Research Article |
Corresponding author: Michael D. Pirie ( michael.pirie@uib.no ) Academic editor: Jaime Fagúndez
© 2023 E. Charles Nelson, E. G. H. Oliver, Michael D. Pirie.
This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0), which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
Citation:
Nelson EC, Oliver EGH, Pirie MD (2023) Erica L. (Ericaceae): homonyms amongst published names for African species and proposed replacement names. PhytoKeys 236: 157-178. https://doi.org/10.3897/phytokeys.236.110498
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In support of ongoing taxonomic work on the large and complex flowering plant genus Erica (Ericaceae), we document nineteen pairs of homonyms representing currently used illegitimate names. We provide replacements for thirteen names and new typifications for five. We relegate five names to synonymy: Erica aemula Guthrie & Bolus under Erica distorta Bartl.; Erica armata Klotzsch ex Benth. under Erica umbrosa H. A. Baker; Erica capensis T.M. Salter under Erica turbiniflora Salisb.; Erica lanata Andrews under Erica flaccida Link; and Erica tomentosa Salisb. under Erica velutina Bartl. Finally, we suggest conservation of Erica aristata Andrews. The new names are: Erica adelopetala E.C. Nelson & E.G.H. Oliv. replacing Erica insignis E.G.H. Oliv.; Erica bombycina E.C. Nelson & Pirie replacing Erica niveniana E.G.H. Oliv.; Erica concordia E.C. Nelson & E.G.H. Oliv. replacing Erica constantia Nois. ex Benth.; Erica didymocarpa E.C. Nelson & E.G.H. Oliv. replacing Erica rugata E.G.H. Oliv.; Erica galantha E.C. Nelson & E.G.H. Oliv. replacing Erica perlata Benth.; Erica mallotocalyx E.C. Nelson & E.G.H. Oliv. replacing Erica flocciflora Benth.; Erica notoporina E.C. Nelson & E.G.H. Oliv. replacing E. autumnalis L.Bolus; Erica oliveranthus E.C. Nelson & Pirie replacing Erica tenuis Salisb.; Erica oraria E.C. Nelson & E.G.H. Oliv. replacing Erica spectabilis Klotzsch ex Benth.; Erica oresbia E.C. Nelson & E.G.H. Oliv. replacing Erica demissa Klotzsch ex Benth.; Erica poculiflora E.C. Nelson & E.G.H. Oliv. replacing Erica stenantha Klotzsch ex Benth.; Erica rhodella E.C. Nelson & E.G.H. Oliv. replacing Erica rhodantha Guthrie & Bolus; Erica supranubia E.C. Nelson & Pirie replacing Erica praecox Klotzsch.
Cape heaths, Erica, Hans Dulfer, International register of heather names, nineteenth-century English nursery catalogues, nomenclature
The nomenclatural history of the genus Erica L. sensu lato (
Work carried out between 1995 and 2004 by ECN for The Heather Society (established in 1963, formally dissolved in 2020) on the second volume of the International Register of Heather Names (
Some of these problems do not arise, in fact, from the unearthing of long-buried names (most of the binomials discussed here were recorded by
Although changes in current names are rarely welcomed, particularly outside the taxonomic community, by following the rules set out in the International Code of Nomenclature for Algae, Fungi and Plants (Shenzhen Code) 2018 (
The Heather Society’s database revealed pairs of homonyms within Erica where the one in current use (compare
Specific epithet | Author(s) and date of currently accepted binomial | Author(s) and date(s) of earlier binomial |
---|---|---|
aemula |
|
Rollisson (1855) |
aristata |
|
|
armata | Klotzsch ex |
|
autumnalis | L. |
Hort. ex |
capensis |
|
|
constantia | Noisette ex |
Hort. ex |
demissa | Klotzsch ex |
Hort. ex |
flocciflora | Bentham (December 1839) | Tausch (October 1839) |
insignis |
|
Hort. (1853) |
lanata |
|
|
niveniana |
|
Hort. ex |
perlata |
|
|
praecox |
|
Hort. ex |
rhodantha |
|
|
rugata |
|
Hort. ex |
spectabilis | Klotzsch ex |
|
stenantha | Klotzsch ex |
|
tenuis |
|
|
tomentosa |
|
|
It should be noted that nurserymen’s catalogues during the 19th and 20th centuries were usually annual publications, reissued with minor amendments, deletions and additions, year after year, decade after decade. The dated catalogues noted in this paper are examples only – they must not be assumed to be the first, the last or the only issue containing a particular name.
