Corresponding author: Ruijiang Wang (
Academic editor: S. Razafimandimbison
A new species of
Wen H, Wang R, Deng S (2015)
The genus
During a field investigation in 2013, we found a semi-erect and hairy herb, which was similar to
All materials were collected by ourselves and deposited at the herbarium of South China Botanical Garden, the Chinese Academy of Sciences (IBSC). The leaf materials were carefully taken from the specimen
Perennial herbs, prostrate when young and decumbent while growing, rooting at nodes adhering to ground; stems terete, densely pubescent. Stipules linear, 3.8–6.5 mm long. Petioles 0.5–2 cm long, sparsely pubescent; leaf blades oval, ovate to broadly ovate, 1.1–5 (–6.2) × 0.5–3 cm, base cuneate to rounded, apex acute, papery, hairy both sides; secondary veins (5–)7–10 on each side, conspicuous, depressed adaxially and projected abaxially; leaf epidermal cells irregularly polygonal both sides, with sinuous anticlinal walls; stomata paracytic. Inflorescences terminal or upper axillary, cymose, (5–) 10–20-flowered, usually condensed into capitate-like; peduncles pubescent, 5–12 mm long, to 3 cm while fruiting; bracts linear, 6–10 mm long. Flowers distylous, 5-merous; pedicels sparsely pubescent, 1–3 mm long, usually condensed. Hypanthium obconical, 1–2 mm long, sparsely pubescent; lobes 5, linear, 2.5–4 mm long, pubescent. Corollas tubular, white, sparsely hairy abaxially, tubes 2–3 mm long, ca. 1 mm wide; lobes subovate, ca. 1.5 × 1 mm; stamens 5; anthers oblong-linear, ca. 1 mm long; stigmas bilobed; ovary 2-celled, ovules on the axile placentas. Long-styled flowers: a ring of pubescence in throat; stamens below the pubescent ring and included, filaments adnate to the base of corolla tube, ca. 1 mm long; styles exserted, ca. 3 mm long, stigma lobes capitate. Short-styled flowers: corolla tubes densely pubescent inside; stamens extend to the corolla throat but included, filaments adnate to the upper portion corolla tube, ca. 1 mm long; styles included, ca. 1 mm long, stigma lobes ovate. Fruits capsular, ovoid to subglobose, 3–4 mm, hairy, dehiscing loculicidally and then septicidally; valves 4, crustaceous, ca. 3.5 mm long; calyx lobes persistent, 1.8–2 mm long. Seeds ca. 30 per capsule, ca. 0.5 mm long, rectangular pyramid, brown, testa papillate.
Morphological characters of
Known only from the type locality. The plants grow in the entrances of limestone caves or moist cliffs of hill sides.
Flowering in May–June and fruiting in June–August.
The number of
Similar to
This work was supported by National Natural Science Foundation of China (31070177). We thank Professor Yigang Wei and Dr. Bo Pan from Guangxi Institute of Botany, Guangxi Zhuangzu Autonomous Region and Chinese Academy of Sciences for their kind help during our field investigation; Dr. Bine Xue and Miss Shuangshuang Zhu for helpful comments on the manuscript.