Corresponding author: Young-Dong Kim (
Academic editor: P. Stoev
Cho S-H, Kim B-Y, Park H-S, Phourin C, Kim Y-D (2017)
In Indochina, there are around 120 species of
Except for
During the recent floristic survey, one species of
CAMBODIA. Kampot Province, Phnum Bokor National Park, sandstone tables in evergreen forest margin,
Comparison of key features of
Taxonomic traits |
|
|
---|---|---|
Leaf | ||
upper surface | pubescent | pubescent with scurfy hairs |
lower surface | glabrous to sparsely pubescent | pubescent on nerves only or tomentose all over |
strigose-ciliate at base | 1–2.7 mm | 1–1.4 mm |
Pedicel | 1.4–2.0 cm | 2.3–3(–5) cm |
Dorsal petal | orbicular-obovate, ca. 6 × 8 mm, horned at apex, horn 2.2–2.3 mm long | cordate, 7 mm long, horned at apex, horn 3 mm long |
Seeds | 3.8–4.6 × 2.6–3.2 mm | 3.5 × 2.5 mm |
Herbs, annual, terrestrial, hermaphroditic. Stems erect, 15–40 cm tall, tinged purplish red, branched, glabrous or sparsely puberulous on the upper part. Leaves simple, alternate; petioles subsessile to 1.4 mm; leaf blade lanceolate to ovate-lanceolate, apex acuminate, base narrowly cuneate to attenuate, 2.5–7.0 × 0.6–2.0 cm, upper surface pubescent, lower surface glabrous to sparsely pubescent, secondary veins pinnate, 6 to 8 on each side of mid-vein, margin serrate, teeth mucronate and purple tinged, strigose-ciliate at base; strigose-ciliate, 4–14, 1–2.7 mm long, purple to purplish black, minutely puberulous at base. Flowers axillary, solitary, rarely 2 fascicled, zygomorphic, minutely puberulous; pedicels slender, erect, purplish red, 1.4–2.0 cm long, glabrous, bracteate at base; bracts linear, up to 4 mm; lateral sepals 2, linear-lanceolate, 2.5–3 mm long, glabrous; lower sepal funnel-form, pink, ca. 5 mm long, ca. 3 mm deep; spur 17–23 mm long, slightly curved; dorsal petal, orbicular-obovate, ca. 6 × 8 mm, horned at apex, horn 2.2–2.3 mm long; lateral united petals separate, bilobed, ca. 11 mm long; upper petals oblong, 6.0–6.5 × ca. 3.0 mm, minutely apiculate; lower petals, 9.0–9.3 × 3.5–3.8 mm; androecium ca. 2.8 × 1.6 mm; stamens 5, connate, surrounding gynoecium; filaments ca. 0.7 mm; ovary fusiform, pubescent, ca. 2.5 × 1.0 mm; style glabrous, ca. 0.2 mm long; stigma 5, ca. 0.25 mm long. Fruit a capsule, fusiform, ca. 15 × 6 mm, pubescent with scurfy hairs, 3[4]-seeded. Seeds obovoid, slightly compressed, 3.8–4.6 × 2.6–3.2 mm, pubescent with spirally sculptured hairs.
CAMBODIA. 16 November 2015, with fruits, Cho et al.
Flowering specimens were collected in August and fruiting specimens in November.
Cho et al.
We thank Ms. Hye-Woo Shin (