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Article title
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Abstract
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Keywords
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Introduction
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Materials and methods
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Specimen collection and morphological character coding
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Biogeographic information for specimens
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DNA extraction, PCR amplification and DNA sequencing
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Character-based DNA barcoding
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Reproductive phenology data
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Results
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Biogeographic patterns
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Comparisons of quantitative and qualitative morphological characters
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Character-based DNA barcoding
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Ovuliferous reproductive phenology
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Integrative taxonomic inference of a new Ceratozamia species
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Taxonomic treatment
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Ceratozamia leptoceras
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Taxonomic keys to Ceratozamia leptoceras and morphologically related species
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Discussion
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Conclusions and future directions
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Acknowledgements
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References
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Appendix I
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Supplementary materials
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