This binomial first appeared in print within an advertisement inserted by Messrs William Rollisson & Sons of Tooting, London, in “The Gardeners’ Chronicle”: 218 (7 April 1855). A brief description, noting the “fine bright crimson” flowers, was included that validates the name. Subsequently, it was published in several other English nurserymen’s catalogues (see Nelson and Small (2004: pt 1: 11)): for example, in William Rollisson and Sons’ catalogue for 1877, which company claimed it as a hybrid raised and sent out by them; James Fraser, Lea Bridge Road Nursery, for 1866–1867; E. G. Henderson and Son for Autumn 1871; James Veitch and Sons for 1873–1874; and B. S. Williams for 1881. Compilers of gardening dictionaries soon included this heath in their works including
Erica aemula Guthrie & Bolus, Fl. Capensis 4,1: 118. 1905 [wfo-0000671313], nom. illeg., non Erica aemula Rollisson, Gard. Chron. 1855: 218 (1855) [wfo-1200040659]. Type. South Africa. “Bains Kloof“ [Bainskloof], Cummings 171 BOL [BOL137142 [https://plants.jstor.org/stable/10.5555/al.ap.specimen.bol137142], syntype; “Fish Hoek, Gordon’s Bay“, Guthrie 3108 BOL (BOL137141) [https://plants.jstor.org/stable/10.5555/al.ap.specimen.bol137141], syntype.
South Africa, “Auf Felsenrücken in der Kluft nach der Platte des Tafelberges in vierter Höhe” (lectotype †GOET destroyed, fide annotation by N. E. Brown on K000314148; isolectotype [fragments only], “ad Cap. b. Spei in monte tabulari leg. Ecklon. Hb. Bg. 1841” K [K000314148].
Richard Anthony Salisbury’s (1761–1829) catalogue (
A decade later the same binomial was employed by Henry Cranke Andrews (fl. 1794–1830) when he illustrated and described the plant that currently bears this name, an inhabitant of the Kleinrivier Mountains (
This species was illustrated by
Erica armata
was validly published by
E. armata Klotzsch ex Benth., Prodr. [A. P. de Candolle] 7(2): 672. 1839 [wfo-0000671419], nom. illeg., non Spreng., Syst. Veg. 2: 184. 1825 [wfo-0000671418]; Guthrie and Bolus, Fl. Capensis 4,1: 113. 1905; Dulfer, Ann. Naturhist. Mus, Wein 68: 67. 1965; Oliver, Strelitzia 29: 489. 2012. Type. South Africa. “in herb. reg. Berol. … in montibus prov. Worcester et Stellenbosch”, Masson, Niven, Drège (lectotypes B destroyed; isolectotype K [Niven 158] (det. E.G.H. Oliver) K000314208).
Specimens in other herbaria collected by Masson and Drège are variously labelled as syntypes or isosyntypes, but their identity has not been confirmed: GDC (Masson, F., s.n., G00494351 https://plants.jstor.org/stable/10.5555/al.ap.specimen.g00494351); GDC (Drège, J.F., s.n., G00494352 https://plants.jstor.org/stable/10.5555/al.ap.specimen.g00494352); HBG (Drège, J.F., s.n., HBG515307 https://plants.jstor.org/stable/10.5555/al.ap.specimen.hbg515307); TUB (Drège, J.F., s.n., TUB003182 left-hand specimen only https://plants.jstor.org/stable/10.5555/al.ap.specimen.tub003182); S (Drège, J.F., s.n., S08-5237 https://plants.jstor.org/stable/10.5555/al.ap.specimen.s08-5237).
South Africa. “Caledon, Elandskloof, Villiersdorp, on a steep S-facing slope in shade in kloof with a large waterfall, 3 April 1961, E.G.H. Oliver 1423 (holotype: BOL; isotype: NBG-0199392-1 [https://plants.jstor.org/stable/10.5555/al.ap.specimen.nbg0199392-1].
This binomial was published under the entry for Erica formosa Thunb. with a validating diagnostic phrase by
A new name is required for the taxon and the new epithet alludes to the autumn, which is the species’ main, Southern-Hemisphere flowering season.
pro Erica autumnalis L.Bolus, Ann. Bolus Herb. 3: 178. 1923 [wfo-0000671451], nom illeg., non E. × autumnalisHort. ex Benth., Prodr. [A. P. de Candolle] 7(2): 659 (1839) [wfo-1000053512]; Regel, Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten 16: 307. 1842; Regel, Kult. Aufz. Eriken, 147. 1843; Dulfer, Ann. Naturhist. Mus, Wein 68: 151 (1965).
