Checklist |
Corresponding author: Ainun Nadhifah ( ainun.nadhifah@brin.go.id ) Academic editor: Peter de Lange
© 2024 Ainun Nadhifah, Ida Haerida, Fandri Sofiana Fastanti, Lars Söderström, Anders Hagborg, Matt von Konrat.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Nadhifah A, Haerida I, Fastanti FS, Söderström L, Hagborg A, von Konrat M (2024) Beyond nutmeg, mace, and cloves: Checklist of the liverworts and hornworts of Maluku Islands (Moluccas), Indonesia. PhytoKeys 239: 107-193. https://doi.org/10.3897/phytokeys.239.116679
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The first ever liverwort and hornwort checklist is provided for the Maluku Islands (Moluccas/Spice Islands) of Indonesia. We report 355 accepted and 16 doubtful species and reject 22 species previously reported for Maluku Islands. The list is based on the specimens housed in the Herbarium Bogoriense (BO) and reports from over 500 literature references, including monographs, regional studies, and molecular investigations. The Maluku Islands are part of the Wallacea Biodiversity Hotspot with many unique species found only in Wallacea. Publications focusing on liverworts and hornworts of Maluku Islands are few and scattered. Considering regionally widespread species that have been recorded elsewhere, we predict that further fieldwork exploring the diversity of habitats coupled with collections unveiled from regional herbaria, a number of new records remain to be reported.
Checklist, hornworts, Indonesia, liverworts, Maluku, Moluccas
The Maluku Islands (Moluccas) is an island archipelago of over 1,000 islands situated in eastern Indonesia south of the Philippine island of Mindanao and east of Sulawesi, scattered in a series of irregular arcs spanning the Maluku, the Banda, and the Seram Seas (
The biogeographical region of Maluku Islands (Fig.
Famously, western exploration of the natural history of the Maluku Islands dates back to the 17th century, starting with Georg Everhard Rumphius, and later, for example, Alfred Russel Wallace (
Maluku Islands are situated in the “Tropical and Subtropical Moist Broadleaved Forests” biome (
Liverworts and hornworts are, together with mosses, commonly referred to as bryophytes. They represent three early diverging clades of land plants forming the second largest group of green land plants after flowering plants and are of critical biological, ecological, and phylogenetic significance. Bryophytes serve as the “macrophytes,” providing a matrix where many microscopic organisms live, including tardigrades, mites, rotifers, micro-molluscs, microalgae, microfungi, and prokaryotes (
Historically, relatively few publications have focused on plant collections from the Maluku Islands, despite the significance of the flora. Approximately 4,442 species of plants and fungi were recorded from Maluku Islands until 2017 (Retnowati and Rugayah 2019). Herbarium Amboinense, a classic work by
We here present the first ever checklist of liverworts and hornworts for Maluku Islands in an effort to further promote bryological research, especially in the eastern part of Indonesia. The checklist complements recently published checklists for Indonesia including
Not less than 113 names (9 of them invalid) have original material from the Maluku Islands. Of the validly published names, 13 have holotypes and 47 of the rest are lectotypified. However, 43 names remain to be typified. Here we designate lectotypes, or validate earlier unsuccessful lectotypifications, for 13 names. For a discussion on lectotypification procedures, including the concept of inadvertent lectotypifications, see
Chandonanthus gracilis Herzog, Hedwigia 66 (6): 341, 1926 (
Original citation. Ceram: Hoale-Paß, ca. 1600 m, leg. E. Stresemann, 1911.
Lectotype (here designated, cf.
Note. Only one specimen has been located but it cannot be ruled out that there exist duplicates elsewhere.
Currently accepted name. Plicanthus hirtellus (F.Weber) R.M.Schust. (see
Lepidozia longifolia Steph., Sp. Hepat. (Stephani) 3: 606, 1909 (
Original citation. Hab. Amboina (Karsten).
Lectotype (here designated). Indonesia, Moluccas, Ambon, Karsten (G00069659 https://www.ville-ge.ch/musinfo/bd/cjb/chg/adetail.php?id=123550&lang=en).
Note. The lectotype is annotated “type” by Mizutani in Feb. 1975.
Lophocolea deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 321, 1921 (
Original citation. Buru: Im Urwald des Kapalamadang, leg. Dr. K. Deninger †, 1906.
Lectotype (here designated). Indonesia, Moluccas, Buru, Im Urwald des Kapalamadang, Deninger 1906 (JE04005314 https://je.jacq.org/JE04005314).
Note. JE04005314 is marked as “holotype” by R. Grolle. We are not aware of any other type material.
Lopholejeunea sagrana var. dentistipula Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19 (1): 246, 1894 (
Original citation. Ins. Amboina, in iisdem locis ac typica, pauca tantum frustula sed c. per. et ♂ inventa sunt (7. 6. 75).
Lectotype (here designated,
Note.
Currently accepted name. Caudalejeunea reniloba (Gottsche) Steph. (see
Madotheca crenilobula Herzog, Beih. Bot. Centralbl. 38 (2): 328, 1921 (
Original citation. Buru: Im Urwald des Kapalamadang, leg. Dr. K. Deninger †, 1906.
Lectotype (here designated). Indonesia, Moluccas, Buru, Im Urwald des Kapalamadang, 1906, K. Deninger (JE04001514 https://je.jacq.org/JE04001514, isotype JE04001515 https://je.jacq.org/JE04001515).
Note. The lectotype was annotated “Type!” and the isotype “isotype” by S. Hattori in 1968 but has apparently never been published.
Currently accepted name. Porella javanica (Gottsche) Inoue (see
Mastigobryum deningeri Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (
Original citation. Buru: In den Bergen Mittelburus, ca. 1500 m, leg. Dr. K. Deninger †, 1911.
Lectotype (here designated). Indonesia, Moluccas, Buru, Mittelburu, ca. 1500 m, 1911, Deninger 10 (JE04006158 https://je.jacq.org/JE04006158, isolectotype JE04006157 https://je.jacq.org/JE04006157).
Note. The selected lectotype is richer than the isolectotype and has a label in Herzog’s handwriting marked “n.sp.”. The isolectotype is marked “Typus” and annotated as type and syn. to Bazzania longicaulis by Kitagawa 18 Nov 1978 but no typification seems to have been published.
Currently accepted name. Bazzania longicaulis (Sande Lac.) Schiffn., syn. nov.>
Mastigobryum nigricans Herzog, Beih. Bot. Centralbl. 38 (2): 322, 1921 (
Original citation. Buru: Im Urwald des Kapalamadang, leg. Dr. K. Deninger †, 1906.
Lectotype (here designated). Indonesia, Moluccas, Buru, Kapalamadang, 1906, K. Deninger 112 (JE04006129 https://je.jacq.org/JE04006129).
Note. The lectotype is marked “n.sp.” on the label and annotated by N. Kitagawa 1991 as type.
Mastigobryum stresemannii Herzog, Beih. Bot. Centralbl. 38 (2): 324, 1921 (
Original citation. Ceram: Paßhöhe zwischen Mansela und Wolu (Useahánpaß), ca. 1750 m, leg. E. Stresemann, 1911.
Lectotype (here designated). Indonesia, Moluccas, Seram, between Mansela and Wolu ca. 1750 m, 1911, E. Stresemann 110, (JE04005944 https://je.jacq.org/JE04005944).
Note. The lectotype is marked “n.sp.” on the label by Herzog and annotated as type by N. Kitagawa in 1978.
Currently accepted name. Bazzania stresemannii (Herzog) N.Kitag.
Mastigobryum vermiculare Herzog, Hedwigia 66 (6): 339, 1926 (
Original citation. Ceram: Hoale, ca.1100m, leg. E. Stresemann, 1911.
Lectotype (here designated). Indonesia, Moluccas, Seram, Hoale, ca. 1100 m, 1911, Stresemann (JE04006130 https://je.jacq.org/JE04006130).
Note. Only one specimen is online in JE and without any annotation.
Pycnolejeunea ventricosa Schiffn. ex P.Syd., Just’s Bot. Jahresber. 19 (1): 246, 1894 (
Original citation. In insula Amboina cum aliis Lejeuniis parcissime, cum fl. ♀ et periantbiis vetustis (7. 6. 75) [cf.
Lectotype (here designated, cf.
Note. “Holotype” by
Currently accepted name. Cheilolejeunea ventricosa (Schiffn. ex P.Syd.) Xiao L.He
Schistochila amboinensis Steph., Sp. Hepat. (Stephani) 4: 77, 1909 (
Original citation. Hab. Amboina.
Lectotype (here designated, cf.
Note.
Currently accepted name. Schistochila beccariana (De Not.) Trevis. (see
Schistochila inversa Herzog, Hedwigia 66 (6): 342, 1926 (
Original citation. Ceram: Hoale-Paß, ca. 1600 m, leg. E. Stresemann, 1911; in Bruchstücken aus Lepidozia ceramensis und L. Ferdinandi Mülleri herausgelesen.
Lectotype (here designated, cf.
Note.
Currently accepted name. Schistochila sciurea (Nees) Schiffn. (see
Schistochila purpurascens Herzog, Hedwigia 66 (6): 341, 1926 (
Original citation. Ceram: Hoale-Paß, ca. 1600 m, leg. E. Stresemann, 1911.
Lectotype (here designated, cf.
Note. “holotype” by
Currently accepted name. Schistochila acuminata Steph. (see
Nomenclature and taxonomy follow the world checklist of hornworts and liverworts (
We here accept reports of 355 species from the area in addition to 16 reported species that we consider as doubtful and 21 species reported whose occurrence we reject from the islands. Species number known varies a lot between the islands/archipelagos (Table
Number of accepted, doubtful and rejected species of liverworts and hornworts from Moluccas and the recognized areas (islands/archipelagos) within the Moluccas.
Area | Accepted | Doubtful | Rejected |
---|---|---|---|
Ambon | 143 | 11 | 10 |
Bacan | 26 | 0 | 0 |
Banda | 7 | 2 | 0 |
Buru | 13 | 0 | 2 |
Halmahera | 31 | 3 | 0 |
Morotai | 2 | 0 | 0 |
Obi | 2 | 0 | 0 |
Seram | 258 | 7 | 5 |
Sulu | 0 | 0 | 0 |
Ternate/Tidore | 22 | 0 | 2 |
Moluccas | 355 | 16 | 22 |
Dendroceros Nees
Dendroceros acutilobus Steph. MOLUCCAS: Ambon: LECTOTYPE [
Dendroceros cavernosus J.Haseg. MOLUCCAS: Seram:
Dendroceros crispus (Sw.) Nees. MOLUCCAS:
Dendroceros foliicola J.Haseg. MOLUCCAS: Seram:
Dendroceros javanicus (Nees) Nees. MOLUCCAS: Seram:
Dendroceros pedunculatus Steph. MOLUCCAS: Ambon: LECTOTYPE [
Dendroceros seramensis J.Haseg. MOLUCCAS: Seram: HOLOTYPE,
Folioceros D.C.Bharadwaj
Folioceros amboinensis (Schiffn.) Piippo. MOLUCCAS: Ambon:
Folioceros appendiculatus (Steph.) J.Haseg. MOLUCCAS: Seram:
Folioceros glandulosus (Lehm. et Lindenb.) D.C.Bharadwaj. MOLUCCAS:
Megaceros Campb.
