 (Stephani 1892d).

*** Solenostomastrictum (Schiffn.) Váňa, Hentschel et Heinrichs, Cryptog. Bryol. 31 (2): 138, 2010 (Váňa et al. 2010a). Bas.: Aploziastricta Schiffn., Denkschr. Kaiserl. Akad. Wiss., Math.-Naturwiss. Kl. 67: 194, 1898 (Schiffner 1898a).

*** Solenostomasubacutum (Herzog) Váňa, Crand.-Stotl. et Stotler, Syst. Bot. 40 (1): 39, 2015 (Shaw et al. 2015). Bas.: Anastrophyllumsubacutum Herzog, Ann. Bryol. 12: 75, 1939 (Herzog 1939b).

*** Solenostomasubrubrum (Schiffn. ex Steph.) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 508, 2009 (Váňa and Long 2009). Bas.: Jungermanniasubrubra Schiffn. ex Steph., Sp. Hepat. (Stephani) 6: 93, 1917 (Stephani 1917a).

* Solenostomasunii Bakalin et Vilnet, Bot. Pacifica 3 (2): 15, 2014 (Bakalin et al. 2014).

* Solenostomatotopapillosum (E.A.Hodgs.) R.M.Schust., Bryologist 100 (3): 366, 1997 (Schuster 1997a). Bas.: Jungermanniatotopapillosa E.A.Hodgs., J. Roy. Soc. New Zealand 2 (1): 111, 1972 (Hodgson 1972). 97

*** Solenostomaudarii (S.C.Srivast. et P.Singh) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermanniaudarii S.C.Srivast. et P.Singh, Recent Stud. Indian Bryoph.: 152, 1995 (Srivastava and Singh 1995).

* Solenostomaventroversum (Grolle) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermanniaventroversa Grolle, Khumbu Himal 1 (4): 284, 1966 (Grolle 1966k). 98

*** Solenostomazantenii (Amakawa) R.M.Schust. ex Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermanniazantenii Amakawa, J. Hattori Bot. Lab. 31: 110, 1968 (Amakawa 1968b).

* Solenostomazengii (C.Gao et X.L.Bai) Váňa et D.G.Long, Nova Hedwigia 89 (3/4): 510, 2009 (Váňa and Long 2009). Bas.: Jungermanniazengii C.Gao et X.L.Bai, Philipp. Scientist 38: 151, 2001 (Gao and Bai 2001).

*** Southbyaceae Váňa, Crand.-Stotl., Stotler et D.G.Long

by J. Váňa

Southbyaceae has traditionally been placed in Arnelliaceae (e.g. Crandall-Stotler et al. 2009) or kept as a separate family (e.g. Vanden Berghen 1957) that has never been validated. However, it was re-established, rearranged and validated in the study by Váňa et al. (2012e).

*** Gongylanthus Nees, Naturgesch. Eur. Leberm. 2: 405, 1836 (Nees 1836).

*** Gongylanthusdusenii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (Stephani 1906a).

*** Gongylanthusericetorum (Raddi) Nees, Naturgesch. Eur. Leberm. 2: 407, 1836 (Nees 1836). Bas.: Calypogeiaericetorum Raddi, Jungermanniogr. Etrusca: 31, 1818 (Raddi 1818a).

*** Gongylanthusgranatensis (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 385 (41), 1906 (Stephani 1906a). Bas.: Lindiginagranatensis Gottsche, Ann. Sci. Nat. Bot. (sér. 5) 1: 138, 1864 (Gottsche 1864).

*** Gongylanthushimalayensis Grolle, Khumbu Himal 1 (4): 285, 1966 (Grolle 1966k).

*** Gongylanthusjavanicus Grolle, J. Jap. Bot. 40 (7): 206, 1965 (Grolle 1965d).

*** Gongylanthusliebmanianus (Lindenb. et Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (Stephani 1906a). Bas.: Gymnantheliebmaniana Lindenb. et Gottsche, Syn. Hepat. 5: 712, 1847 (Gottsche et al. 1847).

*** Gongylanthuslimbatus (Herzog) Grolle et Váňa, Folia Geobot. Phytotax. 9 (2): 198, 1974 (Váňa 1974c). Bas.: Aplozialimbata Herzog, Beih. Bot. Centralbl. 61B (3): 561, 1942 (Herzog 1942d).

