Corresponding author: Xinjian Zhang (
Academic editor: R. Kriebel
A nomenclatural and taxonomic treatment of the name
Zhang X, Dai J-H, Liu X, Li Z, Lee SY, Zhou R, Tan G (2020) Lectotypification of the name
In China, nine
In this study, we are in agreement with the recognition of
During a recent field survey, we collected an unidentified specimen of
Morphological data for identification and description of this taxon were based on observations of specimens in the field (ten individuals) and the herbarium. The voucher specimens are deposited in the
Detailed morphological examination revealed that morphological characters of this unidentified taxon of
The white form of
(designated here): – China. Taiwan. Ilan, Yuensanhsiang, leg.
Perennial shrubs, 0.3–1.5 m tall. Twigs nearly 4-angled to subterete in the younger parts and terete in the old parts, densely covered with appressed to suberect strigose with scales. Leaf blades ovate to elliptic, papery, 3.3–4.8×6–9.5 cm, base broadly cuneate to rounded or subcordate, apex acuminate, margin entire, palmately 7-nerved (the marginal nerves often inconspicuous), adaxially densely strigose, abaxially densely puberulous, strigose along veins; petioles 1.4–2.0 cm, densely strigose with scales. Inflorescences subcapitate corymbose, terminal, 3–5-flowered, with 2 leaf-like bracts at base. Pedicels 8–12 mm, densely strigose with scales; bracteoles 2, opposite, elliptic-lanceolate to elliptic, 6–13 mm, abaxially densely strigose, margin ciliate. Hypanthia 7–12 mm, densely appressed-strigose with scales, margin fimbriate. Sepals lanceolate to ovate-lanceolate, apex acuminate, densely strigose and pubescent on both sides and along the margin. Petals 5, occasionally 6, white, obovate, ca. 27×18 mm, apex rounded. Stamens 10, dimorphic, longer stamens with anthers linear, curved, ca. 9 mm, filaments ca. 10 mm, joined by a connective ca. 9 mm, curved, spur bifid ca. 2 mm, shorter stamens with anthers ca. 8 mm, 2-tuberculate at base, filaments ca. 7 mm, without prolonged connective. Ovary half-inferior, campanulate, with a ring of bristles at apex. Capsule dry, urceolate, apically dehiscent, 9–16×7–10 mm, densely squamose strigose. Seeds numerous, minute, cochleate. Fl. May-Aug, fr. Aug-Oct.
The white-flowered form of
Distribution map of the white form of
The key character of the white form of
The flowers of the white form of
– China. Fujian Province, Pinghe County, in lowland evergreen forest margins, 24°02.66'N, 117°04.75'E, Elev. 276 m. 28 July 2019, X. J. Zhang,
We thank Ms. Siyi Cai, of the National Research Institute of Chinese Medicine, Taiwan, for providing the protologue. We are also grateful to Prof. Wenbo Liao, Prof. Ying Liu and Dr. Wanyi Zhao, all from Sun Yat-sen University, for helpful comments on the taxonomic treatment.