South Africa, “Bought in Adderly St., Cape Town”, 2 February 1922, N.S. Pillans 16784: (holotype: BOL [BOL-137249 (https://plants.jstor.org/stable/10.5555/al.ap.specimen.bol137249); isotype BOL [BOL-137250]).
In his monograph on Erica, Eduard August von Regel (1815–1892) described a plant named E. capensis (
Erica capensis T.M.Salter, J. S. Afr. Bot. 1: 34. 1935 [wfo-0000671621], nom. Illeg., non Regel, Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten 16: 318 (1842) [wfo-1000053513], 158. 1843. Type: South Africa. “Cape Peninsula, marshes on lower Hout and Klaasjager River”, 14 February 1934, T.M. Salter 4292 (holotype: BOL-137252 [https://plants.jstor.org/stable/10.5555/al.ap.specimen.bol137252]).
Without locality, Hibbert ex herb. Salisbury (lectotype, designated here: K [K-314663]).
A Cape heath named “Erica Constantia” was included by Messrs Lee & Kennedy in a manuscript list of species that had been introduced into cultivation by the firm up to 1808 (see
However,
Erica constantia
Nois. ex Benth. is currently applied to a white-flowered heath from rocky slopes at high altitude, ranging from the Hex River Mountains to Klein Swartberg (
Here, we designate the Esterhuysen collection cited by
pro E. constantia Nois. ex Benth., Prodr. [A. P. de Candolle] 7(2): 672. 1839 [wfo-0000671735], nom. illeg., non Hort. ex G.Sinclair, Hort. eric. woburn.: 6, 32. 1825 [wfo-1000055091]; Guthrie and Bolus, Fl. Capensis 4,1: 115–116. 1905; Dulfer, Ann. Naturhist. Mus, Wein 68: 67–68. 1965.
Without locality or collector, Herb. Klotzsch (holotype: B, destroyed). South Africa, Audensberg, S slopes. 15 February 1959, E.E. Esterhuysen 28188 (neotype, designated here, NBG [NBG-0265661-0]).
“Dwarf green-flowered heath” was the English name used by
Sinclair’s description (1825) is more than adequate to validate the name, which
Meanwhile,
A new epithet, derived from the Greek compound ορεσβιος (oresbios) meaning living on mountains (
pro E. demissa Klotzsch ex Benth., Prodr. [A. P. de Candolle] 7(2): 666. 1839 [wfo-0000671862], nom illeg., non Hort. ex G. Sinclair Hort. eric. woburn.: 8. 1825 [wfo-0000671861]; Benth., Prodr. [A. P. de Candolle] 7(2): 621. 1839; Guthrie and Bolus, Fl. Capensis 4,1: 47. 1905; Dulfer, Ann. Naturhist. Mus, Wein 68: 86–87. 1965.
South Africa. “Uitenhaag [Uitenhage], in monitbus Vanstadensrivier”, 1000–4000[ft], C.F. Ecklon and C.L.P. Zeyher s.n. (syntype: not traced); South Africa, “flum. Camtoo [Gamtoos]”, Masson s.n. (syntype: not traced), Burchell 4709 (syntype: HAL [https://plants.jstor.org/stable/10.5555/al.ap.specimen.hal0135770]).
This is an instance of two names published within weeks of each other. Tausch’s binomial was published on 28 October 1839, more than a month before Bentham’s and, thus, has priority. Unfortunately,
Given the fact he had an incorrect (earlier) date of publication,
Erica flocciflora
, as described by Bentham, occurs on the dry, lower slopes and rocky foothills of the Kouga Mountains and has a cream corolla with a distinctive woolly calyx (
Irrespective of synonymy, Bentham’s binomial is illegitimate because it is a later homonym. The new epithet that we provide below continues the allusion to floccus (Latin: tuft of woolly hairs) by adopting a Greek equivalent, μαλλωτoς (mallotos), fleecy.
pro E. flocciflora Benth., Prodr. [A. P. de Candolle] 7(2): 660. 1839 (late December) [wfo-0000672068], nom. illeg., non Tausch, Flora Bot. Zeit. Regensb. 22: 629. 1839 (28 October) [wfo-0000672067]; Guthrie and Bolus, Fl. Capensis 4,1: 310 (1905); Dulfer, Ann. Naturhist. Mus, Wein 68: 98 (1965).