*** Gongylanthusmuelleri (Gottsche) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 388 (44), 1906 (Stephani 1906a). Bas.: Lindigiamuelleri Gottsche, Mexik. Leverm.: 121, 1863 (Gottsche 1863).

** Gongylanthusoniscoides (Spruce) Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 386 (42), 1906 (Stephani 1906a). Bas.: Calypogeiaoniscoides Spruce, Trans. & Proc. Bot. Soc. Edinburgh 15: 448, 1885 (Spruce 1885).

*** Gongylanthusrichardsii E.W.Jones, Trans. Brit. Bryol. Soc. 4 (4): 650, 1964 (Jones 1964).

*** Southbya Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (Spruce 1849).

*** Southbyagollanii Steph., Bull. Herb. Boissier (sér. 2) 6 (5): 381 (37), 1906 (Stephani 1906a).

*** Southbyanigrella (De Not.) Henriq., Bol. Soc. Brot. 4: 244, 1886 [1887] (Henriques 1886). Bas.: Jungermannianigrella De Not., Mem. Reale Accad. Sci. Torino (ser. 2) 1: 315, 1838 (De Notaris 1838).

*** Southbyaorganensis Herzog, Memoranda Soc. Fauna Fl. Fennica 25: 53, 1950 (Herzog 1950c).

*** Southbyatophacea (Spruce) Spruce, Ann. Mag. Nat. Hist. (ser. 2) 3 (18): 501, 1849 (Spruce 1849). Bas.: Jungermanniatophacea Spruce, Hep. Pyr. Exsic.: no. 23, 1847 (Spruce 1847).

** Stephaniellaceae R.M.Schust

by J. Váňa

In the absence of molecular data, the phylogenetic affinities of Stephaniella and Stephaniellidium remain equivocal, but there is no evidence to support their placement in either the Arnelliaceae or Southbyaceae where they have been placed recently. Thus the family as construed by Schuster (2002b) is retained.

*** Stephaniella J.B.Jack, Hedwigia 33 (1): 11, 1894 (Jack 1894).

* Stephaniellaboliviensis Steph., Biblioth. Bot. 87 (2): 182, 1916 (Stephani 1916a).

*** Stephaniellahamata Steph., Bull. Herb. Boissier (sér. 2) 1 (10): 1024 (87), 1901 (Stephani 1901c).

*** Stephaniellaparaphyllina J.B.Jack, Hedwigia 33 (1): 11, 1894 (Jack 1894).

*** Stephaniellarostrata U.Schmitt, Österr. Bot. Z. 115 (2): 124, 1968 (Schmitt and Winkler 1968).

*** Stephaniellauncifolia S.Winkl., Österr. Bot. Z. 115 (2): 124, 1968 (Schmitt and Winkler 1968).

** Stephaniellidium S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (Grolle 1983b). Based on: Stephaniellidium S.Winkl., Mitt. Inst. Colombo-Alemán Invest. Ci. 3: 60, 1969 (Winkler 1969).

*** Stephaniellidiumsleumeri (Müll.Frib.) S.Winkl. ex Grolle, Acta Bot. Fenn. 121: 38, 1983 (Grolle 1983b). Bas.: Stephaniellasleumeri Müll.Frib., Rev. Bryol. Lichénol. 20 (1/2): 177, 1951 (Müller 1951b).

*** Trichotemnomataceae R.M.Schust

*** Trichotemnoma R.M.Schust., Nova Hedwigia 15: 440, 1968 (Schuster 1968b).

*** Trichotemnomacorrugatum (Steph.) R.M.Schust., Nova Hedwigia 15: 440, 1968 (Schuster 1968b). Bas.: Blepharostomacorrugatum Steph., Hedwigia 32 (5): 315, 1893 (Stephani 1893d).

Lophocoleineae Schljakov
*** Blepharostomataceae W.Frey et M.Stech

The recognition of Blepharostomataceae follows Frey and Stech (2008) following reconsideration of molecular and morphological data available at that time.