Megaceros flagellaris (Mitt.) Steph. MOLUCCAS: Ambon:
Phaeoceros Prosk.
Phaeoceros carolinianus (Michx.) Prosk. MOLUCCAS: Seram:
Phaeomegaceros R.J.Duff, J.C.Villarreal, Cargill et Renzaglia
Phaeomegaceros hirticalyx (Steph.) R.J.Duff, J.C.Villarreal, Cargill et Renzaglia. MOLUCCAS: Ambon:
Acrolejeunea (Spruce) Steph.
Acrolejeunea fertilis (Reinw., Blume et Nees) Schiffn. MOLUCCAS: LECTOTYPE of Ptychocoleus tener [
Acrolejeunea pycnoclada (Taylor) Schiffn. MOLUCCAS:
— subsp. pycnoclada. MOLUCCAS: Ambon: LECTOTYPE of Acrolejeunea rostrata [
Acrolejeunea securifolia (Nees) Steph. MOLUCCAS: Seram:
— subsp. hartmannii (Steph.) Gradst. MOLUCCAS: Ambon:
Acromastigum A.Evans
Acromastigum aurescens A.Evans. MOLUCCAS: Ambon:
Acromastigum bancanum (Sande Lac.) A.Evans. MOLUCCAS: Ambon:
Acromastigum divaricatum (Nees) A.Evans ex Reimers. MOLUCCAS: Ambon:
Acromastigum echinatiforme (De Not.) A.Evans. MOLUCCAS: Ambon:
Acromastigum inaequilaterum (Lehm. et Lindenb.) A.Evans. MOLUCCAS: Ambon:
Acromastigum lobuliferum A.Evans. MOLUCCAS: Ambon:
Acroscyphella N.Kitag. et Grolle
Acroscyphella tjiwideiensis (Sande Lac.) N.Kitag. et Grolle. MOLUCCAS:
Allorgella Tixier
Allorgella semperiana (Steph.) Bechteler, G.E.Lee, Schäf.-Verw. et Heinrichs. MOLUCCAS: Seram:
Allorgella zantenii (Grolle) Bechteler, G.E.Lee, Schäf.-Verw. et Heinrichs. MOLUCCAS: Seram:
Anastrophyllopsis (R.M.Schust.) Váňa et L.Söderstr.
Anastrophyllopsis revoluta (Steph.) Váňa et L.Söderstr. MOLUCCAS:
Anastrophyllum (Spruce) Steph.
Anastrophyllum piligerum (Nees) Steph. MOLUCCAS:
Aneura Dumort.
Aneura amboinensis Steph. MOLUCCAS: Ambon: LECTOTYPE [
Aneura pinguis (L.) Dumort. MOLUCCAS: Ambon:
Balantiopsis Mitt.
Balantiopsis ciliaris subsp. novoguineensis S.Hatt. MOLUCCAS: Seram:
Bazzania Gray
Bazzania asymmetrica (Steph.) N.Kitag. MOLUCCAS: Seram:
Bazzania cadens N.Kitag. MOLUCCAS: Seram:
Bazzania cincinnata (De Not.) Trevis. MOLUCCAS: Seram:
Bazzania commutata (Lindenb. et Gottsche) Schiffn. MOLUCCAS: Seram:
Bazzania conophylla (Sande Lac.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Mastigobryum longidens [
Bazzania crassitexta Steph. MOLUCCAS: Ambon: LECTOTYPE [
Bazzania distans (Nees) Trevis. MOLUCCAS: Seram:
Bazzania erosa (Reinw., Blume et Nees) Trevis. MOLUCCAS:
Bazzania fallax (Sande Lac.) Schiffn. MOLUCCAS: Ambon:
Bazzania horridula Schiffn. MOLUCCAS:
Bazzania insignis (De Not.) Trevis. MOLUCCAS: Ambon:
Bazzania intermedia (Gottsche et Lindenb.) Trevis. MOLUCCAS: Ambon:
— var. sarawakiana (De Not.) Schiffn. MOLUCCAS: ?Ambon:
Bazzania irregularis (Steph.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Mastigobryum irregulare [to be designated],
Bazzania javanica (Sande Lac.) Schiffn. MOLUCCAS: Khotimperwati et al. 2018. Seram:
Bazzania kernii Steph. MOLUCCAS: Seram:
Bazzania kokawana N.Kitag. et T.Kodama. MOLUCCAS: Seram:
Bazzania longicaulis (Sande Lac.) Schiffn. MOLUCCAS: Buru: LECTOTYPE of Mastigobryum deningeri [here designated],
Bazzania loricata (Reinw., Blume et Nees) Trevis. MOLUCCAS:
Bazzania merrillana (Steph.) Inoue ex Bonner. MOLUCCAS: ?Buru:
Bazzania parvitexta Steph. MOLUCCAS: Seram:
Bazzania pectinata (Lindenb. et Gottsche) Schiffn. MOLUCCAS:
Bazzania praerupta (Reinw., Blume et Nees) Trevis. MOLUCCAS: Khotimperwati et al. 2018. Seram:
Bazzania spiralis (Reinw., Blume et Nees) Meijer. MOLUCCAS: Halmahera:
Bazzania stresemannii (Herzog) N.Kitag. MOLUCCAS: Seram: LECTOTYPE of Mastigobryum stresemannii [here designated],
Bazzania subtilis (Sande Lac.) Trevis. MOLUCCAS: Ambon:
Bazzania tridens var. tridens. MOLUCCAS: Khotimperwati et al. 2018,
Bazzania uncigera (Reinw., Blume et Nees) Trevis. MOLUCCAS: Ambon:
Bazzania vittata (Gottsche) Trevis. MOLUCCAS: ?Ambon:
Bazzania wallichiana (Lindenb.) Trevis. MOLUCCAS: Ambon:
Calatholejeunea K.I.Goebel
Calatholejeunea paradoxa (Schiffn.) K.I.Goebel. MOLUCCAS: Ambon: LECTOTYPE of Lejeunea paradoxa [
Calypogeia Raddi
Calypogeia goebelii (Schiffn.) Steph. MOLUCCAS: Seram:
Caudalejeunea Schiffn.
Caudalejeunea cristiloba (Steph.) Gradst. MOLUCCAS:
Caudalejeunea reniloba (Gottsche) Steph. MOLUCCAS:
Cephalozia (Dumort.) Dumort.
Cephalozia stolonacea (Herzog) Váňa. MOLUCCAS: Ambon:
Ceratolejeunea (Spruce) J.B.Jack et Steph.
Ceratolejeunea belangeriana (Gottsche) Steph. MOLUCCAS: Seram:
Ceratolejeunea moniliata Herzog. MOLUCCAS: Seram:
Cheilolejeunea (Spruce) Steph.
Cheilolejeunea ceylanica (Gottsche) R.M.Schust. et Kachroo. MOLUCCAS:
Cheilolejeunea falsinervis (Sande Lac.) R.M.Schust. et Kachroo. MOLUCCAS: Bacan:
Cheilolejeunea gigantea (Steph.) R.M.Schust. et Kachroo. MOLUCCAS: Ambon: LECTOTYPE of Pycnolejeunea gigantea [
Cheilolejeunea incisa (Gottsche) R.M.Schust. et Kachroo. MOLUCCAS: Bacan:
Cheilolejeunea intertexta (Lindenb.) Steph. MOLUCCAS: Ambon:
Cheilolejeunea lindenbergii (Gottsche) Mizut. MOLUCCAS: Ambon: LECTOTYPE of Euosmolejeunea integristipula [
Cheilolejeunea longidens (Steph.) R.M.Schust. et Kachroo. MOLUCCAS: Ambon:
Cheilolejeunea occlusa (Herzog) T.Kodama et N.Kitag. MOLUCCAS: Seram:
Cheilolejeunea trapezia (Nees) Kachroo et R.M.Schust. MOLUCCAS: Ambon:
Cheilolejeunea trifaria (Reinw., Blume et Nees) Mizut. MOLUCCAS:
Cheilolejeunea ventricosa (Schiffn. ex P.Syd.) Xiao L.He. MOLUCCAS:
Cheilolejeunea vittata (Steph. ex G.Hoffm.) R.M.Schust. et Kachroo. MOLUCCAS: Seram:
Chiastocaulon Carl
Chiastocaulon braunianum (Nees) S.D.F.Patzak, M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Halmahera:
Chiastocaulon dendroides (Nees) Carl. MOLUCCAS:
Chiastocaulon oppositum (Reinw., Blume et Nees) S.D.F.Patzak, M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Ternate/Tidore: LECTOTYPE of Plagiochila opposita γ filiformis [to be designated],
Cladoradula (Spruce) M.A.M.Renner, Gradst., Ilk.-Borg. et F.R.Oliveira-da-Silva
Cladoradula campanigera (Mont.) M.A.M.Renner, Gradst., Ilk.-Borg. et F.R.Oliveira-da-Silva. MOLUCCAS: Seram:
Cololejeunea (Spruce) Steph.
Cololejeunea aequabilis (Sande Lac.) Schiffn. MOLUCCAS:
Cololejeunea angustiflora (Steph.) Mizut. MOLUCCAS: Seram:
Cololejeunea dozyana (Sande Lac.) Schiffn. MOLUCCAS:
Cololejeunea equialbi Tixier. MOLUCCAS:
Cololejeunea falcata (Horik.) Benedix. MOLUCCAS: Seram:
Cololejeunea floccosa (Lehm. et Lindenb.) Schiffn. MOLUCCAS: Seram:
Cololejeunea hamata Steph. MOLUCCAS: Ambon: LECTOTYPE [
Cololejeunea haskarliana (Lehm.) Schiffn. MOLUCCAS: Seram:
Cololejeunea inflectens (Mitt.) Benedix. MOLUCCAS:
Cololejeunea koponenii (Pócs) Pócs. MOLUCCAS: Seram:
Cololejeunea lanciloba Steph. MOLUCCAS: Seram:
Cololejeunea metzgeriopsis (K.I.Goebel) Gradst., R.Wilson, Ilk.-Borg. et Heinrichs. MOLUCCAS:
Cololejeunea obliqua (Nees et Mont.) Schiffn. MOLUCCAS: Seram:
Cololejeunea papillosa (K.I.Goebel) Mizut. MOLUCCAS: Seram:
Cololejeunea plagiophylla Benedix. MOLUCCAS: Seram:
Cololejeunea planissima var. planissima. MOLUCCAS: Seram:
Cololejeunea raduliloba Steph. MOLUCCAS: Seram:
Cololejeunea stephanii Schiffn. ex Benedix. MOLUCCAS: Seram:
Cololejeunea triapiculata (Herzog) Tixier. MOLUCCAS: Seram:
Cololejeunea trichomanis (Gottsche) Besch. MOLUCCAS: Seram:
Colura (Dumort.) Dumort.
