Corresponding author: Richard K. Rabeler (
Academic editor: C. Morden
Twenty-one new combinations in
Rabeler RK, Wagner WL (2015)
In their study of relationships in
Three subsequent molecular studies have confirmed the placement of the
The information about type specimens of the basionyms of the new combinations that we have included is based on examining protologues and searching major indices (Tropicos –
In the cases where syntypes are cited, we have refrained from designating lectotypes. It is not a requirement for the names to be validly and effectively published and we consider those decisions should be made during the course of a taxon-level revision where a serious study of all specimens would lead to the best selections.
(see McNeill in Notes Roy. Bot. Gard. Edinburgh 24: 120. 20 Sep 1962):
Plants perennial, sometimes densely cespitose or pulvinate or with a stout woody caudex, very rarely annual. Leaves filiform to subulate, often long-linear and grass-like, sometimes short and setaceous or needle-like, congested in vegetative rosettes and at or near base of flowering stems, margin sometimes scarious, apex often apiculate. Inflorescence of one or more terminal cymes, sometimes compressed to head-like, sometimes flowers solitary or paired. Flowers weakly perigynous. Sepals often hardened at base, veins 1–3, margins often white-scarious. Petals white, rarely pink. Floral glands (nectaries) at base of the antisepalous filaments often lobed, conspicuous.
CHINA. Xinjiang: Aksayqing, ca. 4900 m. Northwest Plat. Inst. Biol. Acad. Sin. Exped. 3723 (holotype, NWBI).
TURKEY: Adiyaman, Mt. Nemrut Dagi, 1950 m, 5 July 1983, T. Ҫelebioğlu 83-8/1 (holotype, ISTF).
INDIA: Alpine Sikkim Himalaya, Yeumtong in gravelly places, alt. 15000 ft.; J.D. Hooker s.n. (holotype, K?; possible isotype, GH, GH00353889 [JSTOR image!]).
INDIA: Kumaon, Kali Valley, on rocks near Byans, 2800–3000 m, J.F. Duthie 2762 (BM?).
Kunawar (K?).
Caucasus. circa Kobi (LE?).
CHINA. Xizang: Ngari Diqu, Geze, 4500–4700 m, collector unknown 4346 (holotype, NWBI).
CHINA: Tschili (Hebei). Hsiau Wu tai schan, Felsen des Gipfels, 3050 m, 3 Aug 1912, H.W. Limpricht 600 (WRSL?, WU, WU0029891 [JSTOR image!]); Gipfelfelsen des Pe tai, 4050 m, Kalk, 26 Jun 1915, H.W. Limpricht 2563 (WRSL?, WU, WU0029892 [JSTOR image!]); Felsen des His tai, 3000 m, H.W. Limpricht 2982 (WRSL?). West-Tibet: Hor Tschango, Schtiala, Geröll des Schao kirr bu, 4700 m, H.W. Limpricht 2095 (WRSL?); Kanse, Tsokoma-Stock, Geröllhalden, 5000 m, H.W. Limpricht 2127 (WRSL?). Batang-Litang Pungtschamu, Felsen des Passes Dshagala, 5260 m, H.W. Limpricht 2262 (WRSL?).
This combination is proposed based on the placement of this taxon in subg.
CHINA. Sichuan: Dege, Haizi Shan, 3800 m., Sichuan Exped. D-7475 (holotype, WUG).
CHINA. Xizang: Khamba Fort, 17 July 1903, F.E. Younghusband 107 (holotype, K, K000723883 [JSTOR image!]; E, E00317561 = photo of specimen at K!).
CHINA: northern China, A.A. Tatarinoff (LE?).
INDIA: Kumaon, Ralam valley, 12500 ft., R. Strachey & J.E. Winterbottom 3 (BM, BM000946338 [JSTOR image!]; K, K000742179 [JSTOR image!]; LE?); Ralam valley, 14000-16000 ft., 1884, J.F. Duthie 2757 (LE?); Tihri-Garhwal, J.F. Duthie s.n. (LE?).
CHINA. Yunnan: Weixi, 4000 m, T.T. Yü 8966 (holotype, PE).
NEPAL: Muktinath, 4250 m, T.B. Shrestha & Bista 2462A (holotype, CAL).
Songaria, non procul a finibis Mongoliae, in montibus Kitschni-ne-tau, prope fortalitium Saissan, in rupibus, Potanin (LE?).
INDIA. Sikkim: 14000-17000 ft. [Lana Kayna, 15000 ft., 24 July 1849 (on additional label on K sheet)], J.D. Hooker, Hooker & Thomson, Herb. Ind. Orient. 3 (Holotype, K, K000723992 [JSTOR image!]; isotypes, BM, BM000946339 [JSTOR image!], BM000946343 [JSTOR image!]).
CHINA. Qinghai: Dulan Xian, 4200 m, Qinghai-Gansu Exped. 1194 (holotype, WUG).
CHINA. Tibet: ad fl. Yang-tze, 20 July 1884, Przewalski? (LE?).
CHINA. Xizang: Lunzhe, ca. 4300 m, B.Z. Guo et al. 22435 (holotype, NWBI).
CHINA. Shaanxi: Meixian, side of Dayehai, ca. 4000 m, Shaanxi Chin. Herb. Medic. Exped. 390 (holotype, WUG).
CHINA. Qinghai: Zado, ca. 4400 m, S.W. Liu 110 (holotype, WUG).
We acknowledge Kanchi Gandhi (GH) for nomenclatural advice.