The reticulogram of the D. hachijoense complex. Thin-solid arrow, autopolyploidization; Solid arrows, maternal inheritance; dashed arrows, paternal inheritance; square, triploid apogamous or diploid sexual species; dashed square, hypothesized diploid sexual or triploid apogamous species. *D. okinawaense had only nuclear AK1 allele A of D. dilatatum and B of D. takii, although plastid haplotype was D of D. amaminanum.