Colura acroloba (Prantl) Jovet-Ast. MOLUCCAS: Seram:
Colura amboinensis Steph. MOLUCCAS: Ambon: LECTOTYPE [to be designated], ORIGINAL MATERIAL of Colurolejeunea superba var. typica,
Colura ari (Steph.) Steph. MOLUCCAS: Halmahera:
Colura conica (Sande Lac.) K.I.Goebel. MOLUCCAS: Seram:
Colura imperfecta Steph. MOLUCCAS: Seram:
Colura karstenii K.I.Goebel. MOLUCCAS: Ambon: LECTOTYPE [
Colura maxima Jovet-Ast. MOLUCCAS: Halmahera: LECTOTYPE [to be designated],
Colura ornata K.I.Goebel. MOLUCCAS:
Colura superba (Mont.) Steph. MOLUCCAS:
Conoscyphus Mitt.
Conoscyphus trapezioides (Sande Lac.) Schiffn. MOLUCCAS: Seram:
Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa
Cryptolophocolea costata (Nees) L.Söderstr. MOLUCCAS: Seram:
Cuspidatula Steph.
Cuspidatula contracta (Reinw., Blume et Nees) Steph. MOLUCCAS: Ambon:
Cuspidatula flaccida (Steph.) K.Feldberg, Váňa, Hentschel et Heinrichs. MOLUCCAS: Seram:
Cuspidatula flexicaulis (Nees) Váňa et L.Söderstr. MOLUCCAS:
Denotarisia Grolle
Denotarisia linguifolia (De Not.) Grolle. MOLUCCAS: Ambon: LECTOTYPE of Jungermannia ovifolia [
Dinckleria Trevis.
Dinckleria singularis (Schiffn.) M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Ternate/Tidore:
Diplasiolejeunea (Spruce) Schiffn.
Diplasiolejeunea cavifolia Steph. MOLUCCAS: Seram:
Diplasiolejeunea patelligera Herzog. MOLUCCAS: Seram:
Drepanolejeunea (Spruce) Steph.
Drepanolejeunea blumei var. angustistipa Herzog. MOLUCCAS:
Drepanolejeunea dactylophora (Nees, Lindenb. et Gottsche) J.B.Jack et Steph. MOLUCCAS:
Drepanolejeunea foliicola Horik. MOLUCCAS: Seram:
Drepanolejeunea grandis Herzog. MOLUCCAS: LECTOTYPE [to be designated],
Drepanolejeunea hampeana Steph. MOLUCCAS: Ambon:
Drepanolejeunea intermedia Zwickel. MOLUCCAS:
Drepanolejeunea levicornua Steph. MOLUCCAS: Seram:
Drepanolejeunea longicornua (Herzog) Mizut. MOLUCCAS: Seram:
Drepanolejeunea longicruris (Steph.) Grolle et R.L.Zhu. MOLUCCAS: Seram:
Drepanolejeunea lyrata Grolle. MOLUCCAS: Seram:
Drepanolejeunea moluccensis Herzog. MOLUCCAS: Bacan: LECTOTYPE [
Drepanolejeunea obliqua Steph. MOLUCCAS:
Drepanolejeunea pentadactyla (Mont.) Steph. MOLUCCAS:
— var. dactylophoroides (Herzog) Pócs. ?MOLUCCAS:
Drepanolejeunea spinosocornuta Steph. MOLUCCAS: Seram:
Drepanolejeunea tenera K.I.Goebel. MOLUCCAS:
Drepanolejeunea ternatensis (Gottsche) Schiffn. MOLUCCAS:
Drepanolejeunea thwaitesiana var. thwaitesiana. MOLUCCAS: Seram:
Drepanolejeunea tricornua Herzog. MOLUCCAS: Seram: LECTOTYPE [
Dumortiera Nees
Dumortiera hirsuta (Sw.) Nees. MOLUCCAS: Ambon:
Frullania Raddi
Frullania akiyamae S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
Frullania apiculata (Reinw., Blume et Nees) Nees. MOLUCCAS:
— var. apiculata. MOLUCCAS: Ambon:
— var. goebelii Schiffn. MOLUCCAS:
Frullania armatifolia Verd. MOLUCCAS: Seram:
Frullania attenuata Steph. MOLUCCAS: Seram:
Frullania berthoumieui Steph. MOLUCCAS: Ambon:
Frullania claviloba Steph. MOLUCCAS: Ambon: LECTOTYPE [
Frullania cordistipula (Reinw., Blume et Nees) Nees. MOLUCCAS: Halmahera:
— var. dentistipula S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
Frullania ericoides (Nees) Mont. MOLUCCAS:
Frullania fallax Gottsche. MOLUCCAS: Halmahera:
Frullania gaudichaudii (Nees et Mont.) Nees et Mont. MOLUCCAS: Ambon:
Frullania gracilis (Reinw., Blume et Nees) Nees. MOLUCCAS: Ambon: LECTOTYPE of Frullania lacerifolia [
— var. gracilis. MOLUCCAS: Seram:
— var. vittata S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
Frullania grandistipula Lindenb. MOLUCCAS: Bacan:
Frullania hasskarliana Lindenb. MOLUCCAS:
— var. gracilis S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
— var. hasskarliana. MOLUCCAS: Seram:
— var. parvidentata S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
Frullania heteromorpha Schiffn. MOLUCCAS: Seram:
Frullania integristipula (Nees) Nees
— var. emarginata Verd. MOLUCCAS: Halmahera:
— var. integristipula. MOLUCCAS: Halmahera:
Frullania intermedia (Reinw., Blume et Nees) Nees. MOLUCCAS:
— subsp. intermedia. MOLUCCAS: Ambon: LECTOTYPE of Frullania amboinensis [
Frullania junghuhniana Gottsche
— var. junghuhniana. MOLUCCAS: Seram:
— var. tenella (Sande Lac.) Grolle et S.Hatt. MOLUCCAS: Seram:
Frullania multilacera Steph. MOLUCCAS: Seram:
Frullania nepalensis (Spreng.) Lehm. et Lindenb. MOLUCCAS: Bacan:
Frullania nigricaulis (Reinw., Blume et Nees) Nees. MOLUCCAS: Halmahera:
Frullania nobilis var. cochleata (Steph.) S.Hatt. MOLUCCAS: Seram:
Frullania nodulosa (Reinw., Blume et Nees) Nees. MOLUCCAS:
Frullania orientalis Sande Lac. MOLUCCAS:
Frullania ornithocephala (Reinw., Blume et Nees) Nees. MOLUCCAS:
— var. major (Nees) Schiffn. MOLUCCAS: Ambon:
— var. ornithocephala. MOLUCCAS: Ambon:
Frullania pauciramea Steph. MOLUCCAS: Seram:
Frullania piptophylla var. minor S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
Frullania pulogensis Steph. MOLUCCAS: Seram:
Frullania ramuligera (Nees) Mont. MOLUCCAS: Seram:
Frullania reflexistipula var. reflexistipula MOLUCCAS:
Frullania repandistipula Sande Lac. MOLUCCAS: Ambon:
Frullania serrata Gottsche. MOLUCCAS:
— var. ceramensis S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
— var. hamatispina (S.Hatt.) S.Hatt. MOLUCCAS: Seram:
— var. serrata. MOLUCCAS: Seram:
Frullania subcaduca S.Hatt. MOLUCCAS: Seram:
Frullania sublignosa Steph. MOLUCCAS: Seram:
Frullania subnigricaulis var. subtruncata S.Hatt. MOLUCCAS: Seram:
Frullania subocellata S.Hatt. MOLUCCAS: Seram: HOLOTYPE,
Frullania ternatensis Gottsche. MOLUCCAS:
— var. non-appendiculata S.Hatt. MOLUCCAS: Seram:
— var. ternatensis. MOLUCCAS: Ternate/Tidore:
Frullania tricarinata Sande Lac. MOLUCCAS: Ambon:
Frullania trichodes Mitt. MOLUCCAS: Ambon: LECTOTYPE of Frullania picta [
Frullania vaginata (Sw.) Nees. MOLUCCAS: Ternate/Tidore:
Gottschelia Grolle
Gottschelia schizopleura (Spruce) Grolle. MOLUCCAS:
Hattoriolejeunea Mizut.
Hattoriolejeunea akiyamae Mizut. MOLUCCAS: Seram: HOLOTYPE,
Herbertus Gray
Herbertus longifissus Steph. MOLUCCAS: Seram:
Herbertus longispinus J.B.Jack et Steph. MOLUCCAS: Ambon:
Herbertus pilifer (Steph.) H.A.Mill. MOLUCCAS: Ambon: LECTOTYPE of Schisma pilifera [
Herbertus ramosus (Steph.) H.A.Mill. MOLUCCAS: Seram:
Herbertus sendtneri (Nees) Lindb. MOLUCCAS: Seram: LECTOTYPE of Schisma divaricatum [
Heteroscyphus Schiffn.
Heteroscyphus amboinensis (Schiffn.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Chiloscyphus endlicherianus δ amboinensis [to be designated],
Heteroscyphus argutus (Reinw., Blume et Nees) Schiffn. MOLUCCAS: Seram:
Heteroscyphus aselliformis (Reinw., Blume et Nees) Schiffn. MOLUCCAS: Ambon:
Heteroscyphus coalitus var. coalitus MOLUCCAS: Ambon:
Heteroscyphus splendens (Lehm. et Lindenb.) Grolle. MOLUCCAS: Halmahera:
Heteroscyphus zollingeri (Gottsche) Schiffn. MOLUCCAS: Seram:
Kurzia G.Martens
Kurzia geniculata Mizut. MOLUCCAS: Seram:
Kurzia gonyotricha (Sande Lac.) Grolle. MOLUCCAS:
Leiomitra Lindb.
Leiomitra breviseta (Steph.) R.M.Schust. MOLUCCAS: Seram:
Lejeunea Lib.
Lejeunea albescens (Steph.) Mizut. MOLUCCAS:
Lejeunea anisophylla Mont. MOLUCCAS: Ambon:
Lejeunea discreta Lindenb. MOLUCCAS:
Lejeunea exilis (Reinw., Blume et Nees) Grolle. MOLUCCAS:
Lejeunea fissistipula (Steph.) Steph. MOLUCCAS: Ambon: LECTOTYPE of Eulejeunea fissistipula [
Lejeunea flava subsp. orientalis R.M.Schust. MOLUCCAS: LECTOTYPE of Jungermannia indica [to be designated]. Ternate/Tidore: Reinwardt and Nees 1823 as Jungermannia indica.
