Characters;"L. moorei";"L. longini";"L. boninensis";"L. spectabilis" "Vein apices ending at the laminar margins";yes;no;yes;yes "Widest part at upper pinna";<1/2;<1/3;<1/2;~1/2 "Stipe scales";"dark brown, narrowly lanceolate (usually < 2 mm wide)";"dark brown, broadly lanceolate (usually >3 mm wide)";"dark brown, broadly lanceolate (usually >2 mm wide)";"light brown, narrowly lanceolate (usually <2 mm wide)" "Fertile pinna pairs";10–14;3–9;4–16;6–14 "Swollen ring on articulate at abaxial side, especially upper pinnae (Fig. 6)";obvious;obvious;inconspicuous;inconspicuous "Spore perine (Fig. 4)";spiny;folded;spiny;cristate "Spore number per sporangium";64;32;64;"16 or 32"