South Africa. “In colonia capensi” (cit.
This binomial, overlooked by the compilers of “Index Kewensis”, has been traced in no fewer than six publications issued during the latter half of the 19th century including the “Journal of the Royal Horticultural Society” (1853: 8: xl) and “Revue horticole” (1882: 54: 219–220). Accompanying descriptions indicated it was applied to a scarlet-blossomed heath. It was also traced in catalogues issued by the following British and New Zealand nurseries: James Fraser, Lea Bridge Road Nursery for 1866–1867; William Rollisson & Sons for 1877; B. S. Williams for 1881; James Dickson & Sons, Newton Nurseries, for 1884; Nairn & Sons, Christchurch (New Zealand) for 1896. It is highly improbable that the plant cultivated at least until the end of the 19th century in European and New Zealand gardens was the same species as that first collected by Stokoe in 1935 and described by
The new epithet alludes to the concealed corolla (from Greek: αδελoς (adelos = unseen), πεταλov (petalon = leaf, i.e. petal)) and echoes the name of the section to which this species and E. nabea Guthrie and Bolus were assigned.
pro E. insignis E.G.H. Oliv., Bothalia 13: 446. 1981 [wfo-0000672329], nom. illeg., non hort. in J. Roy. Hort. Soc. 8: xl. 1853 [wfo-1000053515]; Rev. hort. 54: 219–220. 1882.
South Africa, “Swartberg, north slopes below Kangoberg”, 1,400 m, 11 December 1979, E.G.H. Oliver 7469 (holotype, NBG [as STE] [https://plants.jstor.org/stable/10.5555/al.ap.specimen.nbg0133788-0]; isotypes K, PRE).
Messrs Lee & Kennedy had employed the name “Erica lanata” in the manuscript list, mentioned previously (see
Erica lanata
Andrews, Heathery, 3: t. 121 (1806); Col. engr. heaths, 3, t. 179. 1809b [wfo-0000672411], nom. illeg., non E. lanata J.C. Wendl., Bot. Beobach.: 45. 1798 [wfo-0000672410]; Salisb., Trans. Linn. Soc. 6: 360 (1802); Guthrie and Bolus, Fl. Capensis 4,1: 73. 1905; Dulfer, Ann. Naturhist. Mus, Wein 68: 112 (1965); Oliver, Strelitzia 29: 499. 2012. Type. Icontype (illustration in Andrews, Heathery, 3: t. 121. 1806 [cit.
South Africa. “Hab. in Pr. b. sp. [Promontorium Bonae Spei = Cape of Good Hope] … Hort. angl.” (B destroyed). Neotype (here designated). South Africa. Western Cape, George Dist., Outeniqua Pass. June 1960. E.G.H. Oliver 1596 NBG [NBG0112414-0].
This binomial appeared in print two centuries ago (
The new name alludes to the silky hairs that give the inflorescences a fluffy appearance (
pro E. niveniana E.G.H.Oliv., Contrib. Bolus Herb. 19: 225. 2000 [wfo-0000672716], nom. Illeg., non E. niveniiHort. Ex Loudon (as “nivenia”), Hort. Brit.: 147. 1830 [wfo-1000053516] (see
South Africa. Without locality [“Erica N95 on elevated Situations”], J. Niven 95 (holotype K (K-000225736 [https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:1017329-1]).
It appears that no-one has hitherto noticed the inconsistency pertaining to the application of the binomial Erica perlata (meaning “beset with pearls”) (
Fol. 4 | Bractea. rem. | Anth. sub. ex. mut. | Pistill. ex. | Inflorescentia term. br. umbel.; corol. glob. | Color. Corol. R.P.1-3. Anthers R.R.O.7. | T. Flor. Spring, Autumn. |
In other words: leaves ternate; bracts remote; anthers subulate, exserted, muticous; pistil exserted, inflorescences terminal, branching, umbellate; corolla globose, pink-madder; anthers red (between scarlet and Indian red); flowering [in Northern Hemisphere] spring and autumn.