Lejeunea lumbricoides (Nees) Nees. MOLUCCAS: Ambon: LECTOTYPE of Taxilejeunea karstenii [to be designated],
Lejeunea microloba Taylor. MOLUCCAS: Seram:
Lejeunea sordida (Nees) Nees. MOLUCCAS:
Lejeunea stenodentata M.A.M.Renner et Pócs. MOLUCCAS:
Lejeunea umbilicata (Nees) Nees. MOLUCCAS: Ternate/Tidore:
Lepicolea Dumort.
Lepicolea rara (Steph.) Grolle. MOLUCCAS:
Lepicolea yakusimensis (S.Hatt.) S.Hatt. MOLUCCAS: Seram:
Lepidolejeunea R.M.Schust.
Lepidolejeunea bidentula (Steph.) R.M.Schust. MOLUCCAS:
Lepidozia (Dumort.) Dumort.
Lepidozia borneensis Steph. MOLUCCAS: Seram:
Lepidozia brotheri Steph. MOLUCCAS: Seram:
Lepidozia ceramensis Herzog. MOLUCCAS: Seram: LECTOTYPE [to be designated],
Lepidozia cladorhiza (Reinw., Blume et Nees) Nees. MOLUCCAS: Seram:
Lepidozia ferdinandii-muelleri Steph. MOLUCCAS: Halmahera:
Lepidozia griseola Herzog. MOLUCCAS: Seram: LECTOTYPE [to be designated],
Lepidozia haskarliana (Gottsche, Lindenb. et Nees) Steph. MOLUCCAS:
Lepidozia integrifolia Doei. MOLUCCAS: Seram: HOLOTYPE,
Lepidozia longifolia Steph. MOLUCCAS: Ambon: LECTOTYPE [here designated],
Lepidozia massartiana Schiffn. ex Steph. MOLUCCAS: Ambon:
Lepidozia subintegra Lindenb. MOLUCCAS:
Lepidozia subtrichodes Steph. MOLUCCAS: Seram:
Lepidozia trichodes (Reinw., Blume et Nees) Nees. MOLUCCAS: Ambon:
Leptolejeunea (Spruce) Steph.
Leptolejeunea dentistipula Steph. MOLUCCAS: Seram:
Leptolejeunea elliptica (Lehm. et Lindenb.) Besch. MOLUCCAS:
Leptolejeunea epiphylla (Mitt.) Steph. MOLUCCAS:
Leptolejeunea foliicola Steph. MOLUCCAS:
Leptolejeunea maculata (Mitt.) Schiffn. MOLUCCAS:
Leptolejeunea schiffneri (Steph. ex Schiffn.) Steph. MOLUCCAS:
Leptolejeunea vitrea (Nees) Schiffn. MOLUCCAS:
Lobatiriccardia (Mizut. et S.Hatt.) Furuki
Lobatiriccardia coronopus (De Not.) Furuki. MOLUCCAS: Seram:
— subsp. coronopus. MOLUCCAS: Seram:
Lophocolea (Dumort.) Dumort.
Lophocolea deningeri Herzog. MOLUCCAS: Buru: LECTOTYPE [here designated],
Lophocolea kurzii Sande Lac. MOLUCCAS: Seram:
Lopholejeunea (Spruce) Steph.
Lopholejeunea applanata (Reinw., Blume et Nees) Schiffn. MOLUCCAS: Ternate/Tidore:
Lopholejeunea eulopha (Taylor) Schiffn. MOLUCCAS:
Lopholejeunea horticola Schiffn. MOLUCCAS:
Lopholejeunea nigricans (Lindenb.) Schiffn. MOLUCCAS:
Lopholejeunea subfusca (Nees) Schiffn. MOLUCCAS:
Lopholejeunea wiltensii Steph. MOLUCCAS:
Marchantia L.
Marchantia emarginata Reinw., Blume et Nees. MOLUCCAS:
— subsp. emarginata. MOLUCCAS:
Marchantia geminata Reinw., Blume et Nees. MOLUCCAS: Halmahera:
Marchantia polymorpha L. MOLUCCAS: Ambon:
Marchantia rubribarba Steph. MOLUCCAS: Ambon:
Mastigophora Nees
Mastigophora diclados (Brid. ex F.Weber) Nees. MOLUCCAS: Buru:
— var. borneensis (De Not.) Schiffn. MOLUCCAS: Ambon:
— var. diclados. MOLUCCAS: Ambon:
Metalejeunea Grolle
Metalejeunea cucullata (Reinw., Blume et Nees) Grolle. MOLUCCAS:
Metzgeria Raddi
Metzgeria ciliata Raddi. MOLUCCAS: Halmahera:
Metzgeria foliicola Schiffn. MOLUCCAS: Ternate/Tidore:
Metzgeria furcata (L.) Corda. MOLUCCAS: Halmahera:
Metzgeria leptoneura var. leptoneura MOLUCCAS:
Microlejeunea (Spruce) Steph.
Microlejeunea filicuspis (Steph.) Heinrichs, Schäf.-Verw., Pócs et S.Dong. MOLUCCAS:
Mnioloma Herzog
Mnioloma fuscum (Lehm.) R.M.Schust. MOLUCCAS: Seram:
Neolepidozia Fulford et J.Taylor
Neolepidozia cuneifolia (Steph.) Fulford et J.Taylor. MOLUCCAS: Ambon:
Neolepidozia papulosa (Steph.) Fulford et J.Taylor. MOLUCCAS: Seram:
Neolepidozia wallichiana (Gottsche) Fulford et J.Taylor. MOLUCCAS: Ambon:
Pallavicinia Gray
Pallavicinia ambigua (Mitt.) Steph. MOLUCCAS: Seram:
Pallavicinia lyellii (Hook.) Gray. MOLUCCAS:
Plagiochila (Dumort.) Dumort.
Plagiochila abietina (Nees) Mont. et Nees. MOLUCCAS: Ambon:
Plagiochila akiyamae Inoue. MOLUCCAS: Seram: HOLOTYPE,
Plagiochila amboynensis Taylor. MOLUCCAS:
Plagiochila arbuscula (Brid. ex Lehm. et Lindenb.) Lindenb. MOLUCCAS: Halmahera:
Plagiochila bantamensis (Reinw., Blume et Nees) Mont. MOLUCCAS: Seram:
Plagiochila bicornuta Steph. MOLUCCAS:
Plagiochila blepharophora (Nees) Lindenb. MOLUCCAS: Ternate/Tidore:
Plagiochila chauviniana Mont. MOLUCCAS: Seram:
Plagiochila frondescens (Nees) Lindenb. MOLUCCAS: Ambon:
Plagiochila gymnoclada Sande Lac. MOLUCCAS:
Plagiochila hampeana Gottsche. MOLUCCAS: Seram:
Plagiochila junghuhniana Sande Lac. MOLUCCAS: Ambon:
Plagiochila kuhliana Sande Lac. MOLUCCAS: Seram:
Plagiochila kurzii Steph. MOLUCCAS: Seram:
Plagiochila massalongoana Schiffn. MOLUCCAS: Seram:
Plagiochila pulvinata Steph. MOLUCCAS:
Plagiochila renitens (Nees) Lindenb. MOLUCCAS: ?Halmahera:
Plagiochila salacensis Gottsche. MOLUCCAS: Seram:
Plagiochila sandei Dozy. MOLUCCAS:
Plagiochila sciophila Nees. MOLUCCAS: ?Ambon:
Plagiochila teysmannii Sande Lac. MOLUCCAS:
Plagiochila trabeculata Steph. MOLUCCAS: Seram:
Pleurozia Dumort.
Pleurozia conchifolia (Hook. et Arn.) Austin. MOLUCCAS:
Pleurozia gigantea (F.Weber) Lindb. MOLUCCAS: Ambon:
Plicanthus R.M.Schust.
Plicanthus hirtellus (F.Weber) R.M.Schust. MOLUCCAS: Seram: LECTOTYPE of Chandonanthus gracilis [here designated],
Podomitrium Mitt.
Podomitrium malaccense (Steph.) Campb. MOLUCCAS: Ambon:
Porella L.
Porella acutifolia var. acutifolia MOLUCCAS: Halmahera:
Porella geheebii (Steph.) S.Hatt. MOLUCCAS: Seram:
Porella javanica (Gottsche) Inoue. MOLUCCAS:
Psiloclada Mitt.
Psiloclada clandestina subsp. clandestina MOLUCCAS: Ambon: LECTOTYPE of Psiloclada unguligera [to be designated],
— subsp. melanesica R.M.Schust. ?MOLUCCAS: Ambon:
Ptychanthus Nees
Ptychanthus striatus var. striatus. MOLUCCAS:
Pycnolejeunea (Spruce) Schiffn.
Pycnolejeunea cavistipula (Steph.) Mizut. MOLUCCAS: Ambon: LECTOTYPE of Strepsilejeunea cavistipula [
Pycnolejeunea connivens Schiffn. ex P.Syd. MOLUCCAS: Ambon: LECTOTYPE of Pycnolejeunea connivens [to be designated],
Radula Dumort.
Radula anceps Sande Lac. MOLUCCAS: Seram:
Radula densifolia Castle. MOLUCCAS: Seram:
Radula formosa (C.F.W.Meissn. ex Spreng.) Nees. MOLUCCAS: Ambon: LECTOTYPE of Radula pycnolejeuneoides [
Radula javanica Gottsche. MOLUCCAS: Buru: New to Moluccas: Buru, E hoe, 1921.09, Toxopens 299a, det. H. Castle (BO4137]. Seram: LECTOTYPE of Radula ceramensis [to be designated], LECTOTYPE of Radula ovalifolia [
Radula multiflora Gottsche ex Schiffn. MOLUCCAS: Seram:
Radula nymannii Steph. MOLUCCAS: Ambon:
Radula reflexa Nees et Mont. MOLUCCAS: Ambon: LECTOTYPE [
Radula sumatrana Steph. MOLUCCAS: Seram:
Radula tjibodensis K.I.Goebel. MOLUCCAS: Seram:
Radula van-zantenii K.Yamada. MOLUCCAS: Seram:
Radula vrieseana Sande Lac. MOLUCCAS:
Riccardia Gray
Riccardia albomarginata (Steph.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Aneura albomarginata [
Riccardia aspera (Steph.) Grolle. MOLUCCAS: Seram:
Riccardia karstenii (Steph.) Schiffn. MOLUCCAS: Ambon: LECTOTYPE of Aneura karstenii [
Riccia L.
Riccia amboinensis Schiffn. MOLUCCAS: Ambon: LECTOTYPE [to be designated],
Riccia billardierei Mont. et Nees. MOLUCCAS:
Riccia junghuhniana Nees et Lindenb. MOLUCCAS: Ambon:
Saccogynidium Grolle
Saccogynidium muricellum (De Not.) Grolle. MOLUCCAS:
Saccogynidium rigidulum (Nees) Grolle. MOLUCCAS:
Sandeothallus R.M.Schust.
