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  • Article title
  • Abstract
  • Key words
  • Introduction
  • Materials and methods
    • Morphological observation
    • Spore morphology and ornamentation
    • Assessing ploidy via genome size measurement
    • Taxon sampling, DNA extraction, and sequencing
    • Plastome assemblage and annotation
    • Phylogenetic reconstruction
  • Results
    • Plastome characteristics of Huperzia
    • Phylogenetic relationships among Huperzia species
    • Macro- and micro-morphological comparison
    • Taxonomic treatment
  • Discussion
  • Acknowledgements
  • Additional information
    • Conflict of interest
    • Ethical statement
    • Funding
    • Author contributions
    • Author ORCIDs
    • Data availability
  • References
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