Corresponding author: John L. Clark (
Academic editor: Eberhard Fischer
Two new species,
Se describen dos especies nuevas de
Clark JL, Clavijo L (2022) Two new species of
The flowering plant family
General geographic distribution (names in parentheses indicate provinces for Ecuador or departments for Colombia) and comparison of morphological characters to differentiate
|
isophyllous | isophyllous or subequal | anisophyllous | isophyllous or subequal | isophyllous or subequal | isophyllous or subequal |
|
1.5–7.7 cm | 0.3–0.6 cm | 1.0–2.0 cm | 0.7–3.0 cm | 0.47–5.24 cm | 0.5–3.5 cm |
|
eucamptodromous | brochidodromous | brochidodromous | eucamptodromous | eucamptodromous | eucamptodromous |
|
strongly laciniate with unbranched filiform teeth | laciniate with unbranched filiform teeth | strongly laciniate with branched filiform teeth | deeply serrate to slightly laciniate | serrate to laciniate | deeply serrate to pinnatifid |
|
flat | flat | crispate | flat | flat | crispate |
|
2.8–3.5 cm | 2.0–3.5 cm | 3.0–4.6 cm | 6.5–8.0 cm | 2–3.5 cm | 2–4.5 cm |
|
Costa Rica & Panama | western Andean slopes in northern Ecuador (Carchi & Esmeraldas) | western Andean slopes in northern Ecuador (Carchi & Esmeraldas) | western Andean slopes in northern Ecuador (Carchi & Esmeraldas) and Colombia (Choco, Valle de Cauca & Nariño); eastern Andean slopes in southern Ecuador (Morona-Santiago) | widespread in Central and South America | eastern Andean slopes of southern Ecuador (Morona-Santiago) |
Differs from all
Ecuador. Esmeraldas: cantón San Lorenzo, parroquia Alto Tambo, trail from the community Durango to Río Tululbi via trail north of highway San Lorenzo-Ibarra, forest managed by Fundación Sirua,
Terrestrial, hemiepiphytic or epiphytic herb or subshrub, with scandent to horizontal shoots to 1 m long. Stems subquadrangular in cross-section, 0.3–0.5 cm in diameter, strigose to glabrate, internodes 2.5–8 cm long.
This species has been collected with flowers in May and June. Specimens with fruits are only known from May.
The specific epithet is in reference to the intermediate shapes and sizes of the corolla and calyx between
The characters that differentiate
Differs from all
Ecuador. Esmeraldas: cantón San Lorenzo, parroquia Alto Tambo, remnant patch of primary forest on north side of road between Durango and Alto Tambo on highway San Lorenzo-Ibarra,
Terrestrial, hemiepiphytic or epiphytic herb or subshrub, with scandent to horizontal shoots to 1 m long. Stems subquadrangular in cross-section, 0.3–0.6 cm in diameter, strigose to glabrate, internodes 1.0–8 cm long.
This species has been found with flowers in two periods: February to May and August to October.
The specific epithet is in reference to the elongate corolla tube, the longest of any known species of
Our preliminary DNA sequence data strongly support a close relationship in a clade that includes
The National Science Foundation provided support for Clark (DEB-0841958 & DEB-0949169). We are especially grateful to Blanca Bisvicuth from Fundación Sirua (Ecuador) for guiding us on several expeditions throughout the western Andean slopes, including the 2008 expedition that resulted in the type collection. We thank Parques Nacionales Naturales de Colombia, especially Robinson Galindo-Tarazona, the former director of the Parque Nacional Natural Farrallones de Cali, Jaime Celis, CELSIA, in particular Yaneth Hernandez-Montoya, Alejandro Zuluaga (Universidad del Valle), Mathieu Perret (Conservatoire et Jardin botaniques de la Ville de Genève), and the Fundación FELCA for facilitating field expeditions that allowed us to document