Sandeothallus radiculosus (Schiffn.) R.M.Schust. MOLUCCAS: Ambon:
Schiffneria Steph.
Schiffneria hyalina Steph. MOLUCCAS:
Schiffneriolejeunea Verd.
Schiffneriolejeunea cumingiana (Mont.) Gradst. MOLUCCAS: Ambon: Verdoorn 1934a, 1934b both as Ptychocoleus cumingianus,
Schiffneriolejeunea pulopenangensis (Gottsche) Gradst. MOLUCCAS:
Schiffneriolejeunea tumida (Nees) Gradst. MOLUCCAS: Ambon:
— var. haskarliana (Gottsche) Gradst. et Terken. MOLUCCAS:
Schistochila Dumort.
Schistochila acuminata Steph. MOLUCCAS: Seram: LECTOTYPE of Schistochila purpurascens [here designated, cf.
Schistochila aligera (Nees et Blume) J.B.Jack et Steph. MOLUCCAS:
Schistochila beccariana (De Not.) Trevis. MOLUCCAS: Ambon: LECTOTYPE of Schistochila amboinensis [here designated, cf.
Schistochila blumei (Nees) Trevis. MOLUCCAS:
Schistochila doriae (De Not.) Trevis. MOLUCCAS: Ambon:
Schistochila reinwardtii (Nees) Schiffn. MOLUCCAS: Ambon:
Schistochila sciurea var. sciurea. MOLUCCAS: Ambon:
Schizophyllopsis Váňa et L.Söderstr.
Schizophyllopsis bidens (Reinw., Blume et Nees) Váňa et L.Söderstr. MOLUCCAS: Seram:
Solenostoma Mitt.
Solenostoma ariadne (Taylor) R.M.Schust. ex Váňa et D.G.Long. MOLUCCAS: Ambon:
Solenostoma comatum var. comatum. MOLUCCAS: Ternate/Tidore: Sande Lacoste 1856b as Plagiochila comata,
Solenostoma tetragonum (Lindenb.) R.M.Schust. ex Váňa et D.G.Long. MOLUCCAS: Ambon:
Solenostoma truncatum (Nees) R.M.Schust. ex Váňa et D.G.Long. MOLUCCAS: Halmahera:
Spruceanthus Verd.
Spruceanthus polymorphus (Sande Lac.) Verd. MOLUCCAS: Halmahera:
Symphyogynopsis Grolle
Symphyogynopsis gottscheana (Mont. et Nees) Grolle. MOLUCCAS:
Syzygiella Spruce
Syzygiella ovalifolia Inoue. MOLUCCAS: Seram:
Syzygiella securifolia (Nees) Inoue. MOLUCCAS: Halmahera:
Telaranea Spruce ex Schiffn.
Telaranea major (Herzog) J.J.Engel et G.L.Merr. MOLUCCAS:
Telaranea trisetosa (Steph.) Grolle. MOLUCCAS: Seram:
Temnoma Mitt.
Temnoma setigerum (Lindenb.) R.M.Schust. MOLUCCAS: Seram:
Thysananthus Lindenb.
Thysananthus appendiculatus Steph. MOLUCCAS: Ambon:
Thysananthus convolutus var. convolutus. MOLUCCAS:
Thysananthus frauenfeldii Reichardt. MOLUCCAS:
Thysananthus fruticosus (Lindenb. et Gottsche) Schiffn. MOLUCCAS:
Thysananthus humilis (Gottsche) Sukkharak et Gradst. MOLUCCAS:
Thysananthus ligulatus (Lehm. et Lindenb.) Sukkharak et Gradst. MOLUCCAS: Buru:
Thysananthus repletus (Taylor) Sukkharak et Gradst. MOLUCCAS: Ambon:
Thysananthus spathulistipus (Reinw., Blume et Nees) Lindenb. MOLUCCAS:
Thysananthus truncatus (Mizut.) Sukkharak et Gradst. MOLUCCAS: Seram:
Thysananthus virens Ångstr. MOLUCCAS:
Treubia K.I.Goebel
Treubia insignis K.I.Goebel. MOLUCCAS: Seram:
Trichocolea Dumort.
Trichocolea pluma (Reinw., Blume et Nees) Mont. MOLUCCAS: Buru:
Trichocolea tomentella (Ehrh.) Dumort. MOLUCCAS: Morotai: New to Moluccas: Morotai, Sangowo, Exp. Kostermans) (BO13087)]. Ambon:
Tricholepidozia (R.M.Schust.) E.D.Cooper
Tricholepidozia kogiana (Steph.) E.D.Cooper. MOLUCCAS: Seram:
Tricholepidozia neesii (Lindenb.) E.D.Cooper. MOLUCCAS: Halmahera:
Tuyamaella S.Hatt.
Tuyamaella serratistipa S.Hatt. MOLUCCAS: Seram:
Wettsteinia Schiffn.
Wettsteinia inversa (Sande Lac.) Schiffn. MOLUCCAS: Seram:
Zantenia (S.Hatt.) Váňa et J.J.Engel
Zantenia borneensis (Herzog) Váňa et J.J.Engel. MOLUCCAS: Seram:
Zantenia karstenii (Schiffn.) Váňa et J.J.Engel. MOLUCCAS:
Zoopsis Hook.f. ex Gottsche, Lindenb. et Nees
Zoopsis liukiuensis Horik. MOLUCCAS:
Zoopsis setigera K.I.Goebel. MOLUCCAS: Ambon: LECTOTYPE [
We are not able to refer the following published names to any accepted taxon.
Phaeoceros Prosk.
Phaeoceros velutinus J.Haseg. ex H.Akiyama nom. inval. MOLUCCAS: Ambon: ORIGINAL MATERIAL. Seram:
Jungermannia L.
Jungermannia gigantea β laxior (Nees) Lindenb. MOLUCCAS: Ternate/Tidore: SYNTYPE of Jungermannia gigantea β laxior [lectotype to be designated], ORIGINAL MATERIAL of Plagiochila trapezoidea γ major,
Jungermannia tamarisci L. var. β minus-ramosa Schwägr. nom. inval. MOLUCCAS: ORIGINAL MATERIAL,
Plagiochila (Dumort.) Dumort.
Plagiochila opposita (Reinw., Blume et Nees) Lindenb. β falcata (Nees) Schiffn. MOLUCCAS: Ternate/Tidore:
We doubt the identification of the following taxa or we are not aware of any first-hand report from Moluccas.
Acromastigum A.Evans
Acromastigum echinatum (Gottsche) A.Evans. MOLUCCAS: Ambon:
Bazzania Gray
Bazzania ceylanica (Mitt.) Steph. MOLUCCAS: Ambon:
Bazzania japonica (Sande Lac.) Lindb. MOLUCCAS:
Calypogeia Raddi
Calypogeia tosana (Steph.) Steph. MOLUCCAS: Halmahera:
Chiloscyphus Corda
Chiloscyphus ernstianus Steph. MOLUCCAS: Seram:
Cololejeunea (Spruce) Steph.
Cololejeunea cordiflora Steph. MOLUCCAS: Ambon:
Cololejeunea verrucosa Steph. MOLUCCAS:
Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa
Cryptolophocolea ciliolata (Nees) L.Söderstr., Crand.-Stotl., Stotler et Váňa. MOLUCCAS: Seram:
Leptolejeunea (Spruce) Steph.
Leptolejeunea balansae Steph. MOLUCCAS: Ambon:
Leptolejeunea subacuta Steph. ex A.Evans. MOLUCCAS:
Mastigophora Nees
Mastigophora woodsii (Hook.) Nees. MOLUCCAS: Seram: LECTOTYPE of Mastigophora ramentifissa [to be designated], Herzog 1926a, 1926b as Mastigophora ramentifissa,
Plagiochila (Dumort.) Dumort.
Plagiochila gracilis Lindenb. et Gottsche. MOLUCCAS:
Radula Dumort.
Radula amentulosa Mitt. MOLUCCAS: Seram:
Schistochila Dumort.
Schistochila schultzei Steph. MOLUCCAS: Seram:
Spruceanthus Verd.
Spruceanthus semirepandus (Nees) Verd. MOLUCCAS:
Thysananthus Lindenb.
Thysananthus auriculatus (Wilson et Hook.) Sukkharak et Gradst. MOLUCCAS:
The following taxa are reported to occur in Moluccas but the reports are rejected either earlier or here as misidentified or highly unlikely to occur in the area.
Cheilolejeunea (Spruce) Steph.
Cheilolejeunea mariana (Gottsche) B.M.Thiers et Gradst. MOLUCCAS: Seram:
Chiastocaulon Carl
Chiastocaulon fimbriatum (Mitt.) S.D.F.Patzak, M.A.M.Renner, Schäf.-Verw. et Heinrichs. MOLUCCAS: Seram:
Conocephalum Hill
Conocephalum conicum (L.) Dumort. MOLUCCAS: Ambon:
Cryptolophocolea L.Söderstr., Crand.-Stotl., Stotler et Váňa
Cryptolophocolea connata (Sw.) L.Söderstr. et Váňa. MOLUCCAS:
Dendroceros Nees
Dendroceros exalatus Steph. MOLUCCAS: Ambon: LECTOTYPE [
Heteroscyphus Schiffn.
Heteroscyphus caledonicus (Steph.) Schiffn. MOLUCCAS: Ambon:
Lepicolea Dumort.
Lepicolea ochroleuca (Spreng.) Spruce. MOLUCCAS: Seram:
Lepicolea scolopendra (Hook.) Dumort. ex Trevis. MOLUCCAS: Seram:
Lepidozia (Dumort.) Dumort.
Lepidozia microphylla (Hook.) Lindenb. MOLUCCAS: Seram:
Lepidozia rigida Steph. MOLUCCAS: Buru: ORIGINAL MATERIAL of Lepidozia rigida f. minor,
Plagiochila (Dumort.) Dumort.
Plagiochila gigantea Lindenb. MOLUCCAS: Ternate/Tidore:
Plagiochila trapezoidea Lindenb. MOLUCCAS: Ternate/Tidore:
Podomitrium Mitt.
Podomitrium phyllanthus (Hook.) Mitt. MOLUCCAS: Ambon:
Porella L.
Porella nitens (Steph.) S.Hatt. MOLUCCAS:
Pycnolejeunea (Spruce) Schiffn.
Pycnolejeunea schwaneckei (Steph.) Schiffn. ex P.Syd. MOLUCCAS: Ambon:
Radula Dumort.
Radula pinnulata Mitt. MOLUCCAS:
Riccardia Gray
Riccardia latifrons (Lindb.) Lindb. MOLUCCAS: Ambon:
Schistochila Dumort.
Schistochila neesii (Mont.) Lindb. MOLUCCAS: Ambon:
Symbiezidium Trevis.