No herbarium specimens from the Duke of Bedford’s Woburn collection, the basis of Sinclair’s descriptions, are known that could be considered as type material, so the protologue is paramount. At least the colour of the flower signals that the protologue does not match the current application of the binomial. Sinclair’s colour codes were very precise, being determined using a specially constructed “diagram of colours” (a colour-wheel) (
According to
We propose the replacement name Erica galantha for the Riviersonderend species, in allusion to the white, pearl-like flowers. Galanthos, from γάλα (gala = milk) and ἄνθος (anthos = flower), means with a milk-white flower (as in Galanthus L., Amaryllidaceae, the Eurasian snowdrop).
pro E. perlata Benth., Prodr. [A. P. de Candolle] 7(2): 670. 1839 [wfo-1000053517], nom. Illeg., non E. perlata G.Sinclair, Hort. Eric. woburn.: 18 (1825) [wfo-0000672873].
South Africa, “In colonia capensi”, Drège s.n. (lectotype W, effectively designated by
This binomial was printed, without accompanying descriptions, in Conrad Loddiges & Sons’ catalogue for 1807 and in at least two other pre-1825 publications (Anonymous 1808: 195;
Our replacement epithet refers to the species’ occurrence high in the mountains – above the clouds.
pro E. praecox Klotzsch, Linnaea 12: 517. 1838 [wfo-0000672980], nom. illeg., non Hort. ex G.Sinclair, Hort. eric. woburn.: 19, 32. 1825 [wfo-1200010024]; Benth., Prodr. [A. P. de Candolle] 7(2): 678. 1839.
South Africa. “Dutoitskloof”, Drège 1147 (lectotype P (P-00110863), designated here (det. E.G.H. Oliver) [https://plants.jstor.org/stable/10.5555/al.ap.specimen.p00110863]; isolectotype W [https://plants.jstor.org/stable/10.5555/al.ap.specimen.w18890186298, https://plants.jstor.org/stable/10.5555/al.ap.specimen.w18890158225].
The plant described and named by
The new epithet continues the allusion to the rose-red (rhodellus) flowers.
pro E. rhodantha Guthrie and Bolus, Fl. Capensis 4,1: 288. 1905 [wfo-0000673111], nom. illeg., non Regel, Verh. Vereins Beförd. Gartenbaues Königl. Preuss. Staaten 16: 318. 1842 [wfo-1000053518], Regel, Kult. Aufz. Eriken, 158 (1843); Dulfer, Ann. Naturhist. Mus, Wein 68: 154. 1965.
South Africa. Riversdale Div.; Garcias Pass, 1200 ft, Galpin 3706 (lectotype BOL, effectively designated by
This binomial was printed, without accompanying descriptions, in Conrad Loddiges & Sons’ catalogue for 1811 and in at least two other pre-1825 publications (
Unaware of the previous use of this binomial – it is not listed in botanical indexes such as “Index Kewensis”, nor was it noted by
pro E. rugata E.G.H.Oliv., Contrib. Bolus Herb. 19: 368 (2000) [wfo-0000673157], nom. illeg., non Hort. ex G.Sinclair, Hort. eric. woburn.: 22 (1825) [wfo-1000053519] (= E. × rugosa Andrews [wfo-0000673159]).
South Africa. “Cap, im Gebirge bei der Kapstadt” [mountains near Cape Town] [loc. 84], Zeyher s.n. (lectotype K, designated by
The name Erica spectabilis appeared in print more than thirty years before its publication by
As the variant “spectabilia”, the binomial has been traced in the 1804 catalogue issued by Conrad Loddiges & Sons who corrected the spelling to “spectabilis” in 1818.
The handsome, variable, white-, cream- to green-flowered species for which
The new epithet, from the Latin ora (edge or sea coast), reflects the coastal distribution of the species.
pro E. spectabilis Klotzsch ex Benth., Prodr. [A. P. de Candolle] 7(2): 659. 1839 [wfo-0000673283], nom. illeg., non Waitz, Beschreibung der Gattung und Arten der Heiden: 220. 1805 [wfo-0000673282]; Guthrie and Bolus, Fl. Capensis 4,1: 57. 1905.
South Africa. “in Strandweld [Strandveld]”, Drège s.n. (syntypes: †B, GDC [https://plants.jstor.org/stable/10.5555/al.ap.specimen.g00465165], HBG [https://plants.jstor.org/stable/10.5555/al.ap.specimen.hbg507932], K, W [https://plants.jstor.org/stable/10.5555/al.ap.specimen.w18890186305, https://plants.jstor.org/stable/10.5555/al.ap.specimen.w18890321973, https://plants.jstor.org/stable/10.5555/al.ap.specimen.w0005951]).