*** Blepharostoma (Dumort.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermanniasect.Blepharostoma Dumort., Syll. Jungerm. Europ.: 65, 1831 (Dumortier 1831).

** Blepharostomaarachnoideum M.Howe, Mem. Torrey Bot. Club 7: 140, 1899 (Howe 1899).

** Blepharostomaindicum G.Asthana, M.Saxena et Maurya, J. Bryol. 35 (4): 267, 2013 (Asthana et al. 2013).

** Blepharostomaminor Horik., Hikobia 1 (2): 104, 1951 [1952] (Horikawa 1951b).

*** Blepharostomatrichophyllum (L.) Dumort., Recueil Observ. Jungerm.: 18, 1835 (Dumortier 1835). Bas.: Jungermanniatrichophylla L., Sp. Pl. 1: 1135, 1753 (Linnaeus 1753).

** Blepharostomatrichophyllumsubsp.brevirete (Bryhn et Kaal.) R.M.Schust., Bull. Natl. Mus. Canada 164: 16, 1959 (Schuster et al. 1959). Bas.: Blepharostomatrichophyllumvar.brevirete Bryhn et Kaal., Rep. Second Norweg. Arctic Exped. 11: 46, 1906 (Bryhn 1906).

*** Brevianthaceae J.J.Engel et R.M.Schust

by L. Söderström, J. Váňa, R. Stotler, B.J. Crandall-Stotler and J.J. Engel

The treatment of Brevianthaceae follows Söderström et al. (2013b).

*** Brevianthus J.J.Engel et R.M.Schust., Phytologia 47 (4): 317, 1981 (Engel and Schuster 1981).

*** Brevianthusflavus (Grolle) J.J.Engel et R.M.Schust., Phytologia 47 (4): 318, 1981 (Engel and Schuster 1981). Bas.: Jackiellaflava Grolle, J. Hattori Bot. Lab. 33: 222, 1970 (Grolle 1970b).

** Brevianthusflavussubsp.crenulatus J.J.Engel, Nova Hedwigia 93 (3/4): 406, 2011 (Engel 2011).

* Brevianthushypocanthidium M.A.M.Renner et J.J.Engel, PhytoKeys 50: 46, 2015 (Renner et al. 2015).

*** Tetracymbaliella Grolle, Nova Hedwigia 3 (1): 48, 1961 (Grolle 1961b).

** Tetracymbaliellacomptonii (Pearson) Grolle, Rev. Bryol. Lichénol. 32 (3/4): 164, 1963 [1964] (Grolle 1963d). Bas.: Chiloscyphuscomptonii Pearson, J. Linn. Soc., Bot. 46 (305): 23, 1922 (Pearson 1922b).

*** Tetracymbaliellacymbalifera (Hook.f. et Taylor) Grolle, Nova Hedwigia 3 (1): 50, 1961 (Grolle 1961b). Bas.: Jungermanniacymbalifera Hook.f. et Taylor, Bot. Antarct. Voy. I (Fl. Antarct. 1): 151, 1845 (Taylor and Hooker 1845).

*** Tetracymbalielladecipiens (Gottsche) Grolle, Nova Hedwigia 3 (1): 49, 1961 (Grolle 1961b). Bas.: Chiloscyphusdecipiens Gottsche, Syn. Hepat. 2: 176, 1845 (Gottsche et al. 1845a).

*** Tetracymbaliellasubsimplex (Austin) J.J.Engel, Phytotaxa 207 (2): 185, 2015 (Engel 2015b). Bas.: Polyotussubsimplex Austin, Bull. Torrey Bot. Club 6 (7): 46, 1875 (Austin 1875c).

** Chonecoleaceae R.M.Schust. ex Grolle

** Chonecolea Grolle, Rev. Bryol. Lichénol. 25 (3/4): 294, 1956 [1957] (Grolle 1956).

** Chonecoleaacutiloba (Schiffn.) R.M.Schust., Hepat. Anthocerotae N. Amer. 4: 321, 1980 (Schuster 1980c). Bas.: Clasmatocoleaacutiloba Schiffn., Österr. Akad. Wiss., Math.-Naturwiss. Kl., Denkschr. 111: 63, 1964 (Schiffner and Arnell 1964</