Symbiezidium transversale (Sw.) Trevis. MOLUCCAS:
Zoopsis Hook.f. ex Gottsche, Lindenb. et Nees
Zoopsis argentea (Hook.f. et Taylor) Gottsche, Lindenb. et Nees. MOLUCCAS: Ambon:
Zoopsis setulosa Leitg. MOLUCCAS:
Acrolejeunea densifolia (Schiffn.) Schiffn. ex P.Syd. nom. inval. = Schiffneriolejeunea pulopenangensis
Acrolejeunea densifolia Schiffn. nom. inval. = Schiffneriolejeunea pulopenangensis
Acrolejeunea haskarliana (Gottsche) Schiffn. ≡ Schiffneriolejeunea tumida var. haskarliana
Acrolejeunea pulopenangensis (Gottsche) Schiffn. ≡ Schiffneriolejeunea pulopenangensis
Acrolejeunea rostrata Schiffn. nom. inval. = Acrolejeunea pycnoclada subsp. pycnoclada
Acrolejeunea rostrata var. α minor Schiffn. nom. inval. = Acrolejeunea pycnoclada subsp. pycnoclada
Acrolejeunea rostrata var. β major Schiffn. nom. inval. = Acrolejeunea pycnoclada subsp. pycnoclada
Acrolejeunea securifolia (Nees) Steph. nom. inval. ≡ Acrolejeunea securifolia
Acromastigum bidenticulatum A.Evans = Acromastigum bancanum
Anastrophyllum bidens (Reinw., Blume et Nees) Steph. ≡ Schizophyllopsis bidens
Anastrophyllum borneense Herzog ≡ Zantenia borneensis
Anastrophyllum contractum (Reinw., Blume et Nees) Steph. ≡ Cuspidatula contracta
Anastrophyllum karstenii Schiffn. ≡ Zantenia karstenii
Anastrophyllum revolutum Steph. ≡ Anastrophyllopsis revoluta
Aneura albomarginata Steph. ≡ Riccardia albomarginata
Aneura karstenii Steph. ≡ Riccardia karstenii
Aneura latifrons Lindb. ≡ Riccardia latifrons
Aneura pinguis f. normalis Schiffn. nom. inval. = Aneura pinguis
Anomacaulis flaccidus (Steph.) Grolle ≡ Cuspidatula flaccida
Anthoceros amboinensis Schiffn. ≡ Folioceros amboinensis
Anthoceros appendiculatus Steph. ≡ Folioceros appendiculatus
Anthoceros glandulosus Lehm. et Lindenb. ≡ Folioceros glandulosus
Anthoceros grandis Ångstr. = Megaceros flagellaris
Arachniopsis major Herzog ≡ Telaranea major
Aspiromitus amboinensis (Schiffn.) Steph. ≡ Folioceros amboinensis
Bazzania australis (Mont.) Trevis. = Bazzania tridens
Bazzania concinna (De Not.) Trevis. = Bazzania intermedia
Bazzania divaricata (Nees) Trevis. = Acromastigum divaricatum
Bazzania echinatiformis (De Not.) Trevis. ≡ Acromastigum echinatiforme
Bazzania intermedia auct. = Bazzania tridens var. tridens
Bazzania longidens (Steph.) Schiffn. = Bazzania conophylla
Bazzania oshimensis (Steph.) Horik. = Bazzania tridens
Brachiolejeunea molukkensis Steph. = Thysananthus humilis
Bryopteris fruticosa Lindenb. et Gottsche ≡ Thysananthus fruticosus
Calypogeia fusca (Lehm.) Steph. ≡ Mnioloma fuscum
Campylolejeunea ciliatilobula (Schiffn.) S.Hatt. = Cololejeunea inflectens
Campylolejeunea inflectens (Mitt.) Mizut. ≡ Cololejeunea inflectens
Caudalejeunea circinata Steph. = Caudalejeunea cristiloba
Caudalejeunea recurvistipula (Gottsche) Schiffn. = Caudalejeunea reniloba
Cephalozia setulosa (Leitg.) Spruce ≡ Zoopsis setulosa
Chandonanthus gracilis Herzog = Plicanthus hirtellus
Chandonanthus hirtellus (F.Weber) Mitt. ≡ Plicanthus hirtellus
Cheilolejeunea imbricata (Nees) S.Hatt. = Cheilolejeunea trapezia
Cheilolejeunea meyeniana (Nees, Lindenb. et Gottsche) R.M.Schust. et Kachroo = Cheilolejeunea trapezia
Chiloscyphus amboinensis (Schiffn.) Schiffn. ≡ Heteroscyphus amboinensis
Chiloscyphus aselliformis (Reinw., Blume et Nees) Nees ≡ Heteroscyphus aselliformis
Chiloscyphus coalitus (Hook.) Nees ≡ Heteroscyphus coalitus
Chiloscyphus decurrens (Reinw., Blume et Nees) Nees = Heteroscyphus splendens
Chiloscyphus decurrens var. amboinensis (Schiffn.) Schiffn. ex Bonner nom. inval. ≡ Heteroscyphus amboinensis
Chiloscyphus endlicherianus δ amboinensis Schiffn. ≡ Heteroscyphus amboinensis
Chiloscyphus falcifolius Steph. = Acroscyphella tjiwideiensis
Chiloscyphus granulatus Schiffn. = Saccogynidium rigidulum
Clasmatocolea tjiwideiensis (Sande Lac.) Grolle ≡ Acroscyphella tjiwideiensis
Cololejeunea ciliatilobula (Schiffn.) Schiffn. = Cololejeunea inflectens
Cololejeunea crenulata (Herzog) Benedix nom. inval. = Cololejeunea angustiflora
Cololejeunea falcatoides Benedix = Cololejeunea falcata
Cololejeunea goebelii (Gottsche ex Schiffn.) Schiffn. = Cololejeunea trichomanis
Cololejeunea javanica (Steph.) Mizut. = Cololejeunea angustiflora
Cololejeunea karstenii (Steph.) Benedix = Cololejeunea cordiflora
Cololejeunea nymannii (Steph.) Benedix = Cololejeunea obliqua
Cololejeunea peculiaris (Herzog) Benedix = Cololejeunea inflectens
Cololejeunea yulensis (Steph.) Benedix = Cololejeunea aequabilis
Colura paradoxa (Schiffn.) Steph. ≡ Calatholejeunea paradoxa
Colurolejeunea karstenii (K.I.Goebel) Steph. ≡ Colura karstenii
Colurolejeunea karstenii var. latifolia (Schiffn.) Schiffn. nom. inval. = Colura karstenii
Colurolejeunea paradoxa (Schiffn.) Schiffn. ≡ Calatholejeunea paradoxa
Colurolejeunea superba (Mont.) Steph. ≡ Colura superba
Colurolejeunea superba Schiffn. nom. inval. = Colura amboinensis
Colurolejeunea superba var. α typica Schiffn. nom. inval. = Colura amboinensis
Dendroceros karstenii Schiffn. ex Steph. = Dendroceros acutilobus
Drepanolejeunea dactylophora (Nees, Lindenb. et Gottsche) Schiffn. ≡ Drepanolejeunea dactylophora
Drepanolejeunea dentata Steph. ≡ Lejeunea stenodentata
Drepanolejeunea filicuspis Steph. ≡ Microlejeunea filicuspis
Drepanolejeunea levicornua var. longicornua Herzog ≡ Drepanolejeunea longicornua
Drepanolejeunea micholitzii Steph. = Drepanolejeunea pentadactyla
Drepanolejeunea micholitzii var. dactylophoroides Herzog ≡ Drepanolejeunea pentadactyla var. dactylophoroides
Drepanolejeunea micholitzii var. genuina Herzog nom. illeg. = Drepanolejeunea pentadactyla
Drepanolejeunea tenera var. genuina Herzog nom. illeg. = Drepanolejeunea tenera
Drepanolejeunea ternatensis (Gottsche) Steph. nom. inval. ≡ Drepanolejeunea ternatensis
Eucalyx comata (Nees) Verd. ≡ Solenostoma comatum
Eulejeunea fissistipula Steph. ≡ Lejeunea fissistipula
Eulejeunea parallela (Schiffn.) Schiffn. = Metalejeunea cucullata
Euosmolejeunea heterophylla (Sande Lac.) Schiffn. = Cheilolejeunea trifaria
Euosmolejeunea integristipula Steph. = Cheilolejeunea lindenbergii
Euosmolejeunea trifaria (Reinw., Blume et Nees) Steph. ≡ Cheilolejeunea trifaria
Frullania amboinensis Schiffn. = Frullania intermedia subsp. intermedia
Frullania billardiereana Nees et Mont. = Frullania intermedia subsp. intermedia
Frullania dapitana Steph. = Frullania nodulosa
Frullania ericoides var. planescens (Verd.) S.Hatt. = Frullania ericoides
Frullania gracilis var. lacerifolia (Steph.) Verd. = Frullania gracilis
Frullania intermedia f. billardiereana (Nees et Mont.) Verd. = Frullania intermedia subsp. intermedia
Frullania intermedia var. amboinensis (Schiffn.) Verd. = Frullania intermedia subsp. intermedia
Frullania karstenii Schiffn. = Frullania apiculata var. goebelii
Frullania lacerifolia Steph. = Frullania gracilis
Frullania nodulosa f. dapitana (Steph.) Verd. = Frullania nodulosa
Frullania nodulosa f. irreflexa Verd. = Frullania nodulosa
Frullania nodulosa β plana Schiffn. = Frullania nodulosa
Frullania picta Steph. = Frullania trichodes
Frullania replicata (Nees) Mont. = Frullania nodulosa
Frullania rugosa Mitt. = Frullania grandistipula
Frullania serrata f. crispulo-dentata Verd. = Frullania serrata var. serrata
Frullania squarrosa (Mont.) Nees = Frullania ericoides
Frullania squarrosa f. campanuloides Verd. = Frullania ericoides
Frullania squarrosa var. planescens Verd. = Frullania ericoides
Frullania tenuicaulis Mitt. = Frullania trichodes
Frullania thuillieri Nees = Frullania nodulosa
Gamoscyphus aselliformis (Reinw., Blume et Nees) Trevis. ≡ Heteroscyphus aselliformis
Gottschea aligera (Nees et Blume) Nees ≡ Schistochila aligera
Gottschea blumei (Nees) Nees ≡ Schistochila blumei
Harpalejeunea exigua Steph. = Lejeunea exilis
Harpalejeunea filicuspis (Steph.) Mizut. ≡ Microlejeunea filicuspis
Herbertus armitanus (Steph.) H.A.Mill. = Herbertus sendtneri
Herbertus divaricatus (Herzog) H.A.Mill. = Herbertus sendtneri
Herbertus javanicus (Steph.) H.A.Mill. = Herbertus ramosus
Herbertus javanicus (Steph.) Schiffn. nom. inval. = Herbertus ramosus
