Morphological, ecological, and molecular phylogenetic approaches reveal species boundaries and evolutionary history of Goodyera crassifolia (Orchidaceae, Orchidoideae) and its closely related taxa

Goodyera crassifolia (Koji Tanaka KS209, KYO; photographed after immersion in 50 percent ethanol) A flower (lateral view) B flower (dorsal view) C dorsal sepal (abaxial view) D lateral sepals (left: abaxial view, right: adaxial view) E lateral petals (left: abaxial view, right: adaxial view) F lip and column (lateral view) G lip (left: adaxial view, middle: lateral view, right: abaxial view) H column (left: dorsal view, right: obliquely ventral view) I column with partially detached lateral appendages (left: ventral view, right: lateral view) J lateral appendages removed from column (ventral view) K anther cap and pollinarium (ventral view). Arrows indicate the conspicuous lateral appendages. All photographs are derived from the same flower. Scale bars: 3 mm.