As noted by
The new epithet is derived from Latin poculus (cup) and alludes to the cup-shaped flowers.
pro E. stenantha Benth., Prodr. [A. P. de Candolle] 7(2): 685. 1839 [wfo-0000673321], nom. illeg., non Sweet, Hort. Brit.: 340. 1830 [wfo-0000673320] (= E. cylindrica Thunb.).
South Africa. “Berge bei Zwellendam“, C.F.Ecklon and C.L.P. Zeyher 221 (lectotype W, effectively designated by
According to
Mysteriously,
Erica tenuis
Salisb. is the current name for a white-flowered heath (Fig.
Erica oliveranthus E.C. Nelson and Pirie, renamed in honour of E.G.H. (Ted) Oliver and Inge M. Oliver (photo: MDP; https://www.inaturalist.org/observations/19098927).
The new epithet published here is a tribute to our co-author, Dr E.G.H. (‘Ted’) Oliver, pre-eminent authority on the taxonomy of Erica, and to his late wife and collaborator, Inge Magdalene Oliver (1947–2003), who was an authority on Erica in her own right. They have previously been honoured separately in E. ingeana E.G.H. Oliv. and E. oliveri H.A.Baker (
pro E. tenuis Salisb., Trans. linn. Soc 6: 329. 1802 [wfo-0000673389], nom. illeg., non Moench, Methodus: 17. 1802 [wfo-0000673388].
Without locality or collector, Ex herb. R. A. Salisbury (lectotype K [K000314799] [https://powo.science.kew.org/taxon/urn:lsid:ipni.org:names:329729-1]).
Masson only published one account of his botanical explorations at the Cape of Good Hope (
... the hot bath, which is situated at the foot of a ridge of dry mountains: ... Next morning, we went up to the top of this ridge of mountains ... We found here a species of heath remarkable for having its branches and leaves all covered with a fine hoary down or nap, which we thought singular in that genus: we called it Erica tomentosa.
It is often difficult to decide whether a sentence such as Masson’s constitutes a diagnosis as defined in the International Code of Nomenclature: ‘... a statement of that which, in the opinion of its author, distinguishes the taxon from others’ (ICN (Shenzhen Code) 2018, Art. 38.2; Turland et al. 2018). Given that, at this time, the early 1770s, only about fifty Erica species from the Cape Region had been described (
It follows that Salisbury’s binomial is illegitimate and has to be replaced. The heterotypic synonym, Erica velutina Bartl. (fide
Erica tomentosa Salisb., Trans. Linn. Soc. 6: 327. 1802 [wfo-0000673425], nom. illeg., non Masson, Phil. Trans. 66: 299. 1776 [wfo-1000053520] (= E. passerina Montin). Type. South Africa. “Hottentots Holland”, I. Mulder s.n. ex herb. Salisbury (not located).
A label identifying the specimen labelled “C.B.S. Niven 16” (K-000314197) [https://plants.jstor.org/stable/10.5555/al.ap.specimen.k000314197] as a lectotype is incorrect as the protologue cited only a collection from Hottentots Holland by I. Mulder and the Niven collection is, therefore, not original material.
South Africa. “Am Fusse des Babylonschenthurmbergen [Babilonstoring]”, Ecklon s.n. (holotype GOET-003270 [https://plants.jstor.org/stable/10.5555/al.ap.specimen.goet003270]).
The authors thank Dr Alan Elliot (Royal Botanic Garden Edinburgh) for his invaluable input relating to WFO and rediscovery of outstanding nomenclatural issues in Erica addressed here. Thanks are also due to Dr Larry Dorr and Dr John Manning for their careful reviews of the paper and for their many pertinent comments that have improved the paper. We acknowledge the support of The Heather Society (now disbanded) in the development of the International Register of Heather Names which originally brought these issues to light.
The authors have declared that no competing interests exist.
No ethical statement was reported.
The Heather Society.
ECN: Conceptualisation, Investigation, Writing - Original draft, Writing - Review and Editing. EGHO: Investigation, Writing - Review and Editing (prior to 2023). MDP: Investigation, Writing - Review and Editing (from 2023).
E. Charles Nelson https://orcid.org/0000-0002-3909-4388
Michael D. Pirie https://orcid.org/0000-0003-0403-4470
All of the data that support the findings of this study are available in the main text.