Nutlet morphology of Pseudolappula and Lappula A–D Pseudolappula sinaica A fruit lateral view B fruit polar view (growth direction of nutlets indicated by arrows) C nutlet abaxial view D nutlet adaxial view E–H Lappula occultata E fruit lateral view F fruit polar view (arrangement of nutlets indicated by cross lines) G nutlet abaxial view H nutlet adaxial view I–L Lappula patula I fruit lateral view J fruit polar view (arrangement of nutlets indicated by cross lines) K nutlet abaxial view L nutlet adaxial view M adaxial keel of P. sinaica (adaxial view) N adaxial keel of L. occultata (lateral view) O attachment scar of Lappula balchaschensis P attachment scar of Lappula brachycentra. Scale bars: 1 mm.