Details of the traits observed in P. marginatum var. Marginatum (A, C, E, G) and P. marginatum var. humbertii (B, D, F, H) A apex of a sterile pinna in the var. marginatum, adaxial surface (Cowemoy s.n., P01462834) B apex of a sterile pinna in var. humbertii, adaxial surface (Rakotondrainibe 1673, P00100192) C petiole surface in the var. marginatum (Cadet 4050B2, P01462767) D petiole surface in var. humbertii with a scar left by a scale pointed with an arrow (Rakotondrainibe 1673, P00100193) E fertile pinna cross section of the var. marginatum (Bradé 958, P00917035) F fertile pinna cross section of var. humbertii (Rakotondrainibe & Raharimalala 2519, P00904704) G sorus in the var. marginatum (Cadet 4050B1, P01462768) H sori of var. humbertii, with sporangiasters pointed with an arrow (Rakotondrainibe 2743, P00059959). Scale bar: 5 mm (A); 2.5 mm (B); 1 mm (C, D); 800 µm (E, F); 2 mm (G); 500 µm (H).