Heteroscyphus decurrens (Nees) Schiffn. = Heteroscyphus splendens
Hygrolejeunea amboinensis (Schiffn.) Schiffn. nom. inval. = Lejeunea sordida
Hygrolejeunea amboinensis (Schiffn.) Schiffn. ex P.Syd. = Lejeunea sordida
Hygrolejeunea decurvifolia Steph. = Lepidolejeunea bidentula
Hygrolejeunea discreta (Lindenb.) Schiffn. ≡ Lejeunea discreta
Hygrolejeunea discreta (Lindenb.) Steph. nom. inval. ≡ Lejeunea discreta
Hygrolejeunea parkinsonii Steph. = Lejeunea sordida
Hygrolejeunea sordida (Nees) Schiffn. ≡ Lejeunea sordida
Hygrolejeunea sordida (Nees) Steph. nom. inval. ≡ Lejeunea sordida
Hygrolejeunea umbilicata (Nees) Steph. nom. inval. ≡ Lejeunea umbilicata
Hymenophyton malaccense Steph. ≡ Podomitrium malaccense
Hymenophyton phyllanthus (Hook.) Steph. ≡ Podomitrium phyllanthus
Jamesoniella contracta (Reinw., Blume et Nees) N.Kitag. ≡ Cuspidatula contracta
Jamesoniella flexicaulis (Nees) Schiffn. ≡ Cuspidatula flexicaulis
Jamesoniella flexicaulis var. β microphylla (Nees) Schiffn. = Gottschelia schizopleura
Jamesoniella microphylla (Nees) Schiffn. = Gottschelia schizopleura
Jamesoniella ovifolia (Schiffn.) Schiffn. = Denotarisia linguifolia
Jungermannia ariadne Taylor ≡ Solenostoma ariadne
Jungermannia aselliformis Reinw., Blume et Nees ≡ Heteroscyphus aselliformis
Jungermannia comata Nees ≡ Solenostoma comatum
Jungermannia connata Sw. ≡ Cryptolophocolea connata
Jungermannia contracta Reinw., Blume et Nees ≡ Cuspidatula contracta
Jungermannia flexicaulis Nees ≡ Cuspidatula flexicaulis
Jungermannia flexicaulis β microphylla Gottsche, Lindenb. et Nees = Gottschelia schizopleura
Jungermannia gigantea Hook. nom. illeg. ≡ Plagiochila gigantea
Jungermannia gigantea β laxior Nees ≡ Plagiochila gigantea laxior
Jungermannia indica Nees = Lejeunea flava subsp. orientalis
Jungermannia opposita Reinw., Blume et Nees ≡ Chiastocaulon oppositum
Jungermannia ovifolia Schiffn. = Denotarisia linguifolia
Jungermannia simplex Weber nom. illeg. = Plagiochila trapezoidea
Jungermannia tetragona Lindenb. ≡ Solenostoma tetragonum
Jungermannia transversalis Sw. ≡ Symbiezidium transversale
Jungermannia truncata Nees ≡ Solenostoma truncatum
Jungermannia vaginata Sw. ≡ Frullania vaginata
Lejeunea applanata (Reinw., Blume et Nees) Nees ≡ Lopholejeunea applanata
Lejeunea catanduana (Steph.) H.A.Mill., Bonner et Bischl. = Lejeunea anisophylla
Lejeunea chalmersii (Steph.) Mizut. = Lejeunea microloba
Lejeunea connivens Gottsche = Cheilolejeunea ceylanica
Lejeunea cucullata (Reinw., Blume et Nees) Nees ≡ Metalejeunea cucullata
Lejeunea haskarliana (Gottsche) Spruce nom. illeg. ≡ Schiffneriolejeunea tumida var. haskarliana
Lejeunea heterophylla Sande Lac. = Cheilolejeunea trifaria
Lejeunea microstipula (Steph.) Prantl = Lejeunea exilis
Lejeunea paradoxa Schiffn. ≡ Calatholejeunea paradoxa
Lejeunea pusilla (K.I.Goebel) Schiffn. nom. illeg. = Cololejeunea metzgeriopsis
Lejeunea recurvistipula Gottsche = Caudalejeunea reniloba
Lejeunea rostrata var. α minor Schiffn. ex P.Geissler et Bischl. nom. inval. = Acrolejeunea pycnoclada subsp. pycnoclada
Lejeunea sagrana (Mont.) Gottsche, Lindenb. et Nees = Lopholejeunea subfusca
Lejeunea sagrana var. dentistipula Schiffn. = Caudalejeunea reniloba
Lejeunea superba var. α typica Schiffn. nom. inval. = Colura superba
Lejeunea ternatensis Gottsche ≡ Drepanolejeunea ternatensis
Lejeunea trapezia (Nees) Nees ≡ Cheilolejeunea trapezia
Lejeunea trifaria (Reinw., Blume et Nees) Nees ≡ Cheilolejeunea trifaria
Lepidozia papulosa Steph. ≡ Neolepidozia papulosa
Lepidozia rigida f. minor (Steph.) Herzog nom. inval. = Lepidozia rigida
Lepidozia rigida var. minor (Steph.) Herzog nom. inval. = Lepidozia rigida
Lepidozia wallichiana Gottsche ≡ Neolepidozia wallichiana
Leptolejeunea foliicola (Horik.) R.M.Schust. nom. illeg. ≡ Drepanolejeunea foliicola
Leptolejeunea schiffneri f. angustifolia Herzog nom. inval. = Leptolejeunea schiffneri
Leptolejeunea schiffneri var. genuina Herzog nom. inval. = Leptolejeunea schiffneri
Leptolejeunea schiffneri (Steph. ex Schiffn.) Steph. = Leptolejeunea schiffneri
Lobatiriccardia lobata (Schiffn.) Furuki = Lobatiriccardia coronopus
Lophocolea ciliolata (Nees) Gottsche ≡ Cryptolophocolea ciliolata
Lophocolea ernstiana (Steph.) N.Kitag. ≡ Chiloscyphus ernstianus
Lophocolea giulianettii Steph. = Cryptolophocolea costata
Lopholejeunea dentistipula (Schiffn.) Schiffn. = Caudalejeunea reniloba
Lopholejeunea eulopha (Taylor) Steph. nom. inval. ≡ Lopholejeunea eulopha
Lopholejeunea javanica (Nees) Schiffn. = Lopholejeunea nigricans
Lopholejeunea pyriflora Steph. = Lopholejeunea subfusca
Lopholejeunea sagrana (Mont.) Steph. = Lopholejeunea subfusca
Lopholejeunea sagrana var. dentistipula (Schiffn.) Schiffn. ex P.Syd. = Caudalejeunea reniloba
Lopholejeunea sagrana var. dentistipula Schiffn. nom. inval. = Caudalejeunea reniloba
Lopholejeunea sagrana var. β subfusca (Nees) Schiffn. ≡ Lopholejeunea subfusca
Madotheca acutifolia Lehm. et Lindenb. ≡ Porella acutifolia
Madotheca crenilobula Herzog = Porella javanica
Marchantia amboinensis Nees et Mont. = Marchantia emarginata subsp. emarginata
Mastigobryum amboinense Steph. = Bazzania tridens
Mastigobryum asymmetricum Steph. ≡ Bazzania asymmetrica
Mastigobryum ceylanicum Mitt. ≡ Bazzania ceylanica
Mastigobryum concinnum De Not. = Bazzania intermedia
Mastigobryum crassitextum (Steph.) Steph. ≡ Bazzania crassitexta
Mastigobryum deningeri Herzog = Bazzania longicaulis
Mastigobryum echinatiforme De Not. ≡ Acromastigum echinatiforme
Mastigobryum echinatum Gottsche ≡ Acromastigum echinatum
Mastigobryum fallax Sande Lac. ≡ Bazzania fallax
Mastigobryum horridulum (Schiffn.) Steph. ≡ Bazzania horridula
Mastigobryum insigne De Not. ≡ Bazzania insignis
Mastigobryum intermedium Gottsche et Lindenb. ≡ Bazzania intermedia
Mastigobryum irregulare Steph. ≡ Bazzania irregularis
Mastigobryum javanicum Sande Lac. ≡ Bazzania javanica
Mastigobryum longidens Steph. = Bazzania conophylla
Mastigobryum merrillanum Steph. ≡ Bazzania merrillana
Mastigobryum pectinatum (Nees) Lindenb. et Gottsche = Bazzania pectinata
Mastigobryum stresemannii Herzog ≡ Bazzania stresemannii
Mastigobryum subtile Sande Lac. ≡ Bazzania subtilis
Mastigobryum tridens (Reinw., Blume et Nees) Dumort. ≡ Bazzania tridens
Mastigobryum uncigerum (Reinw., Blume et Nees) Nees ≡ Bazzania uncigera
Mastigobryum vittatum Gottsche ≡ Bazzania vittata
Mastigobryum wallichianum Lindenb. ≡ Bazzania wallichiana
Mastigolejeunea amboinensis (Schiffn.) Schiffn. ex P.Syd. = Thysananthus spathulistipus
Mastigolejeunea amboinensis Schiffn. nom. inval. = Thysananthus spathulistipus
Mastigolejeunea amboinensis var. β paucidentata Schiffn. nom. inval. = Thysananthus spathulistipus
Mastigolejeunea atypos Schiffn. ex Sydow = Thysananthus repletus
Mastigolejeunea auriculata (Wilson et Hook.) Steph. ≡ Thysananthus auriculatus
Mastigolejeunea humilis (Gottsche) Schiffn. ≡ Thysananthus humilis
Mastigolejeunea ligulata (Lehm. et Lindenb.) Schiffn. ≡ Thysananthus ligulatus
Mastigolejeunea truncata Mizut. ≡ Thysananthus truncatus
Mastigolejeunea undulata Gradst. et Grolle = Thysananthus frauenfeldii
Mastigolejeunea virens (Ångstr.) Steph. ≡ Thysananthus virens
Mastigophora diclados var. β scorpioides (Reinw., Blume et Nees) Schiffn. = Mastigophora diclados
Mastigophora ramentifissa Herzog = Mastigophora woodsii
Mastigophora vrieseana (Sande Lac.) Schiffn. = Mastigophora diclados
Megaceros grandis (Ångstr.) Steph. = Megaceros flagellaris
Metahygrobiella stolonacea (Herzog) R.M.Schust. ≡ Cephalozia stolonacea
Metzgeria decipiens (C.Massal.) Schiffn. = Metzgeria ciliata
Metzgeria hamatiformis Schiffn. = Metzgeria leptoneura var. leptoneura
Metzgeriopsis pusilla K.I.Goebel = Cololejeunea metzgeriopsis
Microlejeunea microstipula Steph. = Lejeunea exilis
Microlejeunea parallela Schiffn. = Metalejeunea cucullata
Microlejeunea ternatensis (Gottsche) Herzog nom. inval. ≡ Drepanolejeunea ternatensis
Microlepidozia gonyotricha (Sande Lac.) Del Ros. ≡ Kurzia gonyotricha
Nardia ariadne (Taylor) Schiffn. ≡ Solenostoma ariadne
Nardia comata (Nees) Schiffn. ≡ Solenostoma comatum
