Climacosphenia soulonalis sp. nov. All TK4 A–C holotype valve with valvocopula, entire, basal pole and apical pole, respectively, showing seamless craticular bars except near basal pole; arrowheads on B show complex unions D external valve apical pole showing irregular patch of areolae (arrow), spines, and narrowing of the annulus at apical pole E internal aspects of basal and apical poles, showing lack of costae (contrast costae in the piece of Synedrosphenia crystallina to the right) F, G valvocopula with two complex unions at the basal pole (arrowheads, F) and showing the row of pores on the pars interior around the basal pole (arrow). Scale bars: 100 µm (A); 10 µm (B–G).