Introduction
As we continue to emerge from the “Dark Ages of Lumping” it should come as no surprise to any student of the Compositae that morphological and molecular data are being used (separately and together) to address generic limits. As a result we are seeing the breakup of many large non-monophyletic genera. In one such case, the genus Vernonia Schreb., of the tribe Vernonieae, has shrunk to about 20 species from North America and the remaining 1000 or so species from that genus are in the process of being assigned to other genera. Many of these “new” genera were previously described and subsequently sunk into Vernonia, but others needed new names and descriptions. An overview of the tribe was presented by Keeley and Robinson (2009) and major overhauls have taken place for the Americas (Robinson 1999), China (Robinson and Skvarla 2010), Thailand (Bunwong et al. 2014) and Southern Africa (Swelankomo and Manning 2014; Robinson et al. 2016). Still awaiting work are the Vernonieae of Madagascar, tropical Africa, India, and SE Asia.
As part of a larger more comprehensive work on the Vernonieae of Madagascar this effort concerns the establishment of proper limits for the genus Distephanus Cass. Described by Cassini based on a type removed from Conyza (Astereae), Distephanus has long been recognized as distinctive (trees, shrubs or woody vines; yellow or orange flowers; tri-nervate leaf venation) even when it was considered part of Vernonia (herbs or subshrubs; purple, pink or white flowers; pinnate leaf venation). In addition to the type species three other species were described in Distephanus and therefore do not need to be transferred and a number of combinations have already been made: 24 by Robinson and Kahn (1986), one by Robinson (2009), two by Robinson (2012), one by Robinson and Funk (in Funk et al. 2012) and one by Boon and Glen (2013) for a total of 33 species currently in Distephanus. However, because of a lack of available herbarium material some of the Madagascar species names were left as Vernonia. After examining herbarium material from the Muséum National d’Histoire Naturelle (P), nine additional combinations can now be made. The following combinations are needed at this time because of upcoming entries in GenBank and determinations on specimens from a recent field trip in September-October of 2016. Descriptions and synonomy can be found in Humbert (1960: Volume 1, 121–171). With these additions, the number of species in Distephanus now stands at 42 with possible new species to be described based on material collected during the aforementioned field trip.