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Article title
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Abstract
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Keywords
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Introduction
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Materials and methods
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Morphological comparison
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Herbarium resources
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Pollen morphology
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Karyotype analysis
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Distribution map
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Data analysis
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Results
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Macro-morphological differences
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Pollen morphology
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Chromosome cytology
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Distribution
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Discussion
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The differences and the taxonomic status of the unknown Cirsium
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Key to the 14 wild Cirsium taxa of Taiwan (modified from Chang et al. (2019), Chang and Tseng (2019), and Chang and Tseng (2020)
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Taxonomic treatment
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Acknowledgements
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References
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