Noguchia opposita (Reinw., Blume et Nees) Inoue ≡ Chiastocaulon oppositum
Omphalanthus umbilicatus (Nees) Nees ≡ Lejeunea umbilicata
Otolejeunea semperiana (Steph.) Grolle ≡ Allorgella semperiana
Paraschistochila neesii (Mont.) R.M.Schust. ≡ Schistochila neesii
Phaeoceros hirticalyx (Steph.) J.Haseg. ≡ Phaeomegaceros hirticalyx
Phaeoceros laevis subsp. carolinianus (Michx.) Prosk. ≡ Phaeoceros carolinianus
Phaeoceros polyandrus (Steph.) J.Haseg. = Phaeomegaceros hirticalyx
Phragmicoma fertilis (Reinw., Blume et Nees) Nees ex Mont. ≡ Acrolejeunea fertilis
Phragmicoma ligulata (Lehm. et Lindenb.) Gottsche, Lindenb. et Nees ≡ Thysananthus ligulatus
Phragmicoma reniloba Gottsche ≡ Caudalejeunea reniloba
Physocolea ciliatilobula (Schiffn.) Steph. = Cololejeunea inflectens
Physocolea hamata (Steph.) Steph. ≡ Cololejeunea hamata
Plagiochila acanthophylla Gottsche = Plagiochila sciophila
Plagiochila bantamensis var. γ denticulata Sande Lac. = Plagiochila kurzii
Plagiochila batjanensis Schiffn. ex Herzog = Plagiochila teysmannii
Plagiochila belangeriana Lindenb. = Plagiochila arbuscula
Plagiochila blepharophora var. major (Schiffn.) Bonner = Plagiochila sandei
Plagiochila brauniana (Nees) Lindenb. ≡ Chiastocaulon braunianum
Plagiochila ceramica Inoue = Plagiochila pulvinata
Plagiochila comata (Nees) Lindenb. ≡ Solenostoma comatum
Plagiochila decidua Inoue et Grolle = Plagiochila sciophila
Plagiochila dendroides (Nees) Lindenb. ≡ Chiastocaulon dendroides
Plagiochila frondescens var. tenerrima (Nees) Schiffn. = Plagiochila frondescens
Plagiochila frondescens α tenerrima (Nees) Lindenb. = Plagiochila frondescens
Plagiochila frondescens γ rigida (Nees) Lindenb. = Plagiochila frondescens
Plagiochila gedeana Schiffn. = Plagiochila hampeana
Plagiochila novae-guineae Sande Lac. = Plagiochila chauviniana
Plagiochila opposita (Reinw., Blume et Nees) Lindenb. ≡ Chiastocaulon oppositum
Plagiochila opposita var. β falcata (Nees) Schiffn. ≡ Plagiochila conjugata var. falcata
Plagiochila opposita var. γ filiformis (Lindenb.) Schiffn. = Chiastocaulon oppositum
Plagiochila opposita β falcata (Nees) Lindenb. ≡ Plagiochila conjugata var. falcata
Plagiochila opposita γ filiformis Lindenb. nom. inval. = Chiastocaulon oppositum
Plagiochila pseudaberrans Inoue et Grolle = Plagiochila gymnoclada
Plagiochila seemannii Mitt. = Plagiochila sandei
Plagiochila semialata Sande Lac. = Plagiochila teysmannii
Plagiochila singularis Schiffn. ≡ Dinckleria singularis
Plagiochila trapezoidea var. β tenera (Nees) Schiffn. = Dinckleria singularis
Plagiochila trapezoidea var. γ major (Lindenb.) Schiffn. = Plagiochila gigantea laxior
Plagiochila trapezoidea β tenera (Nees) Lindenb. = Dinckleria singularis
Plagiochila trapezoidea γ major Lindenb. nom. illeg. = Plagiochila gigantea laxior
Plagiochila variegata Lindenb. = Syzygiella securifolia
Plagiochilion braunianum (Nees) S.Hatt. ≡ Chiastocaulon braunianum
Plagiochilion fimbriatum (Mitt.) Inoue ≡ Chiastocaulon fimbriatum
Plagiochilion oppositum (Reinw., Blume et Nees) S.Hatt. ≡ Chiastocaulon oppositum
Plectocolea ariadne (Taylor) Mitt. ≡ Solenostoma ariadne
Plectocolea comata (Nees) S.Hatt. ≡ Solenostoma comatum
Porella acutifolia var. subligulifera S.Hatt. = Porella acutifolia var. acutifolia
Psiloclada unguligera Schiffn. = Psiloclada clandestina subsp. clandestina
Ptychanthus javanicus Nees = Ptychanthus striatus var. striatus
Ptychanthus moluccensis Sande Lac. = Ptychanthus striatus var. striatus
Ptychanthus squarrosus Mont. = Ptychanthus striatus
Ptychocoleus brunneus Steph. = Acrolejeunea pycnoclada subsp. pycnoclada
Ptychocoleus cumingianus (Mont.) Trevis. ≡ Schiffneriolejeunea cumingiana
Ptychocoleus densifolius (Schiffn. ex P.Syd.) Steph. nom. illeg. = Schiffneriolejeunea pulopenangensis
Ptychocoleus haskarlianus (Gottsche) Steph. ≡ Schiffneriolejeunea tumida var. haskarliana
Ptychocoleus pulopenangensis (Gottsche) Trevis. ≡ Schiffneriolejeunea pulopenangensis
Ptychocoleus pycnocladus (Taylor) Steph. ≡ Acrolejeunea pycnoclada
Ptychocoleus tener Steph. = Acrolejeunea fertilis
Pycnolejeunea badia Steph. = Lepidolejeunea bidentula
Pycnolejeunea ceylanica (Gottsche) Schiffn. ≡ Cheilolejeunea ceylanica
Pycnolejeunea ceylanica (Gottsche) Steph. nom. inval. ≡ Cheilolejeunea ceylanica
Pycnolejeunea connivens (Gottsche) Gottsche ex Schiffn. nom. inval. = Pycnolejeunea connivens
Pycnolejeunea decurvifolia (Steph.) Steph. = Lepidolejeunea bidentula
Pycnolejeunea demissa Steph. ex Kachroo et R.M.Schust. nom. inval. = Cheilolejeunea longidens
Pycnolejeunea falsinervis (Sande Lac.) Schiffn. ≡ Cheilolejeunea falsinervis
Pycnolejeunea gigantea Steph. ≡ Cheilolejeunea gigantea
Pycnolejeunea incisa (Gottsche) Schiffn. ≡ Cheilolejeunea incisa
Pycnolejeunea meyeniana (Nees, Lindenb. et Gottsche) Steph. nom. inval. = Cheilolejeunea trapezia
Pycnolejeunea schwaneckei Steph. nom. inval. = Pycnolejeunea schwaneckei
Pycnolejeunea trapezia (Nees) Schiffn. ≡ Cheilolejeunea trapezia
Pycnolejeunea ventricosa (Schiffn.) Schiffn. ex P.Syd. = Cheilolejeunea ventricosa
Pycnolejeunea ventricosa Schiffn. nom. inval. = Cheilolejeunea ventricosa
Radula campanigera Mont. ≡ Cladoradula campanigera
Radula ceramensis Steph. = Radula javanica
Radula formosa var. pycnolejeuneoides (Schiffn.) N.Kitag. = Radula formosa
Radula ovalifolia Steph. = Radula javanica
Radula pycnolejeuneoides Schiffn. = Radula formosa
Radula subsimilis Steph. = Radula reflexa
Radula wallichiana Lehm. = Radula javanica
Rhaphidolejeunea longicruris (Steph.) Herzog ≡ Drepanolejeunea longicruris
Riccardia pinguis (L.) Gray ≡ Aneura pinguis
Saccogyna rigidula (Nees) Schiffn. ≡ Saccogynidium rigidulum
Saccogynidium jugatum (Mitt.) Grolle = Saccogynidium rigidulum
Schiffneriolejeunea haskarliana (Gottsche) Gradst. ≡ Schiffneriolejeunea tumida var. haskarliana
Schisma divaricatum Herzog = Herbertus sendtneri
Schisma pilifera (Schiffn.) Steph. = Herbertus pilifer
Schisma piligerum Steph. nom. inval. = Herbertus pilifer
Schistochila amboinensis Steph. = Schistochila beccariana
Schistochila formosana Horik. = Schistochila blumei
Schistochila graeffeana J.B.Jack et Steph. = Schistochila aligera
Schistochila inversa Herzog = Schistochila sciurea
Schistochila philippinensis (Mont.) J.B.Jack et Steph. nom. inval. = Schistochila aligera
Schistochila philippinensis (Mont.) Steph. = Schistochila aligera
Schistochila philippinensis var. transiens H.Buch nom. inval. = Schistochila aligera
Schistochila purpurascens Herzog = Schistochila acuminata
Schistochila recurvata H.Buch nom. inval. = Schistochila aligera
Schistochila sciurea f. robustior Schiffn. = Schistochila sciurea
Schistochilaster aliger (Nees et Blume) H.A.Mill. ≡ Schistochila aligera
Schistochilaster graeffeanus (J.B.Jack et Steph.) H.A.Mill. = Schistochila aligera
Schistochilaster philippinensis (Mont.) H.A.Mill. = Schistochila aligera
Sendtnera diclados α scorpioides (Reinw., Blume et Nees) Nees = Mastigophora diclados
Sendtnera diclados β calcarata (Reinw., Blume et Nees) Gottsche, Lindenb. et Nees nom. inval. = Mastigophora diclados
Sendtnera vrieseana Sande Lac. = Mastigophora diclados
Spruceanthus marianus (Gottsche) Mizut. ≡ Cheilolejeunea mariana
Stenolejeunea dentata (Steph.) Pócs ≡ Lejeunea stenodentata
Strepsilejeunea cavistipula Steph. ≡ Pycnolejeunea cavistipula
Symphyogynopsis filicum (Nadeaud) Grolle = Symphyogynopsis gottscheana
Syzygiella variegata (Lindenb.) Spruce = Syzygiella securifolia
Taxilejeunea albescens Steph. ≡ Lejeunea albescens
Taxilejeunea karstenii Steph. = Lejeunea lumbricoides
Taxilejeunea laxiretis (Steph.) Eifrig = Lejeunea sordida
Taxilejeunea lumbricoides (Nees) Steph. ≡ Lejeunea lumbricoides
Taxilejeunea sordida (Nees) Eifrig ≡ Lejeunea sordida
Telaranea cuneifolia (Steph.) J.J.Engel et G.L.Merr. ≡ Neolepidozia cuneifolia
Telaranea kogiana (Steph.) Grolle ≡ Tricholepidozia kogiana
Telaranea neesii (Lindenb.) Fulford ≡ Tricholepidozia neesii
Telaranea wallichiana (Gottsche) R.M.Schust. ≡ Neolepidozia wallichiana
Thysananthus renilobus (Gottsche) Schiffn. ≡