Schemes (A, C) and cross-sections of the achenes (B, D). A, B Scorzonera longipapposa (Scorzonera polyclada clade), B from Rechinger 35489 (G181688) C, D Koelpinia macrantha (Koelpinia clade), D from Pimenov et al. 447 (MW0891267). Designation of the colours: yellow – thin-walled parenchyma (absent in C), hatched area – thick-walled parenchyma; pink (C) – sclerenchyma obliquely orientated in the region of the wings (present in C); red – sclerenchyma (of parallel orientation); blue – seed coat; violet dots (A, C) – vestiges of vascular bundles in the pericarp and seed coat (A) and in the seed coat (C); green – endosperm; central white area – seed hollow.